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<document id="C7C2F4F818194F56645AF6BBB34E4629" ID-CLB-Dataset="273378" ID-DOI="10.11646/zootaxa.5357.1.3" ID-GBIF-Dataset="ef13171a-ac9d-4abd-bd0b-567af48ea4a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10012545" ID-ZooBank="5DDC347A-3DE3-45F0-8BB5-7EAD1DE87E7C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1697528556513" checkinUser="plazi" docAuthor="Anichtchenko, Alexander &amp; Wiesner, Jürgen" docDate="2023" docId="03BB87D4FFAF2E34FF03946AFBDC0C3F" docLanguage="en" docName="zootaxa.5357.1.3.pdf" docOrigin="Zootaxa 5357 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5357.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Prothyma (Symplecthyma) kapuka Anichtchenko &amp; Wiesner 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="95" masterDocId="FF82FFACFFB92E2CFF94905BFFA60F62" masterDocTitle="Philippine species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae). Part 2. Subgenus Symplecthyma Rivalier, 1964" masterLastPageNumber="99" masterPageNumber="71" pageNumber="93" updateTime="1697658095773" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="ED3532ECF522E12D519064C30EB074D4">Philippine species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae). Part 2. Subgenus Symplecthyma Rivalier, 1964</mods:title>
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<mods:namePart id="0CCBDA249858C1F92F6CB1DDBC2ED105">Wiesner, Jürgen</mods:namePart>
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<mods:date id="4B44CA02BC50B209DD1B738556367852">2023</mods:date>
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<treatment id="03BB87D4FFAF2E34FF03946AFBDC0C3F" ID-DOI="http://doi.org/10.5281/zenodo.10012607" ID-GBIF-Taxon="214039276" ID-Zenodo-Dep="10012607" LSID="urn:lsid:plazi:treatment:03BB87D4FFAF2E34FF03946AFBDC0C3F" httpUri="http://treatment.plazi.org/id/03BB87D4FFAF2E34FF03946AFBDC0C3F" lastPageId="24" lastPageNumber="95" pageId="22" pageNumber="93">
<subSubSection id="C3086549FFAF2E3AFF03946AFCDC0B29" box="[151,890,1073,1100]" pageId="22" pageNumber="93" type="nomenclature">
<paragraph id="8BAD36C2FFAF2E3AFF03946AFCDC0B29" blockId="22.[151,890,1073,1100]" box="[151,890,1073,1100]" pageId="22" pageNumber="93">
<heading id="D0E581AEFFAF2E3AFF03946AFCDC0B29" bold="true" box="[151,890,1073,1100]" fontSize="11" level="1" pageId="22" pageNumber="93" reason="1">
<emphasis id="B966EAD0FFAF2E3AFF03946AFCDC0B29" bold="true" box="[151,890,1073,1100]" pageId="22" pageNumber="93">
<taxonomicName id="4C124D41FFAF2E3AFF03946AFD9C0B2E" authority="Anichtchenko &amp; Wiesner, 2023" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[151,570,1073,1100]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="93" phylum="Arthropoda" rank="species" species="kapuka" status="sp. nov." subGenus="Symplecthyma">
<emphasis id="B966EAD0FFAF2E3AFF03946AFEA80B2E" bold="true" box="[151,270,1073,1100]" italics="true" pageId="22" pageNumber="93">Prothyma</emphasis>
(
<emphasis id="B966EAD0FFAF2E3AFE8B946AFE690B2E" bold="true" box="[287,463,1073,1100]" italics="true" pageId="22" pageNumber="93">Symplecthyma</emphasis>
)
<emphasis id="B966EAD0FFAF2E3AFE74946AFD9C0B2E" bold="true" box="[480,570,1073,1100]" italics="true" pageId="22" pageNumber="93">kapuka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557ABFFAF2E3AFDD5946AFD390B29" box="[577,671,1073,1099]" pageId="22" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13292A47FFAF2E3AFD3A946AFCAD0B29" box="[686,779,1073,1099]" captionStart="FIGURE 1" captionStartId="5.[152,255,1553,1577]" captionTargetBox="[181,1415,198,1509]" captionTargetId="figure-17@5.[151,1436,181,1529]" captionTargetPageId="5" captionText="FIGURE 1. Ventral habitus of Prothyma (Symplecthyma). a) P. inopinata sp. nov.; b) P. kapuka sp. nov.; c) P. aeneoparva Horn, 1934. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10012549" httpUri="https://zenodo.org/record/10012549/files/figure.png" pageId="22" pageNumber="93">Figs 1b</figureCitation>
,
<figureCitation id="13292A47FFAF2E3AFC8D946AFC9E0B29" box="[793,824,1073,1099]" captionStart="FIGURE 12" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[242,1345,186,1908]" captionTargetId="figure-17@23.[230,1358,181,1919]" captionTargetPageId="23" captionText="FIGURE 12. Prothyma (Symplecthyma) kapuka sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, female; d) Head, male; e–f) Aedeagus, left and right lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012587" httpUri="https://zenodo.org/record/10012587/files/figure.png" pageId="22" pageNumber="93">12</figureCitation>
,
<figureCitation id="13292A47FFAF2E3AFCD1946AFCD60B29" box="[837,880,1073,1099]" captionStart="FIGURE 15" captionStartId="27.[152,255,1319,1343]" captionTargetBox="[177,1422,728,1273]" captionTargetId="figure-241@27.[152,1436,705,1296]" captionTargetPageId="27" captionText="FIGURE 15. Head and pronotum of Symplecthyma species. a) P. (S.) incerata Rivalier, 1964; b) P. (S.) inopinata sp. nov.; c) P. (S.) kapuka sp. nov. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012599" httpUri="https://zenodo.org/record/10012599/files/figure.png" pageId="22" pageNumber="93">15c</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3086549FFAF2E3AFF039428FC9C0A7F" pageId="22" pageNumber="93" type="materials_examined">
<paragraph id="8BAD36C2FFAF2E3AFF039428FC9C0A7F" blockId="22.[151,1437,1138,2029]" pageId="22" pageNumber="93">
<emphasis id="B966EAD0FFAF2E3AFF039428FEEC0BEE" bold="true" box="[151,330,1138,1165]" pageId="22" pageNumber="93">Type material.</emphasis>
<materialsCitation id="3B7A3C9FFFAF2E3AFEC39428FCED0BB7" ID-GBIF-Occurrence="4427149351" collectingDate="2023-04-29" collectingDateMax="2023-04-30" collectingDateMin="2023-04-29" collectionCode="DUBC" collectorName="Anichtchenko A." country="Philippines" county="Mindanao" elevation="260" latitude="7.440359" location="Kapuka" longLatPrecision="1" longitude="126.43403" municipality="Ugwad Falls" pageId="22" pageNumber="93" specimenCount="2" stateProvince="Davao" typeStatus="holotype">
<emphasis id="B966EAD0FFAF2E3AFEC39428FDBC0BEF" bold="true" box="[343,538,1139,1165]" pageId="22" pageNumber="93">
<typeStatus id="54A98860FFAF2E3AFEC39428FE600BEF" box="[343,454,1139,1165]" pageId="22" pageNumber="93">Holotype</typeStatus>
, male:
</emphasis>
“
<collectingCountry id="F3057652FFAF2E3AFDA49428FD150BEF" box="[560,691,1139,1165]" name="Philippines" pageId="22" pageNumber="93">Philippines</collectingCountry>
,
<collectingCounty id="62CC4E4EFFAF2E3AFD509428FC9E0BEF" box="[708,824,1139,1165]" pageId="22" pageNumber="93">Mindanao</collectingCounty>
,
<collectingMunicipality id="6BC9ACB8FFAF2E3AFCDD9428FC7B0BEF" box="[841,989,1139,1165]" pageId="22" pageNumber="93">Ugwad Falls</collectingMunicipality>
(
<location id="8ECD6019FFAF2E3AFC659428FBE80BEF" LSID="urn:lsid:plazi:treatment:03BB87D4FFAF2E34FF03946AFBDC0C3F:8ECD6019FFAF2E3AFC659428FBE80BEF" box="[1009,1102,1139,1165]" country="Philippines" county="Mindanao" latitude="7.440359" longLatPrecision="1" longitude="126.43403" municipality="Ugwad Falls" name="Kapuka" pageId="22" pageNumber="93" stateProvince="Davao">Kapuka</location>
),
<location id="8ECD6019FFAF2E3AFBF39428FB1F0BEF" LSID="urn:lsid:plazi:treatment:03BB87D4FFAF2E34FF03946AFBDC0C3F:8ECD6019FFAF2E3AFBF39428FB1F0BEF" box="[1127,1209,1139,1165]" country="Philippines" county="Mindanao" latitude="7.440359" longLatPrecision="1" longitude="126.43403" municipality="Ugwad Falls" name="Caraga" pageId="22" pageNumber="93" stateProvince="Davao">Caraga</location>
,
<collectingRegion id="49D6F820FFAF2E3AFB5D9428FAB40BEF" box="[1225,1298,1139,1165]" country="Philippines" name="Davao" pageId="22" pageNumber="93">Davao</collectingRegion>
Or.,
<quantity id="4CEA9B27FFAF2E3AFAC19428FA3E0BEF" box="[1365,1432,1139,1165]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="22" pageNumber="93" unit="m" value="260.0">
<elevation id="003FD1F1FFAF2E3AFAC19428FA3E0BEF" box="[1365,1432,1139,1165]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="22" pageNumber="93" unit="m" value="260.0">260m</elevation>
</quantity>
,
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,
<geoCoordinate id="EE265005FFAF2E3AFE9894CCFE320BD3" box="[268,404,1175,1201]" degrees="126.434027" direction="east" orientation="longitude" pageId="22" pageNumber="93" precision="1" value="126.43403">126.434027</geoCoordinate>
,
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<collectingDate id="EFE8E9EAFFAF2E3AFE0994CCFDE30BD3" box="[413,581,1175,1201]" pageId="22" pageNumber="93" value="2023-04-29" valueMax="2023-04-30" valueMin="2023-04-29">29–30.IV.2023</collectingDate>
</date>
,
<collectorName id="26E75314FFAF2E3AFDD994CCFCAF0BD3" box="[589,777,1175,1201]" pageId="22" pageNumber="93">Anichtchenko A.</collectorName>
leg.“, “
<typeStatus id="54A98860FFAF2E3AFCCD94CCFC560BD3" box="[857,1008,1175,1201]" pageId="22" pageNumber="93">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C124D41FFAF2E3AFB9594CCFBC80BD3" authorityName=": Horn" authorityYear="1923" box="[1025,1134,1175,1201]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="93" phylum="Arthropoda" rank="genus">Prothyma</taxonomicName>
/ (
<taxonomicName id="4C124D41FFAF2E3AFB1394CCFA950BD3" authorityName="Rivalier" authorityYear="1964" box="[1159,1331,1175,1201]" pageId="22" pageNumber="73" rank="subGenus" subGenus="Symplecthyma">Symplecthyma</taxonomicName>
) /
<taxonomicName id="4C124D41FFAF2E3AFADF94CCFA3A0BD3" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[1355,1436,1175,1201]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="93" phylum="Arthropoda" rank="species" species="kapuka">
<emphasis id="B966EAD0FFAF2E3AFADF94CCFA3A0BD3" box="[1355,1436,1175,1201]" italics="true" pageId="22" pageNumber="93">kapuka</emphasis>
</taxonomicName>
/ des.
<bibRefCitation id="EF834B33FFAF2E3AFF4D94E0FD910BB7" author="Wiesner, J. &amp; Anichtchenko, A." box="[217,567,1211,1237]" pageId="22" pageNumber="93" pagination="1 - 36" refId="ref17139" refString="Wiesner, J. &amp; Anichtchenko, A. (2023) The Philippines species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae) Part 1: Subgenera Prothyma Hope, 1838, Genoprothyma Rivalier, 1964 and Parasymplecthyma subgen. nov. Zootaxa, 5315 (1), 1 - 36." type="journal article" year="2023">Anichtchenko &amp; Wiesner 2023</bibRefCitation>
[printed, red]” (
<collectionCode id="ED03AE07FFAF2E3AFD7B94E0FCE50BB7" box="[751,835,1211,1237]" pageId="22" pageNumber="93">DUBC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7A3C9FFFAF2E3AFCC294E0FA3E0BB7" ID-GBIF-Occurrence="4427149347" box="[854,1432,1211,1237]" collectingDate="2023-04-29" collectingDateMax="2023-04-30" collectingDateMin="2023-04-29" collectionCode="DUBC, JWc" collectorName="Anichtchenko A." country="Philippines" county="Mindanao" elevation="260" latitude="7.440359" location="Kapuka" longLatPrecision="1" longitude="126.43403" municipality="Ugwad Falls" pageId="22" pageNumber="93" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Davao" typeStatus="paratype">
<typeStatus id="54A98860FFAF2E3AFCC294E0FC610BB7" box="[854,967,1211,1237]" pageId="22" pageNumber="93">Paratypes</typeStatus>
:
<specimenCount id="9D14FD4BFFAF2E3AFC4694E0FB8A0BB7" box="[978,1068,1211,1237]" pageId="22" pageNumber="93" type="male">3 males</specimenCount>
,
<specimenCount id="9D14FD4BFFAF2E3AFBAC94E0FB0F0BB7" box="[1080,1193,1211,1237]" pageId="22" pageNumber="93" type="female">3 females</specimenCount>
: idem (
<collectionCode id="ED03AE07FFAF2E3AFB6F94E0FAEB0BB7" box="[1275,1357,1211,1237]" pageId="22" pageNumber="93">DUBC</collectionCode>
,
<collectionCode id="ED03AE07FFAF2E3AFACD94E0FA290BB7" box="[1369,1423,1211,1237]" pageId="22" pageNumber="93">JWc</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7A3C9FFFAF2E3AFF039484FDDC0B9B" ID-GBIF-Occurrence="4427149339" box="[151,634,1247,1273]" collectingDate="2023-04-29" collectingDateMax="2023-04-30" collectingDateMin="2023-04-29" collectionCode="DUBC, JWc" collectorName="M. Medina &amp; All" country="Philippines" county="Mindanao" elevation="260" latitude="7.440359" location="Kapuka" longLatPrecision="1" longitude="126.43403" municipality="Ugwad Falls" pageId="22" pageNumber="93" specimenCount="1" specimenCount-male="1" stateProvince="Davao" typeStatus="paratype">
<specimenCount id="9D14FD4BFFAF2E3AFF039484FF4F0B9B" box="[151,233,1247,1273]" pageId="22" pageNumber="93" type="male">1 male</specimenCount>
: idem, but
<collectorName id="26E75314FFAF2E3AFEFD9484FE410B9B" box="[361,487,1247,1273]" pageId="22" pageNumber="93">M. Medina</collectorName>
leg. (MMc).
</materialsCitation>
<materialsCitation id="3B7A3C9FFFAF2E3AFD159484FC930A7F" ID-GBIF-Occurrence="4427149364" collectingDate="2023-04-29" collectingDateMax="2023-04-30" collectingDateMin="2023-04-29" collectionCode="DUBC, JWc" collectorName="M. Medina &amp; All" country="Philippines" county="Mindanao" elevation="260" latitude="7.440359" location="Kapuka" longLatPrecision="1" longitude="126.43403" municipality="Ugwad Falls" pageId="22" pageNumber="93" specimenCount="1" specimenCount-male="1" stateProvince="Davao" typeStatus="paratype">
<collectorName id="26E75314FFAF2E3AFD159484FD030B9B" box="[641,677,1247,1273]" pageId="22" pageNumber="93">All</collectorName>
paratypes with label “
<typeStatus id="54A98860FFAF2E3AFC309484FB940B9B" box="[932,1074,1247,1273]" pageId="22" pageNumber="93">PARATYPE</typeStatus>
/
<taxonomicName id="4C124D41FFAF2E3AFBDF9484FB1E0B9B" authorityName=": Horn" authorityYear="1923" box="[1099,1208,1247,1273]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="93" phylum="Arthropoda" rank="genus">Prothyma</taxonomicName>
/ (
<taxonomicName id="4C124D41FFAF2E3AFB4D9484FA230B9B" authorityName="Rivalier" authorityYear="1964" box="[1241,1413,1247,1273]" pageId="22" pageNumber="73" rank="subGenus" subGenus="Symplecthyma">Symplecthyma</taxonomicName>
) /
<taxonomicName id="4C124D41FFAF2E3AFF039558FF4E0A7F" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[151,232,1283,1309]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="93" phylum="Arthropoda" rank="species" species="kapuka">
<emphasis id="B966EAD0FFAF2E3AFF039558FF4E0A7F" box="[151,232,1283,1309]" italics="true" pageId="22" pageNumber="93">kapuka</emphasis>
</taxonomicName>
/ des.
<bibRefCitation id="EF834B33FFAF2E3AFEA49558FD2D0A7F" author="Wiesner, J. &amp; Anichtchenko, A." box="[304,651,1283,1309]" pageId="22" pageNumber="93" pagination="1 - 36" refId="ref17139" refString="Wiesner, J. &amp; Anichtchenko, A. (2023) The Philippines species of the genus Prothyma Hope, 1838 (Coleoptera: Cicindelidae) Part 1: Subgenera Prothyma Hope, 1838, Genoprothyma Rivalier, 1964 and Parasymplecthyma subgen. nov. Zootaxa, 5315 (1), 1 - 36." type="journal article" year="2023">Anichtchenko &amp; Wiesner 2023</bibRefCitation>
[printed, red]”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3086549FFAF2E3AFF53957CFB560A23" box="[199,1264,1319,1345]" pageId="22" pageNumber="93" type="etymology">
<paragraph id="8BAD36C2FFAF2E3AFF53957CFB560A23" blockId="22.[151,1437,1138,2029]" box="[199,1264,1319,1345]" pageId="22" pageNumber="93">
<emphasis id="B966EAD0FFAF2E3AFF53957CFEE80A23" bold="true" box="[199,334,1319,1345]" pageId="22" pageNumber="93">Etymology.</emphasis>
The species is named after Kapuka Falls, near which the
<typeStatus id="54A98860FFAF2E3AFC47957CFBA50A23" box="[979,1027,1319,1345]" pageId="22" pageNumber="93">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="C3086549FFAF2E3AFF539511FE4E0AEB" pageId="22" pageNumber="93" type="diagnosis">
<paragraph id="8BAD36C2FFAF2E3AFF539511FE4E0AEB" blockId="22.[151,1437,1138,2029]" pageId="22" pageNumber="93">
<emphasis id="B966EAD0FFAF2E3AFF539511FE9C0A06" bold="true" box="[199,314,1354,1380]" pageId="22" pageNumber="93">Diagnosis</emphasis>
. Dorsal coloration and bright luster resemble
<taxonomicName id="4C124D41FFAF2E3AFCD79510FC0C0A07" authority="Anichtchenko &amp; Wiesner, 2023" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[835,938,1355,1381]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="93" phylum="Arthropoda" rank="species" species="eureka" status="sp. nov.">
<emphasis id="B966EAD0FFAF2E3AFCD79510FC0C0A07" box="[835,938,1355,1381]" italics="true" pageId="22" pageNumber="93">P. eureka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557ABFFAF2E3AFC269510FBAB0A07" box="[946,1037,1355,1381]" pageId="22" pageNumber="93" rank="species">
<emphasis id="B966EAD0FFAF2E3AFC269510FBAE0A07" bold="true" box="[946,1032,1355,1381]" pageId="22" pageNumber="93">sp. nov</emphasis>
.
</taxonomicNameLabel>
, but is readily distinguished by the light brown abdominal sterna.
</paragraph>
</subSubSection>
<subSubSection id="C3086549FFAF2E34FF5395C9FC5C0C5B" lastPageId="24" lastPageNumber="95" pageId="22" pageNumber="93" type="description">
<paragraph id="8BAD36C2FFAF2E3AFF5395C9FB850ACF" blockId="22.[151,1437,1138,2029]" box="[199,1059,1426,1453]" pageId="22" pageNumber="93">
<emphasis id="B966EAD0FFAF2E3AFF5395C9FEFF0ACE" bold="true" box="[199,345,1426,1452]" pageId="22" pageNumber="93">Description.</emphasis>
<emphasis id="B966EAD0FFAF2E3AFEF495C8FE300ACF" box="[352,406,1427,1453]" italics="true" pageId="22" pageNumber="93">Size:</emphasis>
TL (without labrum)
<quantity id="4CEA9B27FFAF2E3AFD1E95C9FCAC0ACF" box="[650,778,1426,1453]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" pageId="22" pageNumber="93" unit="mm" value="9.25" valueMax="10.0" valueMin="8.5">8.5–10 mm</quantity>
(mean =
<quantity id="4CEA9B27FFAF2E3AFCE195C8FC680ACF" box="[885,974,1427,1453]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="22" pageNumber="93" unit="mm" value="9.3">9.3 mm</quantity>
; n = 8).
</paragraph>
<paragraph id="8BAD36C2FFAF2E3AFF5395ECFAC8088F" blockId="22.[151,1437,1138,2029]" pageId="22" pageNumber="93">
<emphasis id="B966EAD0FFAF2E3AFF5395ECFEAA0AB3" box="[199,268,1463,1489]" italics="true" pageId="22" pageNumber="93">Head:</emphasis>
(
<figureCitation id="13292A47FFAF2E3AFE8F95ECFEDB0AB3" box="[283,381,1463,1489]" captionStart="FIGURE 12" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[242,1345,186,1908]" captionTargetId="figure-17@23.[230,1358,181,1919]" captionTargetPageId="23" captionText="FIGURE 12. Prothyma (Symplecthyma) kapuka sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, female; d) Head, male; e–f) Aedeagus, left and right lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012587" httpUri="https://zenodo.org/record/10012587/files/figure.png" pageId="22" pageNumber="93">Figs 12c</figureCitation>
,
<figureCitation id="13292A47FFAF2E3AFE1D95EDFE140AB3" box="[393,434,1462,1489]" captionStart="FIGURE 15" captionStartId="27.[152,255,1319,1343]" captionTargetBox="[177,1422,728,1273]" captionTargetId="figure-241@27.[152,1436,705,1296]" captionTargetPageId="27" captionText="FIGURE 15. Head and pronotum of Symplecthyma species. a) P. (S.) incerata Rivalier, 1964; b) P. (S.) inopinata sp. nov.; c) P. (S.) kapuka sp. nov. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012599" httpUri="https://zenodo.org/record/10012599/files/figure.png" pageId="22" pageNumber="93">15c</figureCitation>
) slightly wider than elytra (mean EW/HW = 0.87). Dorsally color shiny copper; with gold and copper reflections; orbital plates with copper reflections; area behind the eye and near orbital plates with blue and green reflections; genae blue-violet; clypeus with greenish-blue reflections; surface glabrous, with two setigerous punctures next to eye, one along frontal margin and one dorsally at center; strong longitudinal striae on orbital plates and vertex; weakly transverse, moderately zigzag striae behind the eyes and on neck; striae becoming weak on genae, frons, and clypeus. Labrum golden-red. Labrum of male (
<figureCitation id="13292A47FFAF2E3AFCB99630FC2B09E7" box="[813,909,1643,1669]" captionStart="FIGURE 12" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[242,1345,186,1908]" captionTargetId="figure-17@23.[230,1358,181,1919]" captionTargetPageId="23" captionText="FIGURE 12. Prothyma (Symplecthyma) kapuka sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, female; d) Head, male; e–f) Aedeagus, left and right lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012587" httpUri="https://zenodo.org/record/10012587/files/figure.png" pageId="22" pageNumber="93">Fig. 12c</figureCitation>
) wider than long (mean LW/LL = 1.23); with two setae located near the lateral margins and two setae centrally, between midline and lateral margins in anterior fourth; with two obtuse anterolateral teeth and two outward directed thorn-like anterior teeth at the anteromedian lobe, intermediate space slightly convex. Labrum of female (
<figureCitation id="13292A47FFAF2E3AFCC3968CFC1C0993" box="[855,954,1751,1777]" captionStart="FIGURE 12" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[242,1345,186,1908]" captionTargetId="figure-17@23.[230,1358,181,1919]" captionTargetPageId="23" captionText="FIGURE 12. Prothyma (Symplecthyma) kapuka sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, female; d) Head, male; e–f) Aedeagus, left and right lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012587" httpUri="https://zenodo.org/record/10012587/files/figure.png" pageId="22" pageNumber="93">Fig. 12d</figureCitation>
) almost as long as wide (mean LW/LL = 1.04), with two setae located near the lateral margins and two setae centrally, between midline and lateral margins in anterior fourth; with two blunt anterolateral teeth and two small forward directed anterior teeth at the anteromedian lobe, intermediate space with a long and slightly downturned triangular tooth. Mandibles light brownish, with four brown teeth. Labial and maxillary palpi testaceous, the last segment of both light brown. Antennae slender, extending posteriorly slightly past the middle of the elytra in the male, shorter in the females; scape and antennomere two light brown; antennomeres 3–4 with metallic purple luster, brown in the anterior third; scape with one apical seta, the succeeding three antennomeres almost glabrous; antennomeres 5–11 black, matte, finely, and evenly pubescent.
</paragraph>
<caption id="DF6D664AFFAE2E3BFF0C97CCFA2F08B1" ID-DOI="http://doi.org/10.5281/zenodo.10012587" ID-Zenodo-Dep="10012587" httpUri="https://zenodo.org/record/10012587/files/figure.png" pageId="23" pageNumber="94" startId="23.[152,255,1943,1967]" targetBox="[242,1345,186,1908]" targetPageId="23" targetType="figure">
<paragraph id="8BAD36C2FFAE2E3BFF0C97CCFA2F08B1" blockId="23.[151,1436,1943,2004]" pageId="23" pageNumber="94">
<emphasis id="B966EAD0FFAE2E3BFF0C97CCFE8108CD" bold="true" box="[152,295,1943,1967]" pageId="23" pageNumber="94">FIGURE 12.</emphasis>
<taxonomicName id="4C124D41FFAE2E3BFEBB97C3FD2808D2" authority="Anichtchenko &amp; Wiesner, 2023" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[303,654,1944,1968]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="94" phylum="Arthropoda" rank="species" species="kapuka" status="sp. nov." subGenus="Symplecthyma">
<emphasis id="B966EAD0FFAE2E3BFEBB97C3FE3408D2" box="[303,402,1944,1968]" italics="true" pageId="23" pageNumber="94">Prothyma</emphasis>
(
<emphasis id="B966EAD0FFAE2E3BFE3697C3FD9208D2" box="[418,564,1944,1968]" italics="true" pageId="23" pageNumber="94">Symplecthyma</emphasis>
)
<emphasis id="B966EAD0FFAE2E3BFDD197C3FD2808D2" box="[581,654,1944,1968]" italics="true" pageId="23" pageNumber="94">kapuka</emphasis>
</taxonomicName>
<emphasis id="B966EAD0FFAE2E3BFD0297C3FCA008CD" bold="true" box="[662,774,1943,1968]" pageId="23" pageNumber="94">
<taxonomicNameLabel id="A25557ABFFAE2E3BFD0297C3FD4E08D2" box="[662,744,1944,1968]" pageId="23" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
a)
</emphasis>
Dorsal habitus, Holotype, male. Scale = 5 mm.
<emphasis id="B966EAD0FFAE2E3BFB6997CCFAB208CD" bold="true" box="[1277,1300,1943,1967]" pageId="23" pageNumber="94">b)</emphasis>
Left elytron, paratype, female. Scale = 5 mm.
<emphasis id="B966EAD0FFAE2E3BFE7297E0FE5F08B1" bold="true" box="[486,505,1979,2003]" pageId="23" pageNumber="94">c)</emphasis>
Labrum, female;
<emphasis id="B966EAD0FFAE2E3BFD3B97E0FD6008B1" bold="true" box="[687,710,1979,2003]" pageId="23" pageNumber="94">d)</emphasis>
Head, male;
<emphasis id="B966EAD0FFAE2E3BFCD897E0FCD308B1" bold="true" box="[844,885,1979,2003]" pageId="23" pageNumber="94">e–f)</emphasis>
Aedeagus, left and right lateral view. Scale = 1 mm.
</paragraph>
</caption>
<paragraph id="8BAD36C2FFA12E34FF5390CCFA950F9B" blockId="24.[151,1437,150,861]" pageId="24" pageNumber="95">
<emphasis id="B966EAD0FFA12E34FF5390CCFE870FD3" box="[199,289,151,177]" italics="true" pageId="24" pageNumber="95">Thorax:</emphasis>
pronotum slightly longer than wide (mean PW/PL = 0.85); sides weakly rounded between the transvers constrictions, nearly parallel-sided; glabrous; transverse discal wrinkles weakly impressed throughout; shiny copper with weak greenish reflections dorsally; transverse constrictions and margins with light green reflections.
</paragraph>
<paragraph id="8BAD36C2FFA12E34FF539158FA8B0E97" blockId="24.[151,1437,150,861]" pageId="24" pageNumber="95">
<emphasis id="B966EAD0FFA12E34FF539158FEB00E7F" box="[199,278,259,285]" italics="true" pageId="24" pageNumber="95">Elytra:</emphasis>
(
<figureCitation id="13292A47FFA12E34FEB19158FE040E7F" box="[293,418,259,285]" captionStart="FIGURE 12" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[242,1345,186,1908]" captionTargetId="figure-17@23.[230,1358,181,1919]" captionTargetPageId="23" captionText="FIGURE 12. Prothyma (Symplecthyma) kapuka sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, female; d) Head, male; e–f) Aedeagus, left and right lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012587" httpUri="https://zenodo.org/record/10012587/files/figure.png" pageId="24" pageNumber="95">Figs 12a, b</figureCitation>
) twice as long as wide (mean EL/EW = 2.08), parallel-sided, with a dorsally flattened basal hump, dorsal surface without microreticulation, very shining coppery with greenish reflections behind humeri; testaceous elytral maculation in males consists of humeral, lateromedial and preapical spots; the humeral spot is relatively large and located at the humeri; the lateromedial spot is wide and oblique; the preapical spot is relatively large, oblique and reaching margins; in females humeral spot absent. Elytra densely covered by narrow transverse punctures, becoming confluent near the suture. Apical margin minutely serrate, distinctly rounded and restricted towards middle suture, with a minute sutural spine. Epipleurae brown, on anterior half with bluish luster.
</paragraph>
<paragraph id="8BAD36C2FFA12E34FF5391A4FDD70DCB" blockId="24.[151,1437,150,861]" pageId="24" pageNumber="95">
<emphasis id="B966EAD0FFA12E34FF5391A4FED50D7A" box="[199,371,511,537]" italics="true" pageId="24" pageNumber="95">Ventral aspect:</emphasis>
venter glabrous; sternae and episternae glabrous; trochanter glabrous, coxa with two setae; femur, tibia, and tarsi setose. Prosternum, mesosternum and episterna violet. Metasternum greenish violet with brown central part. Trochanter, pro- and mesocoxa, femurs and tibiae testaceous; tarsa and apices of protibia dark brown. Metacoxa brown and partially blue; all abdominal sterna light brown. Protarsomeres
<quantity id="4CEA9B27FFA12E34FB3D9230FB570DE7" box="[1193,1265,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="24" pageNumber="95" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3 in</quantity>
males laterally and ventrally covered by long white setae.
</paragraph>
<paragraph id="8BAD36C2FFA12E34FF5392E8FC5C0C5B" blockId="24.[151,1437,150,861]" pageId="24" pageNumber="95">
<emphasis id="B966EAD0FFA12E34FF5392E8FE920DAF" box="[199,308,691,717]" italics="true" pageId="24" pageNumber="95">Aedeagus</emphasis>
: (
<figureCitation id="13292A47FFA12E34FEDF92E8FE610DAF" box="[331,455,691,717]" captionStart="FIGURE 12" captionStartId="23.[152,255,1943,1967]" captionTargetBox="[242,1345,186,1908]" captionTargetId="figure-17@23.[230,1358,181,1919]" captionTargetPageId="23" captionText="FIGURE 12. Prothyma (Symplecthyma) kapuka sp. nov. a) Dorsal habitus, Holotype, male. Scale = 5 mm. b) Left elytron, paratype, female. Scale = 5 mm. c) Labrum, female; d) Head, male; e–f) Aedeagus, left and right lateral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10012587" httpUri="https://zenodo.org/record/10012587/files/figure.png" pageId="24" pageNumber="95">Figs 12e, f</figureCitation>
) in general appearance similar to
<taxonomicName id="4C124D41FFA12E34FCD392E8FC170DAF" authority="Anichtchenko &amp; Wiesner, 2023" authorityName="Anichtchenko &amp; Wiesner" authorityYear="2023" box="[839,945,691,717]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="95" phylum="Arthropoda" rank="species" species="sciakyi" status="sp. nov.">
<emphasis id="B966EAD0FFA12E34FCD392E8FC170DAF" box="[839,945,691,717]" italics="true" pageId="24" pageNumber="95">P. sciakyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25557ABFFA12E34FC2D92E8FBB50DAF" box="[953,1043,691,717]" pageId="24" pageNumber="95" rank="species">
<emphasis id="B966EAD0FFA12E34FC2D92E8FBA80DAF" bold="true" box="[953,1038,691,717]" pageId="24" pageNumber="95">sp. nov</emphasis>
.
</taxonomicNameLabel>
, i.e. median lobe short and robust, strongly widened immediately from the base; ventral side of median lobe, just behind the middle towards to apex, with rounded protrusion, in which the coil of spiral sclerite is situated; apex short and wide, with thick hook; spiral sclerite of endophallus relatively wide, with one and half or two coils at base.
</paragraph>
</subSubSection>
<subSubSection id="C3086549FFA12E34FF539319FBDC0C3F" box="[199,1146,834,861]" pageId="24" pageNumber="95" type="distribution">
<paragraph id="8BAD36C2FFA12E34FF539319FBDC0C3F" blockId="24.[151,1437,150,861]" box="[199,1146,834,861]" pageId="24" pageNumber="95">
<emphasis id="B966EAD0FFA12E34FF539319FEC70C3E" bold="true" box="[199,353,834,860]" pageId="24" pageNumber="95">Distribution.</emphasis>
Known only from
<typeStatus id="54A98860FFA12E34FDA39318FDC10C3F" box="[567,615,835,861]" pageId="24" pageNumber="95">type</typeStatus>
locality in
<collectingRegion id="49D6F820FFA12E34FD729318FC330C3F" box="[742,917,835,861]" country="Philippines" name="Davao Oriental" pageId="24" pageNumber="95">Davao Oriental</collectingRegion>
, Mindanao (
<figureCitation id="13292A47FFA12E34FBB49318FBCF0C3F" box="[1056,1129,835,861]" captionStart="MAP 1" captionStartId="8.[152,211,993,1017]" captionTargetBox="[151,1436,181,970]" captionTargetId="figure-25@8.[151,1436,181,970]" captionTargetPageId="8" captionText="MAP 1. Distribution of species of Prothyma (Symplecthyma)." figureDoi="http://doi.org/10.5281/zenodo.10019734" httpUri="https://zenodo.org/record/10019734/files/figure.png" pageId="24" pageNumber="95">Map 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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