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<mods:title id="A42D6556BA05DA2318BF66E3A3F5A983">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
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<mods:namePart id="DABDD9F4FDE524F75C6318393746C23D">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="99ED21B08121BAC40B22CEE48E03ECC0">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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<mods:namePart id="D02CB0BD2A5149CE9242B757172326BE">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:affiliation id="7A8A0248BBEE5787F4692CF212573A0D">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:title id="7848C0695C4A6BCF6298A216B121FD3C">Proceedings of the California Academy of Sciences</mods:title>
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<mods:date id="D043B2DFD8E074C3E8D5BEC46D7CAF49">2023</mods:date>
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<mods:number id="48D8B111EF10D51B2B240F2061DD4BB2">2023-02-15</mods:number>
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<treatment id="1C1187FCEB66ED57FF9EFB71611A8501" ID-DOI="http://doi.org/10.5281/zenodo.12726723" ID-GBIF-Taxon="235039992" ID-Zenodo-Dep="12726723" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501" httpUri="http://treatment.plazi.org/id/1C1187FCEB66ED57FF9EFB71611A8501" lastPageId="18" lastPageNumber="511" pageId="14" pageNumber="507">
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<subSubSection id="DCA26561EB66ED4BFF9EFB7162E9828E" box="[123,462,1246,1270]" pageId="14" pageNumber="507" type="nomenclature">
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<paragraph id="940736EAEB66ED4BFF9EFB7162E9828E" blockId="14.[123,1174,1246,1726]" box="[123,462,1246,1270]" pageId="14" pageNumber="507">
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<heading id="CF4F8186EB66ED4BFF9EFB7162E9828E" box="[123,462,1246,1270]" fontSize="10" level="2" pageId="14" pageNumber="507" reason="4">
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<taxonomicName id="53B84D69EB66ED4BFF9EFB7162E9828E" ID-CoL="32X57" authority="Andrewes" authorityName="Andrewes" box="[123,462,1246,1270]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="inustus">
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<emphasis id="A6CCEAF8EB66ED4BFF9EFB7162E9828E" bold="true" box="[123,462,1246,1270]" pageId="14" pageNumber="507">
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<emphasis id="A6CCEAF8EB66ED4BFF9EFB716271828D" bold="true" box="[123,342,1246,1270]" italics="true" pageId="14" pageNumber="507">Cyclosomus inustus</emphasis>
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Andrewes
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB66ED4BFF9EFB5162F2836F" box="[123,469,1278,1300]" pageId="14" pageNumber="507" type="description">
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<paragraph id="940736EAEB66ED4BFF9EFB5162F2836F" blockId="14.[123,1174,1246,1726]" box="[123,469,1278,1300]" pageId="14" pageNumber="507">
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<figureCitation id="0C832A6FEB66ED4BFF9EFB5163F2836F" box="[123,213,1278,1300]" captionStart="FIGURE" captionStartId="1.[171,183,963,982]" captionTargetBox="[120,1176,216,952]" captionTargetId="figure-323@1.[120,1176,216,952]" captionTargetPageId="1" captionText="FIGURE 1. Cyclosomus inustus Andrewes, adult male feeding on a small fly (photograph reproduced from Wang et al. 2017 with permission of Liang-Jong Wang)." figureDoi="http://doi.org/10.5281/zenodo.11512956" httpUri="https://zenodo.org/record/11512956/files/figure.png" pageId="14" pageNumber="507">Figures 1</figureCitation>
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,
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<figureCitation id="0C832A6FEB66ED4BFF07FB5063D7836F" box="[226,240,1279,1300]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="14" pageNumber="507">7</figureCitation>
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,
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<figureCitation id="0C832A6FEB66ED4BFF1FFB516203836F" box="[250,292,1278,1300]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="14" pageNumber="507">14C</figureCitation>
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,
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<figureCitation id="0C832A6FEB66ED4BFECAFB51627E836F" box="[303,345,1278,1300]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="14" pageNumber="507">16C</figureCitation>
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,
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<figureCitation id="0C832A6FEB66ED4BFE81FB5162A9836F" box="[356,398,1278,1300]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="14" pageNumber="507">17C</figureCitation>
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,
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<figureCitation id="0C832A6FEB66ED4BFE7DFB516295836F" box="[408,434,1278,1300]" captionStart="FIGURE" captionStartId="38.[155,167,1584,1603]" captionTargetBox="[120,1176,216,1564]" captionTargetId="figure-9@38.[120,1176,216,1564]" captionTargetPageId="38" captionText="FIGURE 19. Photographs of habitats for Cyclosomus inustus Andrewes. A. Lingshanbi, Shihmen District, New Taipei City, Taiwan; B. Shalun, Danshui District, New Taipei City, Taiwan. Photographs reproduced from Wang et al. 2017 with permission of Liang-Jong Wang." figureDoi="http://doi.org/10.5281/zenodo.11512998" httpUri="https://zenodo.org/record/11512998/files/figure.png" pageId="14" pageNumber="507">19</figureCitation>
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,
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<figureCitation id="0C832A6FEB66ED4BFE58FB5162F2836F" box="[445,469,1278,1300]" captionStart="FIGURE" captionStartId="39.[171,183,837,856]" captionTargetBox="[168,1128,216,825]" captionTargetId="figure-14@39.[168,1128,216,825]" captionTargetPageId="39" captionText="FIGURE 20. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow circles = C. acutangulus Kavanaugh and Cueva-Dabkoski; white triangles = C. flexuosus Dejean; yellow triangle = locality where C. acutangulus and C. flexuosus have been found together; white squares = C. inustus Andrewes. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/ Whole_world_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513000" httpUri="https://zenodo.org/record/11513000/files/figure.png" pageId="14" pageNumber="507">20</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB66ED4BFF9EFA8C675F832D" pageId="14" pageNumber="507" type="reference_group">
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<paragraph id="940736EAEB66ED4BFF9EFA8C675F832D" blockId="14.[123,1174,1246,1726]" pageId="14" pageNumber="507">
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<treatmentCitationGroup id="B4A811C4EB66ED4BFF9EFA8C67B18342" box="[123,1174,1315,1337]" pageId="14" pageNumber="507">
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<treatmentCitation id="151910FBEB66ED4BFF9EFA8C612E8342" author="ANDREWES, H. E." box="[123,521,1315,1337]" page="464" pageId="14" pageNumber="507" year="1924">
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<taxonomicName id="53B84D69EB66ED4BFF9EFA8C612E8342" ID-CoL="32X57" authority="Andrewes, 1924: 464" authorityName="Andrewes" authorityPageNumber="464" authorityYear="1924" box="[123,521,1315,1337]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="inustus">
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<emphasis id="A6CCEAF8EB66ED4BFF9EFA8C621E8342" box="[123,313,1315,1337]" italics="true" pageId="14" pageNumber="507">Cyclosomus inustus</emphasis>
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<bibRefCitation id="F0294B1BEB66ED4BFEDBFA8C612E8342" author="ANDREWES, H. E." box="[318,521,1315,1337]" pageId="14" pageNumber="507" pagination="459 - 468" refId="ref23019" refString="ANDREWES, H. E. 1924. XXII. On the Oriental Carabidae of the " Reise Novara. " Transactions of the Entomological Society of London 1923: 459 - 468." type="journal article" year="1924">Andrewes, 1924:464</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="4B038848EB66ED4BFDF7FA8C61B48342" box="[530,659,1315,1337]" pageId="14" pageNumber="507" type="holotype">HOLOTYPE</typeStatus>
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, a male, deposited in
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<collectionCode id="F2A9AE2FEB66ED4BFC80FA8C60E48343" box="[869,963,1315,1336]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="507" type="Museum">NHMUK</collectionCode>
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. Type locality:
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<collectingCountry id="ECAF767AEB66ED4BFBB3FA8C67B58342" box="[1110,1170,1315,1337]" name="China" pageId="14" pageNumber="507">China</collectingCountry>
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,
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</treatmentCitationGroup>
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<collectingCountry id="ECAF767AEB66ED4BFF4EFAEF6238832E" box="[171,287,1344,1365]" name="Hong Kong" pageId="14" pageNumber="507">Hong Kong</collectingCountry>
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.
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<bibRefCitation id="F0294B1BEB66ED4BFECCFAEF62FC832D" author="CSIKI, E." box="[297,475,1344,1366]" pageId="14" pageNumber="507" pagination="1279 - 1598" refId="ref23258" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1295)</bibRefCitation>
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;
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<bibRefCitation id="F0294B1BEB66ED4BFE03FAEF61B9832D" author="LORENZ, W." box="[486,670,1344,1366]" pageId="14" pageNumber="507" refId="ref23840" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz (2005:452)</bibRefCitation>
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;
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<bibRefCitation id="F0294B1BEB66ED4BFD4DFAEF6050832D" author="BOUSQUET, Y." box="[680,887,1344,1366]" pageId="14" pageNumber="507" refId="ref23176" refString="BOUSQUET, Y. 2017. Tribe Cyclosomini. Pages 498 - 499 in I. Lobl and D. Lobl, eds., Catalogue of Palaearctic Coleoptera. Volume 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden, xxxiv + 1143 pp." type="book" year="2017">Bousquet (2017:498)</bibRefCitation>
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;
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<bibRefCitation id="F0294B1BEB66ED4BFC64FAEF6708832D" author="WANG, L. J. & J. J. CHERNG & D. H. KAVANAUGH" box="[897,1071,1344,1366]" pageId="14" pageNumber="507" pagination="107 - 116" refId="ref23969" refString="WANG, L. J., J. J. CHERNG, AND D. H. KAVANAUGH. 2017. First records for Cyclosomus inustus Andrewes (Coleoptera: Carabidae: Cyclosomini) for Taiwan, with notes on habitat and behavior. Proceedings of the California Academy of Sciences (Series 4), 64: 107 - 116." type="journal article" year="2017">Wang et al. (2017</bibRefCitation>
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,
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<bibRefCitation id="F0294B1BEB66ED4BFBDCFAEF674A832D" author="WANG, L. J. & J. J. CHERNG & U. ONG" box="[1081,1133,1344,1366]" pageId="14" pageNumber="507" pagination="112 - 115" refId="ref24032" refString="WANG, L. J., J. J. CHERNG, AND U. ONG. 2022. New records of Cyclosomus inustus Andrewes (Coleoptera: Carabidae: Cyclosomini) in Taiwan and Kinmen. Japanese Journal of Systematic Entomology 28: 112 - 115." type="journal article" year="2022">2022</bibRefCitation>
|
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|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB66ED4BFF4EFACA62B180C6" pageId="14" pageNumber="507" type="description">
|
|||
|
|
<paragraph id="940736EAEB66ED4BFF4EFACA62B180C6" blockId="14.[123,1174,1246,1726]" pageId="14" pageNumber="507">
|
|||
|
|
<emphasis id="A6CCEAF8EB66ED4BFF4EFACA61188307" bold="true" box="[171,575,1381,1404]" pageId="14" pageNumber="507">Notes on types and nomenclature.</emphasis>
|
|||
|
|
<location id="91676031EB66ED4BFDAFFACA61478307" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB66ED4BFDAFFACA61478307" box="[586,608,1381,1404]" country="Hong Kong" county="Andrewes" municipality="Andrewes'" name="In" pageId="14" pageNumber="507">In</location>
|
|||
|
|
his original description of this species,
|
|||
|
|
<bibRefCitation id="F0294B1BEB66ED4BFBCCFACB63D683E7" author="ANDREWES, H. E." pageId="14" pageNumber="507" pagination="459 - 468" refId="ref23019" refString="ANDREWES, H. E. 1924. XXII. On the Oriental Carabidae of the " Reise Novara. " Transactions of the Entomological Society of London 1923: 459 - 468." type="journal article" year="1924">Andrewes (1924:464)</bibRefCitation>
|
|||
|
|
indicated that he had compared “
|
|||
|
|
<specimenCount id="82BEFD63EB66ED4BFDB0FA2B603F83E7" box="[597,792,1412,1436]" count="16" pageId="14" pageNumber="507" type="generic">sixteen specimens</specimenCount>
|
|||
|
|
labeled “
|
|||
|
|
<collectingCountry id="ECAF767AEB66ED4BFC99FA2B60D883E6" box="[892,1023,1412,1437]" name="Hong Kong" pageId="14" pageNumber="507">Hong Kong</collectingCountry>
|
|||
|
|
: and “
|
|||
|
|
<collectingCountry id="ECAF767AEB66ED4BFBA4FA2B67AF83E7" box="[1089,1160,1412,1436]" name="China" pageId="14" pageNumber="507">China</collectingCountry>
|
|||
|
|
”: these latter include examples in the
|
|||
|
|
<location id="91676031EB66ED4BFDE6FA0B619383C7" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB66ED4BFDE6FA0B619383C7" box="[515,692,1444,1468]" country="Hong Kong" county="Andrewes" municipality="Andrewes'" name="British Museum" pageId="14" pageNumber="507">British Museum</location>
|
|||
|
|
,
|
|||
|
|
<location id="91676031EB66ED4BFD24FA0B673E83C7" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB66ED4BFD24FA0B673E83C7" box="[705,1049,1444,1468]" country="Hong Kong" county="Andrewes" municipality="Andrewes'" name="Cambridge University Museum" pageId="14" pageNumber="507">Cambridge University Museum</location>
|
|||
|
|
, the “Reise Novara” example, and specimens from my own collection” with specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB66ED4BFC05FA6A674283A7" authorityName=", Kavanaugh" authorityYear="2015" box="[992,1125,1476,1500]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="A6CCEAF8EB66ED4BFC05FA6A674283A7" box="[992,1125,1476,1500]" italics="true" pageId="14" pageNumber="507">C. flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="53B84D69EB66ED4BFF9EFA4A63D18387" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[123,246,1508,1532]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB66ED4BFF9EFA4A63D18387" box="[123,246,1508,1532]" italics="true" pageId="14" pageNumber="507">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. He added that “
|
|||
|
|
<location id="91676031EB66ED4BFE4DFA4B62E08387" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB66ED4BFE4DFA4B62E08387" box="[424,455,1508,1532]" country="Hong Kong" county="Andrewes" municipality="Andrewes'" name="As" pageId="14" pageNumber="507">As</location>
|
|||
|
|
a result of this I think that the
|
|||
|
|
<collectingCountry id="ECAF767AEB66ED4BFCE9FA4A60AE8386" box="[780,905,1509,1533]" name="Hong Kong" pageId="14" pageNumber="507">Hong Kong</collectingCountry>
|
|||
|
|
examples form a distinct species.”
|
|||
|
|
<location id="91676031EB66ED4BFF05F9AA62248067" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB66ED4BFF05F9AA62248067" box="[224,259,1541,1564]" country="Hong Kong" county="Andrewes" municipality="Andrewes'" name="We" pageId="14" pageNumber="507">We</location>
|
|||
|
|
examined
|
|||
|
|
<specimenCount id="82BEFD63EB66ED4BFE93F9AB61068067" box="[374,545,1540,1564]" count="3" pageId="14" pageNumber="507" type="generic">three specimens</specimenCount>
|
|||
|
|
from “
|
|||
|
|
<collectingCountry id="ECAF767AEB66ED4BFD8EF9AB61CA8067" box="[619,749,1540,1564]" name="Hong Kong" pageId="14" pageNumber="507">Hong Kong</collectingCountry>
|
|||
|
|
”, one labeled “Type” (i.e., the specimen we list above as
|
|||
|
|
<typeStatus id="4B038848EB66ED4BFED8F98A62BB8046" box="[317,412,1573,1597]" pageId="14" pageNumber="507" type="holotype">holotype</typeStatus>
|
|||
|
|
) and the other two labeled “Cotype”,
|
|||
|
|
<specimenCount id="82BEFD63EB66ED4BFCB4F98A60D48046" box="[849,1011,1573,1597]" count="2" pageId="14" pageNumber="507" type="generic">two specimens</specimenCount>
|
|||
|
|
from “
|
|||
|
|
<collectingCountry id="ECAF767AEB66ED4BFBADF98A67AB8046" box="[1096,1164,1573,1597]" name="China" pageId="14" pageNumber="507">China</collectingCountry>
|
|||
|
|
” labeled “Cotype,” as well as single specimens from
|
|||
|
|
<location id="91676031EB66ED4BFD59F9EA601C8026" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB66ED4BFD59F9EA601C8026" box="[700,827,1605,1629]" country="Hong Kong" county="Andrewes" municipality="Andrewes'" name="Chandipore" pageId="14" pageNumber="507">Chandipore</location>
|
|||
|
|
, (
|
|||
|
|
<collectingCountry id="ECAF767AEB66ED4BFCB5F9EA60AD8026" box="[848,906,1605,1629]" name="India" pageId="14" pageNumber="507">India</collectingCountry>
|
|||
|
|
) and
|
|||
|
|
<location id="91676031EB66ED4BFC2FF9E967608026" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB66ED4BFC2FF9E967608026" box="[970,1095,1606,1629]" country="Hong Kong" county="Andrewes" municipality="Andrewes'" name="Dom Toum" pageId="14" pageNumber="507">Dom Toum</location>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="ECAF767AEB66ED4BFBBDF9EA67A88026" box="[1112,1167,1605,1629]" name="Laos" pageId="14" pageNumber="507">Laos</collectingCountry>
|
|||
|
|
) labeled “Cotype”.
|
|||
|
|
<location id="91676031EB66ED4BFEDAF9CA62468006" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB66ED4BFEDAF9CA62468006" box="[319,353,1637,1661]" country="Hong Kong" county="Andrewes" municipality="Andrewes'" name="All" pageId="14" pageNumber="507">All</location>
|
|||
|
|
of these cotypes were mentioned in
|
|||
|
|
<collectingMunicipality id="7463AC90EB66ED4BFD01F9CA60708006" box="[740,855,1637,1661]" pageId="14" pageNumber="507">Andrewes’</collectingMunicipality>
|
|||
|
|
description as conspecific, so each of these, as well as any
|
|||
|
|
<typeStatus id="4B038848EB66ED4BFE49F92A617F80E6" box="[428,600,1669,1693]" isOtherTypeStatus="true" pageId="14" pageNumber="507">other specimens</typeStatus>
|
|||
|
|
labeled “Cotype” by
|
|||
|
|
<collectingCounty id="7D664E66EB66ED4BFCD9F92A608C80E6" box="[828,939,1669,1693]" pageId="14" pageNumber="507">Andrewes</collectingCounty>
|
|||
|
|
, should be considered as
|
|||
|
|
<typeStatus id="4B038848EB66ED4BFF7DF90963D980C5" box="[152,254,1702,1726]" pageId="14" pageNumber="507" type="paratype">paratypes</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="53B84D69EB66ED4BFEC6F90962A880C6" box="[291,399,1702,1725]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB66ED4BFEC6F90962A880C6" box="[291,399,1702,1725]" italics="true" pageId="14" pageNumber="507">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB67ED57FF4EFF73611A8501" lastPageId="18" lastPageNumber="511" pageId="15" pageNumber="508" type="diagnosis">
|
|||
|
|
<paragraph id="940736EAEB67ED4AFF4EFF73606B854D" blockId="15.[123,1174,219,1725]" pageId="15" pageNumber="508">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFF4EFF7362388688" bold="true" box="[171,287,220,243]" pageId="15" pageNumber="508">Diagnosis.</emphasis>
|
|||
|
|
Adults of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFE78FF74612A8688" box="[413,525,219,243]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFE78FF74612A8688" box="[413,525,219,243]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be distinguished from those of all other
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFBF1FF7463EF8768" pageId="15" pageNumber="508">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFBF1FF7467B28688" box="[1044,1173,219,243]" italics="true" pageId="15" pageNumber="508">Cyclosomus</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
in Asia by the following combination of character states: Body size medium for genus, BL males = 7.3 to 8.0 mm, females =
|
|||
|
|
<quantity id="53409B0FEB67ED4AFE0DFEB461A68748" box="[488,641,283,307]" metricMagnitude="-3" metricUnit="m" metricValue="7.199999999999999" metricValueMax="7.8" metricValueMin="6.6" pageId="15" pageNumber="508" unit="mm" value="7.199999999999999" valueMax="7.8" valueMin="6.6">6.6 to 7.8 mm</quantity>
|
|||
|
|
; body form (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFCEEFEB4608E8748" box="[779,937,283,307]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="15" pageNumber="508">Figs. 7A, C, D</figureCitation>
|
|||
|
|
) more elongate-ovoid (ratio BL/EW = 1.47 to 1.53), with elytra widest at or slightly anterior to mid-length; pronotum (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFF66FEF363C1870F" box="[131,230,348,372]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="15" pageNumber="508">Fig. 14C</figureCitation>
|
|||
|
|
) relatively longer and narrower (ratio PWM/PL = 2.33 to 2.93) and more broadened basally (ratio PWM/PWA = 1.63 to 1.75), with disc varied in color (rufous to piceous) and lateral pale bands of medium width and well-defined, anterior angles broad, lateral margins not sinuate near anterior angles; free apex of prosternal intercoxal process long (as in
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFC4FFE13672387AF" box="[938,1028,444,468]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="15" pageNumber="508">Fig.15C</figureCitation>
|
|||
|
|
); elytral dark pattern varied in both form and color, with dark maculae rufous to piceous in color and with basal and middle bands of average (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFE26FE526130846E" box="[451,535,509,533]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="15" pageNumber="508">Fig. 7D</figureCitation>
|
|||
|
|
) to less than average (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFCE8FE526045846E" box="[781,866,509,533]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="15" pageNumber="508">Fig. 7A</figureCitation>
|
|||
|
|
) thickness and continuity or middle band absent (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFEBFFDB26289844E" box="[346,430,541,565]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="15" pageNumber="508">Fig. 7C</figureCitation>
|
|||
|
|
), preapical dark spot absent or only faintly present (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFC39FDB26708844E" box="[988,1071,541,565]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="15" pageNumber="508">Fig. 7D</figureCitation>
|
|||
|
|
) in a few specimens (from inland areas in Indochina); elytral striae very deeply impressed, elytral intervals (especially intervals 3 to 5) slightly but distinctly convex; elytral epipleura with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; median lobe of male genitalia long and more slender, with shaft of approximately equal thickness throughout, ventral curvature gradually and only slightly arcuate in apical two-thirds in lateral aspect (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFC13FD11677F84AD" box="[1014,1112,702,726]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="15" pageNumber="508">Fig. 16C</figureCitation>
|
|||
|
|
), apical lamella longer, smoothly rounded apically, and not or only slightly narrowed basally in dorsal aspect (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFF29FD516209856D" box="[204,302,766,790]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="15" pageNumber="508">Fig. 17C</figureCitation>
|
|||
|
|
); specimen from southern Asia, from northeastern coastal
|
|||
|
|
<collectingCountry id="ECAF767AEB67ED4AFC78FD5160F1856D" box="[925,982,766,790]" name="India" pageId="15" pageNumber="508">India</collectingCountry>
|
|||
|
|
, through Indochina, to coastal southeastern and eastern
|
|||
|
|
<collectingCountry id="ECAF767AEB67ED4AFDF8FCB1617A854D" box="[541,605,798,822]" name="China" pageId="15" pageNumber="508">China</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="ECAF767AEB67ED4AFD74FCB161F8854D" box="[657,735,798,822]" name="Taiwan" pageId="15" pageNumber="508">Taiwan</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFD0BFCB16018854D" box="[750,831,798,822]" captionStart="FIGURE" captionStartId="39.[171,183,837,856]" captionTargetBox="[168,1128,216,825]" captionTargetId="figure-14@39.[168,1128,216,825]" captionTargetPageId="39" captionText="FIGURE 20. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow circles = C. acutangulus Kavanaugh and Cueva-Dabkoski; white triangles = C. flexuosus Dejean; yellow triangle = locality where C. acutangulus and C. flexuosus have been found together; white squares = C. inustus Andrewes. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/ Whole_world_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513000" httpUri="https://zenodo.org/record/11513000/files/figure.png" pageId="15" pageNumber="508">Fig. 20</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB67ED4AFF4EFCEF62A28243" blockId="15.[123,1174,219,1725]" pageId="15" pageNumber="508">
|
|||
|
|
In his original description of this species,
|
|||
|
|
<bibRefCitation id="F0294B1BEB67ED4AFD80FC906038852C" author="ANDREWES, H. E." box="[613,799,831,855]" pageId="15" pageNumber="508" pagination="459 - 468" refId="ref23019" refString="ANDREWES, H. E. 1924. XXII. On the Oriental Carabidae of the " Reise Novara. " Transactions of the Entomological Society of London 1923: 459 - 468." type="journal article" year="1924">Andrewes (1924)</bibRefCitation>
|
|||
|
|
included specimens demonstrating variation among populations in different parts of the range of the species. He noted that a specimen from Pak Lay (
|
|||
|
|
<collectingCountry id="ECAF767AEB67ED4AFEC7FCD0627E85EC" box="[290,345,895,919]" name="Laos" pageId="15" pageNumber="508">Laos</collectingCountry>
|
|||
|
|
) was exceptionally dark and convex and that specimens from Nanoa Island (
|
|||
|
|
<collectingRegion id="567CF808EB67ED4AFF67FC30624D85CC" box="[130,362,927,951]" country="China" name="Guangdong" pageId="15" pageNumber="508">Guangdong Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="ECAF767AEB67ED4AFE90FC30629185CC" box="[373,438,927,951]" name="China" pageId="15" pageNumber="508">China</collectingCountry>
|
|||
|
|
) were exceptionally small and pale. Our examination of genitalia of males from these areas and the others listed below found that they all have similar overall form but that the shape and length of the apical lamella is slightly varied both within and among populations. Nonetheless, Andrewes concluded that these diverse populations were all conspecific and we tentatively follow him here.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB67ED4AFF4EFBEF625783E1" blockId="15.[123,1174,219,1725]" pageId="15" pageNumber="508">
|
|||
|
|
Specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFEACFBEE629F8223" box="[329,440,1089,1112]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFEACFBEE629F8223" box="[329,440,1089,1112]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
with the elytral preapical pale spot absent are similar to those of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFF9EFBCE62328202" box="[123,277,1121,1145]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFF9EFBCE62328202" box="[123,277,1121,1145]" italics="true" pageId="15" pageNumber="508">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFEC6FBCE62BD8202" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[291,410,1121,1145]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="pallidus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFEC6FBCE62BD8202" box="[291,410,1121,1145]" italics="true" pageId="15" pageNumber="508">C. pallidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFE3DFBCE614D8202" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[472,618,1121,1145]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="vespertilio">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFE3DFBCE614D8202" box="[472,618,1121,1145]" italics="true" pageId="15" pageNumber="508">C. vespertilio</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. They are distinguished from
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFC51FBCE670C8202" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[948,1067,1121,1145]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="pallidus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFC51FBCE670C8202" box="[948,1067,1121,1145]" italics="true" pageId="15" pageNumber="508">C. pallidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
members by their more elongate body form and somewhat narrower pronotum (compare
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFC1CFB2E677C82E2" box="[1017,1115,1153,1177]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="15" pageNumber="508">Fig. 14C</figureCitation>
|
|||
|
|
with
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFF9EFB0E63FF82C2" box="[123,216,1185,1209]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="15" pageNumber="508">Fig. 14E</figureCitation>
|
|||
|
|
) with lateral pale bands much narrower and lateral areas less broadly flattened. Most (but not all) of these
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFECCFB6D62B382A2" box="[297,404,1218,1241]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFECCFB6D62B382A2" box="[297,404,1218,1241]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
specimens have the middle dark band, or at least a vestige of it, evident on the elytra, whereas
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFE99FB4D62D28281" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[380,501,1250,1274]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="pallidus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFE99FB4D62D28281" box="[380,501,1250,1274]" italics="true" pageId="15" pageNumber="508">C. pallidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
specimens lack this band altogether (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFC7AFB4D60DE8281" box="[927,1017,1250,1274]" captionStart="FIGURE" captionStartId="21.[171,183,1155,1174]" captionTargetBox="[120,1176,216,1138]" captionTargetId="figure-233@21.[120,1176,216,1147]" captionTargetPageId="21" captionText="FIGURE 9. Cyclosomus pallidus Kavanaugh and Cueva-Dabkoski sp. nov. A. Holotype female, dorsal habitus; B. Labels associated with holotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512976" httpUri="https://zenodo.org/record/11512976/files/figure.png" pageId="15" pageNumber="508">Fig. 9A</figureCitation>
|
|||
|
|
). Members of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFF9EFAAD63CE8361" box="[123,233,1282,1306]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFF9EFAAD63CE8361" box="[123,233,1282,1306]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be distinguished from those of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFD99FAAD60288361" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[636,783,1282,1306]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="vespertilio">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFD99FAAD60288361" box="[636,783,1282,1306]" italics="true" pageId="15" pageNumber="508">C. vespertilio</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by their more elongate body form, slightly broader basal dark band on the elytra, and narrower reflexed lateral elytral margins. Males of these two species also differ distinctly in the shape of the median lobe of the genitalia, with the shaft much shorter and thicker in
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFE11FACC61A08301" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[500,647,1378,1402]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="vespertilio">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFE11FACC61A08301" box="[500,647,1378,1402]" italics="true" pageId="15" pageNumber="508">C. vespertilio</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
males (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFD05FACD60638301" box="[736,836,1378,1402]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="15" pageNumber="508">Fig. 16H</figureCitation>
|
|||
|
|
) than in
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFC49FACC673D8301" box="[940,1050,1379,1402]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFC49FACC673D8301" box="[940,1050,1379,1402]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFBCEFACD67A98301" box="[1067,1166,1378,1402]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="15" pageNumber="508">Fig. 16C</figureCitation>
|
|||
|
|
) males in lateral aspect.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB67ED54FF4EFA0C639587AF" blockId="15.[123,1174,219,1725]" lastBlockId="17.[123,1174,219,1725]" lastPageId="17" lastPageNumber="510" pageId="15" pageNumber="508">
|
|||
|
|
Distinguishing specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFE06FA0B616883C0" box="[483,591,1444,1467]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFE06FA0B616883C0" box="[483,591,1444,1467]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from those of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFD0AFA0B60AF83C0" box="[751,904,1444,1467]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFD0AFA0B60AF83C0" box="[751,904,1444,1467]" italics="true" pageId="15" pageNumber="508">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is more difficult, at least partly because there are so few specimens of the latter known (just three) that it remains difficult to fully characterize its members. As presently conceived, members of the two species differ slightly in body shape (slightly more elongate-ovoid in
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFD6BF9AB61DF8060" box="[654,760,1540,1563]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFD6BF9AB61DF8060" box="[654,760,1540,1563]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), pronotal shape (relatively longer and narrower and with narrower lateral pale bands in
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFD63F98B61D68047" box="[646,753,1572,1596]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFD63F98B61D68047" box="[646,753,1572,1596]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; compare
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFC84F98B60998047" box="[865,958,1572,1596]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="15" pageNumber="508">Fig. 14C</figureCitation>
|
|||
|
|
with
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFC1FF98B677C8047" box="[1018,1115,1572,1596]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="15" pageNumber="508">Fig. 14D</figureCitation>
|
|||
|
|
), and length of the free apex of the prosternal intercoxal process (longer in
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFC93F9EA60FF8027" authorityName="Andrewes" authorityYear="1924" box="[886,984,1605,1628]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustu">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFC93F9EA60FF8027" box="[886,984,1605,1628]" italics="true" pageId="15" pageNumber="508">C. inustu</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
s, as in
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFBCAF9EB67B68027" box="[1071,1169,1604,1628]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="15" pageNumber="508">Fig. 15C</figureCitation>
|
|||
|
|
; medium length in
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFEADF9CA62C38007" box="[328,484,1637,1660]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFEADF9CA62C38007" box="[328,484,1637,1660]" italics="true" pageId="15" pageNumber="508">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, as in
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFDD6F9CB61BF8007" box="[563,664,1636,1660]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="15" pageNumber="508">Fig. 15B</figureCitation>
|
|||
|
|
). Because the
|
|||
|
|
<typeStatus id="4B038848EB67ED4AFCD9F9CB60B98007" box="[828,926,1636,1660]" pageId="15" pageNumber="508" type="lectotype">lectotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFC2DF9CA67438007" box="[968,1124,1637,1660]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFC2DF9CA67438007" box="[968,1124,1637,1660]" italics="true" pageId="15" pageNumber="508">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is a female, we cannot be certain that the male specimen we tentatively identify as that species is conspecific with the type. Genitalia of that male (
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFD95F90B61F680C7" box="[624,721,1700,1724]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="15" pageNumber="508">Fig. 16D</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="0C832A6FEB67ED4AFCEDF90B600680C7" box="[776,801,1700,1724]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="15" pageNumber="508">17</figureCitation>
|
|||
|
|
FD) and those of
|
|||
|
|
<taxonomicName id="53B84D69EB67ED4AFC06F90A677780C6" box="[995,1104,1701,1725]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="508" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB67ED4AFC06F90A677780C6" box="[995,1104,1701,1725]" italics="true" pageId="15" pageNumber="508">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="0C832A6FEB67ED54FB85F90B63808688" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" lastPageId="17" lastPageNumber="510" pageId="15" pageNumber="508">Figs. 16C</figureCitation>
|
|||
|
|
and
|
|||
|
|
<figureCitation id="0C832A6FEB79ED54FF05FF74622A8688" box="[224,269,219,243]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="17" pageNumber="510">17C</figureCitation>
|
|||
|
|
) are similar in form but differ slightly in the shape of the apical lamella in dorsal aspect (compare
|
|||
|
|
<figureCitation id="0C832A6FEB79ED54FED5FF5462D98768" box="[304,510,251,275]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="17" pageNumber="510">Figs. 17C and 17D</figureCitation>
|
|||
|
|
). However, the shape of the apical lamella in one of
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FBD6FF5367B18768" box="[1075,1174,251,275]" count="2" pageId="17" pageNumber="510" type="male">two male</specimenCount>
|
|||
|
|
specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FEF5FEB3625A8748" box="[272,381,284,307]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FEF5FEB3625A8748" box="[272,381,284,307]" italics="true" pageId="17" pageNumber="510">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FE25FEB461098748" box="[448,558,283,307]" name="Cambodia" pageId="17" pageNumber="510">Cambodia</collectingCountry>
|
|||
|
|
is similar to that of the male we identify as
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FBF6FEB363BD8728" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FBF6FEB363BD8728" italics="true" pageId="17" pageNumber="510">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, although the two differ in all the external features noted above to distinguish these two species. We certainly need to consider that
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FE09FEF36170870F" box="[492,599,348,372]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FE09FEF36170870F" box="[492,599,348,372]" italics="true" pageId="17" pageNumber="510">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FD6FFEF36005870F" box="[650,802,348,372]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FD6FFEF36005870F" box="[650,802,348,372]" italics="true" pageId="17" pageNumber="510">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
may, in fact, be conspecific. However, the paucity of material referable to the latter and the uncertainty (see below) about its actual geographical and habitat distributions renders a taxonomic decision in this regard premature at this time.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="C0C76662EB78ED55FF4EF98463848009" ID-DOI="http://doi.org/10.5281/zenodo.11512970" ID-Zenodo-Dep="11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="16" pageNumber="509" startId="16.[171,183,1579,1598]" targetBox="[182,1114,216,1560]" targetPageId="16" targetType="figure">
|
|||
|
|
<paragraph id="940736EAEB78ED55FF4EF98463848009" blockId="16.[123,1174,1579,1650]" pageId="16" pageNumber="509">
|
|||
|
|
<smallCapsWord id="92E1A036EB78ED55FF4EF98463CF8046" baselines="1593,1593" box="[171,232,1579,1598]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="16" pageNumber="509">FIGURE</smallCapsWord>
|
|||
|
|
7.
|
|||
|
|
<taxonomicName id="53B84D69EB78ED55FEE0F984610A8045" authority="Andrewes. A." authorityName="Andrewes. A." box="[261,557,1579,1598]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="509" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB78ED55FEE0F98462888045" box="[261,431,1579,1598]" italics="true" pageId="16" pageNumber="509">Cyclosomus inustus</emphasis>
|
|||
|
|
Andrewes. A.
|
|||
|
|
</taxonomicName>
|
|||
|
|
Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="940736EAEB79ED54FF4EFE7362FC84AD" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="510">
|
|||
|
|
Those few atypical specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FDF4FE72615B878F" box="[529,636,477,500]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FDF4FE72615B878F" box="[529,636,477,500]" italics="true" pageId="17" pageNumber="510">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
that we have seen with a faint preapical dark spot on the elytra (
|
|||
|
|
<figureCitation id="0C832A6FEB79ED54FEEBFE526246846E" box="[270,353,509,533]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="17" pageNumber="510">Fig. 7D</figureCitation>
|
|||
|
|
) (see section on Geographical Variation below) are externally very similar to members of
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FEE1FDB162A4844E" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[260,387,541,565]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FEE1FDB162A4844E" box="[260,387,541,565]" italics="true" pageId="17" pageNumber="510">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and reliably distinguished from the latter only by reference to the male genitalia. The apical lamella of such
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FDEBFD91615D842E" box="[526,634,574,597]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FDEBFD91615D842E" box="[526,634,574,597]" italics="true" pageId="17" pageNumber="510">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
males is rounded and slightly (as in
|
|||
|
|
<figureCitation id="0C832A6FEB79ED54FBEAFD926757842E" box="[1039,1136,573,597]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="17" pageNumber="510">Fig. 17D</figureCitation>
|
|||
|
|
) to moderately elongate (
|
|||
|
|
<figureCitation id="0C832A6FEB79ED54FE80FDF262E0840E" box="[357,455,605,629]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="17" pageNumber="510">Fig. 17C</figureCitation>
|
|||
|
|
), whereas that of
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FD6DFDF16021840E" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[648,774,605,629]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FD6DFDF16021840E" box="[648,774,605,629]" italics="true" pageId="17" pageNumber="510">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
males is shorter and bluntly triangular (
|
|||
|
|
<figureCitation id="0C832A6FEB79ED54FF42FDD1622B84ED" box="[167,268,638,662]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="17" pageNumber="510">Fig. 17G</figureCitation>
|
|||
|
|
). It is certainly possible that these specimens with preapical pale spots from inland localities in Indochina, which we identify here as
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FD6AFD3161DD84CD" box="[655,762,670,694]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FD6AFD3161DD84CD" box="[655,762,670,694]" italics="true" pageId="17" pageNumber="510">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, actually represent a separate, perhaps undescribed species (see below).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB79ED54FF4EFD7063D982C2" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="510">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FF4EFD7062BC848D" bold="true" box="[171,411,735,758]" pageId="17" pageNumber="510">Habitat distribution.</emphasis>
|
|||
|
|
<bibRefCitation id="F0294B1BEB79ED54FE40FD716148848D" author="WATERHOUSE, J. O." box="[421,623,734,758]" pageId="17" pageNumber="510" pagination="102 - 107" refId="ref24081" refString="WATERHOUSE, J. O. 1850 - 51. Proceeding of the Entomological Society of London, September 1, 1851. Transactions of the Entomological Society of London (New Series) 1: 102 - 107." type="journal article" year="1850">Waterhouse (1850</bibRefCitation>
|
|||
|
|
-51:104) quoted text from a letter written by J.C. Bowring and dated
|
|||
|
|
<date id="E006102AEB79ED54FEA9FD5062F4856D" box="[332,467,766,790]" pageId="17" pageNumber="510" value="1851-06-09">June 9, 1851</date>
|
|||
|
|
describing his encounter with this species in
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FC53FD506711856C" box="[950,1078,767,791]" name="Hong Kong" pageId="17" pageNumber="510">Hong Kong</collectingCountry>
|
|||
|
|
, the
|
|||
|
|
<typeStatus id="4B038848EB79ED54FB82FD5067B2856C" box="[1127,1173,767,791]" pageId="17" pageNumber="510">type</typeStatus>
|
|||
|
|
locality, as follows: “I inclose [sic] a pair of
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FD96FCB0608D854C" authority=", White" authorityName="White" box="[627,938,799,823]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="insularis">
|
|||
|
|
Cyclosomus insularis,
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FC89FCB0608D854C" box="[876,938,799,823]" italics="true" pageId="17" pageNumber="510">White</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
[a nomen nudum], a species I met yesterday morning for the first time. This beetle burrows to some depth in the sand by the seashore; it is very active in its movements, and when exposed on the surface disappears beneath the sand with truly wonderful rapidity, diving down head foremost. I captured about
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FBBCFC2F622285CC" count="20" pageId="17" pageNumber="510" type="generic">twenty specimens</specimenCount>
|
|||
|
|
by turning up the sand for some distance to a depth of five or six inches”.
|
|||
|
|
<bibRefCitation id="F0294B1BEB79ED54FBC6FC0F63E685A3" author="WANG, L. J. & J. J. CHERNG & D. H. KAVANAUGH" pageId="17" pageNumber="510" pagination="107 - 116" refId="ref23969" refString="WANG, L. J., J. J. CHERNG, AND D. H. KAVANAUGH. 2017. First records for Cyclosomus inustus Andrewes (Coleoptera: Carabidae: Cyclosomini) for Taiwan, with notes on habitat and behavior. Proceedings of the California Academy of Sciences (Series 4), 64: 107 - 116." type="journal article" year="2017">Wang et al (2017)</bibRefCitation>
|
|||
|
|
reported on the discovery of
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FDE7FC6F614985A3" box="[514,622,960,984]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FDE7FC6F614985A3" box="[514,622,960,984]" italics="true" pageId="17" pageNumber="510">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in coastal sun dune areas (
|
|||
|
|
<figureCitation id="0C832A6FEB79ED54FC77FC6F60C785A3" box="[914,992,960,984]" captionStart="FIGURE" captionStartId="38.[155,167,1584,1603]" captionTargetBox="[120,1176,216,1564]" captionTargetId="figure-9@38.[120,1176,216,1564]" captionTargetPageId="38" captionText="FIGURE 19. Photographs of habitats for Cyclosomus inustus Andrewes. A. Lingshanbi, Shihmen District, New Taipei City, Taiwan; B. Shalun, Danshui District, New Taipei City, Taiwan. Photographs reproduced from Wang et al. 2017 with permission of Liang-Jong Wang." figureDoi="http://doi.org/10.5281/zenodo.11512998" httpUri="https://zenodo.org/record/11512998/files/figure.png" pageId="17" pageNumber="510">Fig. 19</figureCitation>
|
|||
|
|
) in northern and northeastern
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FEE3FC4F62708583" box="[262,343,992,1016]" name="Taiwan" pageId="17" pageNumber="510">Taiwan</collectingCountry>
|
|||
|
|
, and
|
|||
|
|
<bibRefCitation id="F0294B1BEB79ED54FE75FC4E617E8583" author="WANG, L. J. & J. J. CHERNG & U. ONG" box="[400,601,992,1017]" pageId="17" pageNumber="510" pagination="112 - 115" refId="ref24032" refString="WANG, L. J., J. J. CHERNG, AND U. ONG. 2022. New records of Cyclosomus inustus Andrewes (Coleoptera: Carabidae: Cyclosomini) in Taiwan and Kinmen. Japanese Journal of Systematic Entomology 28: 112 - 115." type="journal article" year="2022">Wang et al. (2022)</bibRefCitation>
|
|||
|
|
found addition coastal dune localities in
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FBFCFC4F67408583" box="[1049,1127,992,1016]" name="Taiwan" pageId="17" pageNumber="510">Taiwan</collectingCountry>
|
|||
|
|
and on
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FF78FBAF63D48263" box="[157,243,1024,1048]" country="Taiwan" name="Kinmen" pageId="17" pageNumber="510">Kinmen</collectingRegion>
|
|||
|
|
Island just off the Chinese mainland. They also searched extensively for these beetles in similarly exposed sandy habitats along river and lake shores in both areas but without success. Additional records (see below) from Fuzhou and Ningbo (
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FD01FBEE600E8222" box="[740,809,1089,1113]" name="China" pageId="17" pageNumber="510">China</collectingCountry>
|
|||
|
|
), and probably also from Ban Hua Hin (
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FF51FBCE623F8202" box="[180,280,1121,1145]" name="Thailand" pageId="17" pageNumber="510">Thailand</collectingCountry>
|
|||
|
|
), and Chandipur (
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FE3AFBCE613E8202" box="[479,537,1121,1145]" name="India" pageId="17" pageNumber="510">India</collectingCountry>
|
|||
|
|
) likely also represent coastal sandy beach or dune occurrence. At least in
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FEA5FB2E62A982E2" box="[320,398,1153,1177]" name="Taiwan" pageId="17" pageNumber="510">Taiwan</collectingCountry>
|
|||
|
|
and on
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FE09FB2E616582E2" box="[492,578,1153,1177]" country="Taiwan" name="Kinmen" pageId="17" pageNumber="510">Kinmen</collectingRegion>
|
|||
|
|
Island, this species appears to be restricted to coastal sandy areas.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB79ED54FF4EFB6E60A28341" blockId="17.[123,1174,219,1725]" pageId="17" pageNumber="510">
|
|||
|
|
The habitats for specimens collected at inland localities in Indochina (i.e., in
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FC05FB6E677782A2" box="[992,1104,1217,1241]" name="Cambodia" pageId="17" pageNumber="510">Cambodia</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FBBEFB6D67B682A2" box="[1115,1169,1218,1241]" name="Laos" pageId="17" pageNumber="510">Laos</collectingCountry>
|
|||
|
|
, and central
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FF1DFB4D627E8281" box="[248,345,1250,1274]" name="Thailand" pageId="17" pageNumber="510">Thailand</collectingCountry>
|
|||
|
|
) are unknown but likely were along streams with exposed sandy banks. This may reflect either a broader habitat range for
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FD65FAAC61D78361" box="[640,752,1283,1306]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FD65FAAC61D78361" box="[640,752,1283,1306]" italics="true" pageId="17" pageNumber="510">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in this region or a species difference between these inland populations and the coastal
|
|||
|
|
<taxonomicName id="53B84D69EB79ED54FD6BFA8C61DD8341" box="[654,762,1315,1338]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="510" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FD6BFA8C61DD8341" box="[654,762,1315,1338]" italics="true" pageId="17" pageNumber="510">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
populations.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB79ED57FF4EFAEC60578748" blockId="17.[123,1174,219,1725]" lastBlockId="18.[123,1174,219,890]" lastPageId="18" lastPageNumber="511" pageId="17" pageNumber="510">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FF4EFAEC62FA8321" bold="true" box="[171,477,1347,1370]" pageId="17" pageNumber="510">Geographical distribution.</emphasis>
|
|||
|
|
We have examined a total of
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FCD1FAED60EF8321" box="[820,968,1346,1370]" count="41" pageId="17" pageNumber="510" type="generic">41 specimens</specimenCount>
|
|||
|
|
(
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FC3EFAED67188321" box="[987,1087,1346,1370]" count="23" pageId="17" pageNumber="510" type="male">23 males</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FB9EFAED63F78300" count="18" pageId="17" pageNumber="510" type="female">18 females</specimenCount>
|
|||
|
|
)
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FF07FACC615483E0" ID-GBIF-Occurrence="4905152360" collectingDate="2012-05-07" collectingDateMax="2012-05-10" collectingDateMin="2012-05-07" collectionCode="NHMUK" collectorName="J. K. Li" country="Cambodia" location="Steong Treng" pageId="17" pageNumber="510" specimenCount="2" specimenCount-male="2">
|
|||
|
|
from the following localities:
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FDD1FACC61EC8301" bold="true" box="[564,715,1379,1402]" pageId="17" pageNumber="510">
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FDD1FACC61EC8301" box="[564,715,1379,1402]" name="Cambodia" pageId="17" pageNumber="510">CAMBODIA</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
:
|
|||
|
|
<location id="91676031EB79ED54FD3BFACC60538300" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FD3BFACC60538300" box="[734,884,1379,1403]" country="Cambodia" name="Steong Treng" pageId="17" pageNumber="510">Steong Treng</location>
|
|||
|
|
, “Hat Baoho” (
|
|||
|
|
<date id="E006102AEB79ED54FBC2FACC639483E0" pageId="17" pageNumber="510" value="2012-05-07" valueMax="2012-05-10" valueMin="2012-05-07">
|
|||
|
|
<collectingDate id="F042E9C2EB79ED54FBC2FACC639483E0" pageId="17" pageNumber="510" value="2012-05-07" valueMax="2012-05-10" valueMin="2012-05-07">7-10 May 2012</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="394D533CEB79ED54FF5BFA2B622F83E0" box="[190,264,1411,1435]" pageId="17" pageNumber="510">J.K. Li</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FE9EFA2B62C983E0" box="[379,494,1411,1435]" count="2" pageId="17" pageNumber="510" type="male">two males</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FE1CFA2C614283E0" box="[505,613,1411,1435]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FD9BFA2B614883C0" ID-GBIF-Occurrence="4905152333" collectionCode="NHMUK, NHRS, IRSNB" country="China" location="CHINA" pageId="17" pageNumber="510" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FD9BFA2B61F083E0" bold="true" box="[638,727,1412,1435]" pageId="17" pageNumber="510">
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FD9BFA2B61F083E0" box="[638,727,1412,1435]" name="China" pageId="17" pageNumber="510">CHINA</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
([
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FD08FA2C607583E0" box="[749,850,1411,1435]" count="1" pageId="17" pageNumber="510" type="male">one male</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FC62FA2B672B83E0" box="[903,1036,1411,1435]" count="2" pageId="17" pageNumber="510" type="female">two females</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FBFDFA2C67A383E0" box="[1048,1156,1411,1435]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
]), ([
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FF6CFA0C63D983C0" box="[137,254,1443,1467]" count="2" pageId="17" pageNumber="510" type="male">two males</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FEEFFA0C627083C0" box="[266,343,1443,1467]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="17" pageNumber="510">NHRS</collectionCode>
|
|||
|
|
]), ([
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FE65FA0C62D983C0" box="[384,510,1443,1467]" count="1" pageId="17" pageNumber="510" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FDEFFA0C614783C0" box="[522,608,1443,1467]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="17" pageNumber="510">IRSNB</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
:
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FD9FFA0C62F983A0" ID-GBIF-Occurrence="4905152310" collectingDate="1935-05-31" collectionCode="NHMUK" collectorName="M. S. Yang" country="China" location="Fuzhou" pageId="17" pageNumber="510" specimenCount="3" specimenCount-female="3" stateProvince="Fujian">
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FD9FFA0C61E583C0" box="[634,706,1443,1467]" country="China" name="Fujian" pageId="17" pageNumber="510">Fujian</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<location id="91676031EB79ED54FD28FA0C603A83C0" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FD28FA0C603A83C0" box="[717,797,1443,1467]" country="China" name="Fuzhou" pageId="17" pageNumber="510" stateProvince="Fujian">Fuzhou</location>
|
|||
|
|
(
|
|||
|
|
<date id="E006102AEB79ED54FCCBFA0C60E583C0" box="[814,962,1443,1468]" pageId="17" pageNumber="510" value="1935-05-31">
|
|||
|
|
<collectingDate id="F042E9C2EB79ED54FCCBFA0C60E583C0" box="[814,962,1443,1468]" pageId="17" pageNumber="510" value="1935-05-31">31 May 1935</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
-36,
|
|||
|
|
<collectorName id="394D533CEB79ED54FC17FA0C674383C7" box="[1010,1124,1443,1468]" pageId="17" pageNumber="510">M.S. Yang</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FF21FA6C627E83A0" box="[196,345,1475,1499]" count="3" pageId="17" pageNumber="510" type="female">three females</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FE80FA6C62F683A0" box="[357,465,1475,1499]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FE0FFA6C625F8380" ID-GBIF-Occurrence="4905152345" collectionCode="NHMUK" collectorName="J. J. Walker" country="China" location="Nanoa Island" pageId="17" pageNumber="510" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Guangdong">
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FE0FFA6C614C83A0" box="[490,619,1475,1499]" country="China" name="Guangdong" pageId="17" pageNumber="510">Guangdong</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<location id="91676031EB79ED54FD91FA6B602583A0" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FD91FA6B602583A0" box="[628,770,1475,1499]" country="China" name="Nanoa Island" pageId="17" pageNumber="510" stateProvince="Guangdong">Nanoa Island</location>
|
|||
|
|
(
|
|||
|
|
<collectorName id="394D533CEB79ED54FCF7FA6C60A183A0" box="[786,902,1475,1499]" pageId="17" pageNumber="510">J.J. Walker</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FC1DFA6B674F83A0" box="[1016,1128,1475,1499]" count="2" pageId="17" pageNumber="510" type="male">two males</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FF9EFA4B63D38380" box="[123,244,1507,1531]" count="1" pageId="17" pageNumber="510" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FF1AFA4C624C8380" box="[255,363,1507,1531]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FE61FA4B60E78380" ID-GBIF-Occurrence="4905152342" box="[388,960,1507,1532]" collectionCode="NHMUK" country="China" location="Hong Kong" pageId="17" pageNumber="510" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Guangdong">
|
|||
|
|
<location id="91676031EB79ED54FE61FA4B61228387" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FE61FA4B61228387" box="[388,517,1508,1532]" country="China" name="Hong Kong" pageId="17" pageNumber="510" stateProvince="Guangdong">Hong Kong</location>
|
|||
|
|
: ([
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FDFBFA4C61A98380" box="[542,654,1507,1531]" count="2" pageId="17" pageNumber="510" type="male">two males</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FD26FA4B601B8380" box="[707,828,1507,1531]" count="1" pageId="17" pageNumber="510" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FCA2FA4C60948380" box="[839,947,1507,1531]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FC29FA4C62578067" ID-GBIF-Occurrence="4905152340" collectionCode="MFNB" country="China" location="Ningobo" pageId="17" pageNumber="510" specimenCount="1" specimenCount-female="1" stateProvince="Zhejiang">
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FC29FA4C67098380" box="[972,1070,1507,1531]" country="China" name="Zhejiang" pageId="17" pageNumber="510">Zhejiang</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<location id="91676031EB79ED54FBDDFA4C67B28380" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FBDDFA4C67B28380" box="[1080,1173,1507,1531]" country="China" name="Ningobo" pageId="17" pageNumber="510" stateProvince="Zhejiang">Ningobo</location>
|
|||
|
|
([
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FF6CF9AB62228067" box="[137,261,1540,1564]" count="1" pageId="17" pageNumber="510" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FEEAF9AB62458067" box="[271,354,1540,1564]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="17" pageNumber="510">MFNB</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FE9CF9AB61C18060" ID-GBIF-Occurrence="4905152367" box="[377,742,1540,1564]" collectionCode="NHMUK" country="China" isTailEnumeration="true" location="China" pageId="17" pageNumber="510" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang">
|
|||
|
|
“W
|
|||
|
|
<location id="91676031EB79ED54FE41F9AB62C18067" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FE41F9AB62C18067" box="[420,486,1540,1564]" name="China" pageId="17" pageNumber="510">China</location>
|
|||
|
|
” ([
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FDE6F9AB614D8067" box="[515,618,1540,1564]" count="1" pageId="17" pageNumber="510" type="male">one male</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FD90F9AB61C78060" box="[629,736,1540,1563]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
]
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FD0AF9AB628E8047" ID-GBIF-Occurrence="4905152316" collectionCode="MFNB, NHMUK" country="India" location="INDIA" pageId="17" pageNumber="510" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FD0AF9AB60198060" bold="true" box="[751,830,1540,1563]" pageId="17" pageNumber="510">
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FD0AF9AB60198060" box="[751,830,1540,1563]" name="India" pageId="17" pageNumber="510">INDIA</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FCAFF9AB60888067" box="[842,943,1540,1564]" count="1" pageId="17" pageNumber="510" type="male">one male</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FC5CF9AB672C8067" box="[953,1035,1540,1564]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="17" pageNumber="510">MFNB</collectionCode>
|
|||
|
|
]), ([
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FBD7F9AB67B28067" box="[1074,1173,1540,1564]" count="1" pageId="17" pageNumber="510" type="male">one male</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FF4CF98A62048047" box="[169,291,1572,1596]" count="1" pageId="17" pageNumber="510" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FECAF98B62BB8047" box="[303,412,1572,1596]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
:
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FE51F98B62EE8027" ID-GBIF-Occurrence="4905152303" collectingDate="1915-06-03" collectingDateMax="1915-06-07" collectingDateMin="1915-06-03" collectionCode="NHMUK" collectorName="F. H. Gravely" country="India" location="Balasore" municipality="Chandipur" pageId="17" pageNumber="510" specimenCount="1" specimenCount-female="1" stateProvince="Odisha">
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FE51F98B61238047" box="[436,516,1572,1596]" country="India" name="Odisha" pageId="17" pageNumber="510">Odisha</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<collectingMunicipality id="7463AC90EB79ED54FDEAF98B61A78047" box="[527,640,1572,1596]" pageId="17" pageNumber="510">Chandipur</collectingMunicipality>
|
|||
|
|
(near
|
|||
|
|
<location id="91676031EB79ED54FD23F98B60018047" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FD23F98B60018047" box="[710,806,1572,1596]" country="India" municipality="Chandipur" name="Balasore" pageId="17" pageNumber="510" stateProvince="Odisha">Balasore</location>
|
|||
|
|
,
|
|||
|
|
<date id="E006102AEB79ED54FCD7F98B60E98047" box="[818,974,1572,1596]" pageId="17" pageNumber="510" value="1915-06-03" valueMax="1915-06-07" valueMin="1915-06-03">
|
|||
|
|
<collectingDate id="F042E9C2EB79ED54FCD7F98B60E98047" box="[818,974,1572,1596]" pageId="17" pageNumber="510" value="1915-06-03" valueMax="1915-06-07" valueMin="1915-06-03">3-7 June 1915</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="394D533CEB79ED54FC3CF98A67438047" box="[985,1124,1572,1596]" pageId="17" pageNumber="510">F.H. Gravely</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FF23F9EA62648027" box="[198,323,1604,1628]" count="1" pageId="17" pageNumber="510" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FEAAF9EB629C8027" box="[335,443,1604,1628]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FE30F9EA61CD8007" ID-GBIF-Occurrence="4905152371" collectingDate="1916-10-24" collectionCode="NHMUK" collectorName="R. V. de Salvaza" country="Laos" location="Dom Toum" municipality="Khone area" pageId="17" pageNumber="510" specimenCount="1" specimenCount-male="1" stateProvince="Champasak">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FE30F9EA61398027" bold="true" box="[469,542,1605,1628]" pageId="17" pageNumber="510">
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FE30F9EA61398027" box="[469,542,1605,1628]" name="Laos" pageId="17" pageNumber="510">LAOS</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FDC8F9EB60338027" box="[557,788,1604,1628]" country="Laos" name="Champasak" pageId="17" pageNumber="510">Champasak Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="7463AC90EB79ED54FCFAF9EB60BC8027" box="[799,923,1604,1628]" pageId="17" pageNumber="510">Khone area</collectingMunicipality>
|
|||
|
|
(
|
|||
|
|
<location id="91676031EB79ED54FC4FF9EB670D8027" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FC4FF9EB670D8027" box="[938,1066,1604,1628]" country="Laos" municipality="Khone area" name="Dom Toum" pageId="17" pageNumber="510" stateProvince="Champasak">Dom Toum</location>
|
|||
|
|
,
|
|||
|
|
<date id="E006102AEB79ED54FBD3F9EB63FC8007" pageId="17" pageNumber="510" value="1916-10-24">
|
|||
|
|
<collectingDate id="F042E9C2EB79ED54FBD3F9EB63FC8007" pageId="17" pageNumber="510" value="1916-10-24">24 October 1916</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="394D533CEB79ED54FF03F9CA62A98007" box="[230,398,1636,1660]" pageId="17" pageNumber="510">R.V. de Salvaza</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FE1AF9CA61418007" box="[511,614,1636,1660]" count="1" pageId="17" pageNumber="510" type="male">one male</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FD94F9CB61FA8007" box="[625,733,1636,1660]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="17" pageNumber="510" type="Museum">NHMUK</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FD10F9CB61F980E7" ID-GBIF-Occurrence="4905152349" collectingDate="1916-01" collectionCode="MFNB" collectorName="R. V. de Salvaza" country="Laos" location="Pak Lay" pageId="17" pageNumber="510" specimenCount="1" specimenCount-female="1" stateProvince="Xaignabouli">
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FD10F9CB60F68007" box="[757,977,1636,1660]" country="Laos" name="Xaignabouli" pageId="17" pageNumber="510">Sainyabuli Province</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<location id="91676031EB79ED54FC3EF9CB67148006" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FC3EF9CB67148006" box="[987,1075,1636,1661]" country="Laos" name="Pak Lay" pageId="17" pageNumber="510" stateProvince="Xaignabouli">Pak Lay</location>
|
|||
|
|
(
|
|||
|
|
<date id="E006102AEB79ED54FBA4F9CB639480E7" pageId="17" pageNumber="510" value="1916-01">
|
|||
|
|
<collectingDate id="F042E9C2EB79ED54FBA4F9CB639480E7" pageId="17" pageNumber="510" value="1916-01">January 1916</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="394D533CEB79ED54FF27F92A625380E7" box="[194,372,1668,1692]" pageId="17" pageNumber="510">R.V. de Salvaza</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FE0BF92A614980E7" box="[494,622,1668,1692]" count="1" pageId="17" pageNumber="510" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FD9BF92B61F680E7" box="[638,721,1668,1692]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="17" pageNumber="510">MFNB</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED54FD08F92A671D80C6" ID-GBIF-Occurrence="4905152364" collectingDate="2013-10-03" collectionCode="CAS" collectorName="L. J. Wang" country="Taiwan" location="Lingshanbi" municipality="Shihmen" pageId="17" pageNumber="510" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="New Taipei">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED54FD08F92A607E80E7" bold="true" box="[749,857,1669,1692]" pageId="17" pageNumber="510">
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED54FD08F92A607E80E7" box="[749,857,1669,1692]" name="Taiwan" pageId="17" pageNumber="510">TAIWAN</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="567CF808EB79ED54FC8EF92A670F80E7" box="[875,1064,1668,1692]" country="Taiwan" name="New Taipei" pageId="17" pageNumber="510">New Taipei City</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<collectingMunicipality id="7463AC90EB79ED54FBD2F92B67B180E7" box="[1079,1174,1668,1692]" pageId="17" pageNumber="510">Shihmen</collectingMunicipality>
|
|||
|
|
(
|
|||
|
|
<location id="91676031EB79ED54FF67F90A63D980C6" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB79ED54FF67F90A63D980C6" box="[130,254,1701,1725]" country="Taiwan" municipality="Shihmen" name="Lingshanbi" pageId="17" pageNumber="510" stateProvince="New Taipei">Lingshanbi</location>
|
|||
|
|
,
|
|||
|
|
<date id="E006102AEB79ED54FEE8F90A629A80C6" box="[269,445,1701,1725]" pageId="17" pageNumber="510" value="2013-10-03">
|
|||
|
|
<collectingDate id="F042E9C2EB79ED54FEE8F90A629A80C6" box="[269,445,1701,1725]" pageId="17" pageNumber="510" value="2013-10-03">3 October 2013</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="394D533CEB79ED54FE2EF90A611C80C6" box="[459,571,1701,1725]" pageId="17" pageNumber="510">L.J. Wang</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FD56F90A603A80C6" box="[691,797,1701,1725]" count="6" pageId="17" pageNumber="510" type="male">six males</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB79ED54FCBCF90A60C480C6" box="[857,995,1701,1725]" count="5" pageId="17" pageNumber="510" type="female">five females</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB79ED54FC17F90A670B80C6" box="[1010,1068,1701,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="510" type="Museum">CAS</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB79ED57FBADF909622F8768" ID-GBIF-Occurrence="4905152343" collectingDate="1990-01-15" collectionCode="CAS" collectorName="J. Nielsen" country="Thailand" lastPageId="18" lastPageNumber="511" location="Ban Hua Hin" pageId="17" pageNumber="510" specimenCount="1" specimenCount-male="1" stateProvince="Prachuap Khiri Khan">
|
|||
|
|
<emphasis id="A6CCEAF8EB79ED57FBADF90963E08688" bold="true" lastPageId="18" lastPageNumber="511" pageId="17" pageNumber="510">
|
|||
|
|
<collectingCountry id="ECAF767AEB79ED57FBADF90963E08688" lastPageId="18" lastPageNumber="511" name="Thailand" pageId="17" pageNumber="510">THAILAND</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="567CF808EB7AED57FF30FF7461028688" box="[213,549,219,243]" country="Thailand" name="Prachuap Khiri Khan" pageId="18" pageNumber="511">Prachuap Khiri Khan Province</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<location id="91676031EB7AED57FDCAFF7461998688" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB7AED57FDCAFF7461998688" box="[559,702,219,243]" country="Thailand" name="Ban Hua Hin" pageId="18" pageNumber="511" stateProvince="Prachuap Khiri Khan">Ban Hua Hin</location>
|
|||
|
|
(
|
|||
|
|
<date id="E006102AEB7AED57FD2BFF7460A38688" box="[718,900,219,243]" pageId="18" pageNumber="511" value="1990-01-15">
|
|||
|
|
<collectingDate id="F042E9C2EB7AED57FD2BFF7460A38688" box="[718,900,219,243]" pageId="18" pageNumber="511" value="1990-01-15">15 January 1990</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="394D533CEB7AED57FC6AFF7460DE8688" box="[911,1017,219,243]" pageId="18" pageNumber="511">J. Nielsen</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB7AED57FB89FF7463958768" count="1" pageId="18" pageNumber="511" type="male">one male</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB7AED57FF25FF5463DD8768" box="[192,250,251,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="511" type="Museum">CAS</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="24D03CB7EB7AED57FEF3FF5463FB8748" ID-GBIF-Occurrence="4905152361" collectingDate="1990-12" collectionCode="MFNB" country="Thailand" elevation="110" location="Lan-Sak" pageId="18" pageNumber="511" specimenCount="2" specimenCount-male="2" stateProvince="Uthai Thani">
|
|||
|
|
<collectingRegion id="567CF808EB7AED57FEF3FF5461228768" box="[278,517,251,275]" country="Thailand" name="Uthai Thani" pageId="18" pageNumber="511">Uthai Thani Province</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<location id="91676031EB7AED57FDF7FF54614B8768" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB7AED57FDF7FF54614B8768" box="[530,620,251,275]" country="Thailand" name="Lan-Sak" pageId="18" pageNumber="511" stateProvince="Uthai Thani">Lan-Sak</location>
|
|||
|
|
(
|
|||
|
|
<quantity id="53409B0FEB7AED57FD9BFF5461E28768" box="[638,709,251,275]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="511" unit="km" value="25.0">25 km</quantity>
|
|||
|
|
NW,
|
|||
|
|
<quantity id="53409B0FEB7AED57FCECFF5460768768" box="[777,849,251,275]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="511" unit="m" value="110.0">
|
|||
|
|
<elevation id="1F95D1D9EB7AED57FCECFF5460768768" box="[777,849,251,275]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="511" unit="m" value="110.0">110 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="E006102AEB7AED57FCBAFF54672C8768" box="[863,1035,251,275]" pageId="18" pageNumber="511" value="1990-12">
|
|||
|
|
<collectingDate id="F042E9C2EB7AED57FCBAFF54672C8768" box="[863,1035,251,275]" pageId="18" pageNumber="511" value="1990-12">December 1990</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
[
|
|||
|
|
<specimenCount id="82BEFD63EB7AED57FBF9FF5367B58768" box="[1052,1170,251,275]" count="2" pageId="18" pageNumber="511" type="male">two males</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB7AED57FF9EFEB463E98748" box="[123,206,283,307]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="18" pageNumber="511">MFNB</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB7AED57FF03FEB4604B8748" ID-GBIF-Occurrence="4905152362" box="[230,876,283,307]" collectionCode="MFNB" country="Thailand" location="Specimens" pageId="18" pageNumber="511" specimenCount="1" specimenCount-male="1" stateProvince="Uthai Thani">
|
|||
|
|
<location id="91676031EB7AED57FF03FEB4627D8748" LSID="urn:lsid:plazi:treatment:1C1187FCEB66ED57FF9EFB71611A8501:91676031EB7AED57FF03FEB4627D8748" box="[230,346,283,307]" country="Thailand" name="Specimens" pageId="18" pageNumber="511" stateProvince="Uthai Thani">Specimens</location>
|
|||
|
|
with illegible locality data: ([
|
|||
|
|
<specimenCount id="82BEFD63EB7AED57FD72FEB461D88748" box="[663,767,283,307]" count="1" pageId="18" pageNumber="511" type="male">one male</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB7AED57FCEEFEB460798748" box="[779,862,283,307]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="18" pageNumber="511">MFNB</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB7AED57FF4EFE93618787AE" blockId="18.[123,1174,219,890]" pageId="18" pageNumber="511">
|
|||
|
|
As presently conceived, the known geographical range of
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FCC5FE9260AB872F" box="[800,908,317,340]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FCC5FE9260AB872F" box="[800,908,317,340]" italics="true" pageId="18" pageNumber="511">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="0C832A6FEB7AED57FC7EFE9360CD872F" box="[923,1002,316,340]" captionStart="FIGURE" captionStartId="39.[171,183,837,856]" captionTargetBox="[168,1128,216,825]" captionTargetId="figure-14@39.[168,1128,216,825]" captionTargetPageId="39" captionText="FIGURE 20. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow circles = C. acutangulus Kavanaugh and Cueva-Dabkoski; white triangles = C. flexuosus Dejean; yellow triangle = locality where C. acutangulus and C. flexuosus have been found together; white squares = C. inustus Andrewes. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/ Whole_world_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513000" httpUri="https://zenodo.org/record/11513000/files/figure.png" pageId="18" pageNumber="511">Fig. 20</figureCitation>
|
|||
|
|
) extends across Southeast Asia from the coast of
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FE0FFEF36106870F" box="[490,545,348,372]" name="India" pageId="18" pageNumber="511">India</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<collectingRegion id="567CF808EB7AED57FDD7FEF36198870F" box="[562,703,348,372]" country="India" name="Odisha" pageId="18" pageNumber="511">Odisha State</collectingRegion>
|
|||
|
|
) on the Bay of Bengal eastward to coastal
|
|||
|
|
<collectingRegion id="567CF808EB7AED57FF9EFED263E587EE" box="[123,194,381,405]" country="China" name="Fujian" pageId="18" pageNumber="511">Fujian</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="567CF808EB7AED57FF2AFED2626987EE" box="[207,334,381,405]" country="China" name="Guangdong" pageId="18" pageNumber="511">Guangdong</collectingRegion>
|
|||
|
|
, and
|
|||
|
|
<collectingRegion id="567CF808EB7AED57FE68FED262CB87EE" box="[397,492,381,405]" country="China" name="Zhejiang" pageId="18" pageNumber="511">Zhejiang</collectingRegion>
|
|||
|
|
provinces (
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FD8BFED2619687EE" box="[622,689,381,405]" name="China" pageId="18" pageNumber="511">China</collectingCountry>
|
|||
|
|
) and
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FD17FED2606487EE" box="[754,835,381,405]" name="Taiwan" pageId="18" pageNumber="511">Taiwan</collectingCountry>
|
|||
|
|
, and south into
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FC1DFED2674187EE" box="[1016,1126,381,405]" name="Cambodia" pageId="18" pageNumber="511">Cambodia</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FF9EFE3263FA87CE" box="[123,221,413,437]" name="Thailand" pageId="18" pageNumber="511">Thailand</collectingCountry>
|
|||
|
|
. This species has not yet been recorded from
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FD16FE31607B87CD" box="[755,860,414,438]" name="Myanmar" pageId="18" pageNumber="511">Myanmar</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FC89FE3260EE87CE" box="[876,969,413,437]" name="Vietnam" pageId="18" pageNumber="511">Vietnam</collectingCountry>
|
|||
|
|
, or
|
|||
|
|
<collectingRegion id="567CF808EB7AED57FC1EFE32676F87CE" box="[1019,1096,413,437]" country="China" name="Hainan" pageId="18" pageNumber="511">Hainan</collectingRegion>
|
|||
|
|
Island (
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FF67FE1263E087AE" box="[130,199,445,469]" name="China" pageId="18" pageNumber="511">China</collectingCountry>
|
|||
|
|
), but it is likely to occur also in these areas.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB7AED57FF4EFE7061FD8482" blockId="18.[123,1174,219,890]" pageId="18" pageNumber="511">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FF4EFE706298878D" bold="true" box="[171,447,479,502]" pageId="18" pageNumber="511">Geographical variation.</emphasis>
|
|||
|
|
Most specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FD4CFE71603E878D" box="[681,793,478,502]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FD4CFE71603E878D" box="[681,793,478,502]" italics="true" pageId="18" pageNumber="511">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from localities in
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FC0AFE71671A878D" box="[1007,1085,478,502]" name="Taiwan" pageId="18" pageNumber="511">Taiwan</collectingCountry>
|
|||
|
|
and on
|
|||
|
|
<collectingRegion id="567CF808EB7AED57FF9EFE5163F6846D" box="[123,209,510,534]" country="Taiwan" name="Kinmen" pageId="18" pageNumber="511">Kinmen</collectingRegion>
|
|||
|
|
and Nanao Islands (Guangdong) are smaller than those from mainland areas and have the elytral dark pattern paler (rufous or rufopiceous rather than piceous) and reduced in extent (see
|
|||
|
|
<bibRefCitation id="F0294B1BEB7AED57FF9EFDEF6217842C" author="WANG, L. J. & J. J. CHERNG & D. H. KAVANAUGH" box="[123,304,575,600]" pageId="18" pageNumber="511" pagination="107 - 116" refId="ref23969" refString="WANG, L. J., J. J. CHERNG, AND D. H. KAVANAUGH. 2017. First records for Cyclosomus inustus Andrewes (Coleoptera: Carabidae: Cyclosomini) for Taiwan, with notes on habitat and behavior. Proceedings of the California Academy of Sciences (Series 4), 64: 107 - 116." type="journal article" year="2017">Wang et al. 2017</bibRefCitation>
|
|||
|
|
for images showing this variation), with the middle dark band absent from some specimens (
|
|||
|
|
<figureCitation id="0C832A6FEB7AED57FF18FDF06272840C" box="[253,341,607,631]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="18" pageNumber="511">Fig. 7C</figureCitation>
|
|||
|
|
). At least some specimens from inland parts of Indochina (
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FC11FDF06744840C" box="[1012,1123,607,631]" name="Cambodia" pageId="18" pageNumber="511">Cambodia</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FF9EFD2F63FB84E3" box="[123,220,640,664]" name="Thailand" pageId="18" pageNumber="511">Thailand</collectingCountry>
|
|||
|
|
) have a small and faint preapical dark spot present (
|
|||
|
|
<figureCitation id="0C832A6FEB7AED57FCE0FD2F607D84E3" box="[773,858,640,664]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="18" pageNumber="511">Fig. 7D</figureCitation>
|
|||
|
|
). Specimens from these areas (and
|
|||
|
|
<collectingCountry id="ECAF767AEB7AED57FF52FD0F63CB84C3" box="[183,236,672,696]" name="Laos" pageId="18" pageNumber="511">Laos</collectingCountry>
|
|||
|
|
) are more darkly pigmented (pronotum piceous rather than rufopiceous or rufous, middle dark band of the elytra black rather than rufopiceous). However, all of these specimens have male genitalia similar to those of other
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FDC7FD4E61A98482" box="[546,654,737,761]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FDC7FD4E61A98482" box="[546,654,737,761]" italics="true" pageId="18" pageNumber="511">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
males.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB7AED57FF4EFCAD611A8501" blockId="18.[123,1174,219,890]" pageId="18" pageNumber="511">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FF4EFCAD60768562" bold="true" box="[171,849,769,793]" pageId="18" pageNumber="511">
|
|||
|
|
Geographical relationships with other
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FD8BFCAE60768562" box="[622,849,769,793]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="undetermined">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FD8BFCAE61D58562" bold="true" box="[622,754,769,793]" italics="true" pageId="18" pageNumber="511">Cyclosomus</emphasis>
|
|||
|
|
species.
|
|||
|
|
</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
To date, this species has not been found syntopic with any other congeneric species; however, it may be sympatric with
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FBA2FC8D639F8521" authorityName=", Kavanaugh" authorityYear="2015" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FBA2FC8D639F8521" italics="true" pageId="18" pageNumber="511">C. flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FF22FCEC62478521" box="[199,352,835,858]" italics="true" pageId="18" pageNumber="511">
|
|||
|
|
C,
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FF02FCEC62478521" box="[231,352,835,858]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="marginatus">marginatus</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
, and/or
|
|||
|
|
<taxonomicName id="53B84D69EB7AED57FE5EFCEC611E8521" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[443,569,834,858]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="511" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB7AED57FE5EFCEC611E8521" box="[443,569,834,858]" italics="true" pageId="18" pageNumber="511">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in the western part of its range, but none of these relationships have been demonstrated to date.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|