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382 lines
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<mods:titleInfo id="AF7D66EA894152EE3EF85248B1A29E6B">
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<mods:title id="29391C9B2FDD55637D4A6F5AA5B567BC">Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications</mods:title>
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<mods:namePart id="A0CD85557711A72E76F698F29AE101A4">Ivanov, Martin</mods:namePart>
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<mods:namePart id="206966EA50E6EA16E043239091DFA0C3">Čerňanský, Andrej</mods:namePart>
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<mods:namePart id="9BFE7EEAED1DCE1A3B52391DF59A176D">Bonilla-Salomón, Isaac</mods:namePart>
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<mods:namePart id="EE8FFEC1755FA9BBFC04DDCA3BA1068B">Luján, Àngel Hernández</mods:namePart>
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<mods:typeOfResource id="FAFF03B84110341197859517D9F1CCB5">text</mods:typeOfResource>
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<mods:title id="AE38CFFB69500D0A52A1F670A7DF4FCD">Geodiversitas</mods:title>
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<mods:part id="3A7765B9D59EFA213BE25982E1F28810">
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<mods:date id="FDA7E9E51DAA70D7043C512A711CF2FE">2020</mods:date>
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<mods:number id="CD4021840FB145C1937F27F6755488DF">2020-08-28</mods:number>
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<mods:number id="2CDEA3530D7A73F7D67D2E5343E37CF4">42</mods:number>
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<mods:number id="8819BDEB7C745451BBB40FABE3D2EF96">20</mods:number>
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<mods:start id="30E416AFB6BF2D29F6C804F4FDD15FA0">343</mods:start>
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<mods:classification id="2071AAA5ABD8D1F492B754B2AAB55F27">journal article</mods:classification>
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<mods:identifier id="E69F834FBC799167DF5DC3872C892C9F" type="CLB-Dataset">8562</mods:identifier>
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<mods:identifier id="E338EC6841979EA1AE771E69339E1C2A" type="DOI">10.5252/geodiversitas2020v42a20</mods:identifier>
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<mods:identifier id="0A918D201FEE0A9B73A1E12DA636CD8B" type="GBIF-Dataset">ad4dbfb6-14ce-48e2-8fe9-99fa26a9bcf4</mods:identifier>
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<mods:identifier id="E53C904C328347BEE911125E509DC655" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="E47438E3DE064B2FA975A6237EB62CF5" type="Zenodo-Dep">4447563</mods:identifier>
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<treatment id="03C587C7430FFFE9FEC3FB8849C5F91E" ID-DOI="http://doi.org/10.5281/zenodo.4447716" ID-GBIF-Taxon="177200742" ID-Zenodo-Dep="4447716" LSID="urn:lsid:plazi:treatment:03C587C7430FFFE9FEC3FB8849C5F91E" httpUri="http://treatment.plazi.org/id/03C587C7430FFFE9FEC3FB8849C5F91E" lastPageId="13" lastPageNumber="354" pageId="11" pageNumber="352">
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<subSubSection id="C376655A430FFFEFFEC3FB884A41FBFD" box="[383,525,1044,1071]" pageId="11" pageNumber="352" type="nomenclature">
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<paragraph id="8BD336D1430FFFEFFEC3FB884A41FBFD" blockId="11.[383,525,1044,1071]" box="[383,525,1044,1071]" pageId="11" pageNumber="352">
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<heading id="D09B81BD430FFFEFFEC3FB884A41FBFD" box="[383,525,1044,1071]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="352" reason="2">
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<taxonomicName id="4C6C4D52430FFFEFFEC3FB8849A4FBFC" authorityName="Merrem" authorityYear="1820" box="[383,488,1044,1070]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="sp.">
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<emphasis id="B918EAC3430FFFEFFEC3FB8849A4FBFC" box="[383,488,1044,1070]" italics="true" pageId="11" pageNumber="352">Pseudopus</emphasis>
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</taxonomicName>
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sp.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A430FFFEFFE39FBA84A4AFB9C" box="[389,518,1076,1103]" pageId="11" pageNumber="352" type="description">
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<paragraph id="8BD336D1430FFFEFFE39FBA84A4AFB9C" blockId="11.[389,518,1076,1103]" box="[389,518,1076,1103]" pageId="11" pageNumber="352">
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(
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<figureCitation id="13572A54430FFFEFFE32FBA84981FB9D" box="[398,461,1076,1103]" captionStart="FIG" captionStartId="11.[133,144,1917,1935]" captionTargetId="figure-20@12.[151,1436,215,1878]" captionTargetPageId="12" captionText="FIG. 5. — Pseudopus from the early Miocene (MN 4) of MWQ. Pseudopus laurillardi: parietal Pal. 1574 (2/2003 Reptile Joint) in (A) dorsal and (B) ventral aspects; parietal Pal. 1402 (1/2001 Turtle Joint) in (C) dorsal and (D) ventral aspects. Pseudopus sp.: left maxilla Pal. 1575 (2/2003 Reptile Joint) in (E) lateral and (F) medial aspects. Left quadrate Pal. 1576 (2/2003 Reptile Joint) in (G) lateral and (H) medial aspects. Right dentary Pal. 1403 (1/2001 Turtle Joint) in (I) lateral, (J) medial and (K) dorsal aspects. Trunk vertebra Pal. 1404 (1/2001 Turtle Joint) in (L) lateral, (M) dorsal, (N) ventral, (O) anterior and (P) posterior aspects. Trunk vertebra 1405 (1/2001 Turtle Joint) in (Q) anterior aspect. Abbreviations: are, arcuate edge; cc, cephalic condyle; cd, condyle; cf, facet for" figureDoi="http://doi.org/10.5281/zenodo.4487033" httpUri="https://zenodo.org/record/4487033/files/figure.png" pageId="11" pageNumber="352">Fig. 5</figureCitation>
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E-Q)
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A430FFFEFFF37FBF14AFFFB57" box="[139,691,1133,1157]" pageId="11" pageNumber="352" type="reference_group">
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<paragraph id="8BD336D1430FFFEFFF37FBF14AFFFB57" blockId="11.[139,691,1133,1157]" box="[139,691,1133,1157]" pageId="11" pageNumber="352">
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<treatmentCitationGroup id="AB7C11FF430FFFEFFF37FBF14AFFFB57" box="[139,691,1133,1157]" pageId="11" pageNumber="352">
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<taxonomicName id="4C6C4D52430FFFEFFF37FBF148A5FB57" authorityName="Merrem" authorityYear="1820" box="[139,233,1133,1157]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="sp.">
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<emphasis id="B918EAC3430FFFEFFF37FBF148A5FB57" box="[139,233,1133,1157]" italics="true" pageId="11" pageNumber="352">Pseudopus</emphasis>
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</taxonomicName>
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sp. –
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<treatmentCitation id="0ACD10C0430FFFEFFE9FFBF24A4BFB57" author="IVANOV M. & MUSIL R. & BRZOBOHATY R." box="[291,519,1133,1157]" page="229" pageId="11" pageNumber="352" year="2006">
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<bibRefCitation id="EFFD4B20430FFFEFFE9FFBF24A4BFB57" author="IVANOV M. & MUSIL R. & BRZOBOHATY R." box="[291,519,1133,1157]" pageId="11" pageNumber="352" pagination="223 - 239" refId="ref24955" refString="IVANOV M., MUSIL R. & BRZOBOHATY R. 2006. - Terrestrial and Marine Faunas from the Miocene Deposits of the Mokra Plateau (Drahany Upland, Czech Republic) - Impact on Palaeogeography. Beitrage zur Palaontologie 30: 223 - 239." type="journal article" year="2006">
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Ivanov
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<emphasis id="B918EAC3430FFFEFFED7FBF249D6FB57" box="[363,410,1133,1157]" italics="true" pageId="11" pageNumber="352">et al.</emphasis>
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2006: 229
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</bibRefCitation>
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</treatmentCitation>
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, table 2 (in part).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A430FFFEFFF38FB014A0CFAD6" pageId="11" pageNumber="352" type="materials_examined">
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<paragraph id="8BD336D1430FFFEFFF38FB014A0CFAD6" blockId="11.[131,777,1181,1284]" pageId="11" pageNumber="352">
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MATERIAL. —
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<materialsCitation id="3B043C8C430FFFEFFEA3FB014A0CFAD6" ID-GBIF-Occurrence="3027945315" pageId="11" pageNumber="352">
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MWQ, early Miocene, Burdigalian, Orleanian,
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<collectionCode id="ED7DAE14430FFFEFFF3AFB2448FDFB1D" box="[134,177,1208,1231]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="11" pageNumber="352">MN</collectionCode>
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4: maxilla, 1/2001 Turtle Joint: one right dentary (Pal. 1403), three trunk vertebrae (Pal. 1404-1406); 2/2003 Reptile Joint: left maxilla (Pal. 1575), left quadrate (Pal. 1576).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A430FFFE9FF38FAA84ABDFC5D" lastPageId="13" lastPageNumber="354" pageId="11" pageNumber="352" type="description">
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<paragraph id="8BD336D1430FFFEFFF38FAA84959FA9E" blockId="11.[131,777,1332,1740]" box="[132,277,1332,1358]" pageId="11" pageNumber="352">DESCRIPTION</paragraph>
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<paragraph id="8BD336D1430FFFEFFF38FACF489AFABF" blockId="11.[131,777,1332,1740]" box="[132,214,1363,1389]" pageId="11" pageNumber="352">
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<emphasis id="B918EAC3430FFFEFFF38FACF489AFABF" box="[132,214,1363,1389]" italics="true" pageId="11" pageNumber="352">Maxilla</emphasis>
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</paragraph>
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<paragraph id="8BD336D1430FFFEFFF38FAEF4A62F91E" blockId="11.[131,777,1332,1740]" pageId="11" pageNumber="352">Only the posterior region of a left maxilla is partly preserved. It bears four teeth still attached to the bone. An oval superior alveolar foramen is located at the level of the third tooth (counted from posterior). The whole preserved section of the maxilla gradually narrows posteriorly rather than being stepped, but the posterior termination is broken off. Here, a wedge-shaped facet for the jugal is present. It reaches almost to the level of the last preserved tooth. The lateral surface of the preserved portion is pierced by three labial foramina located dorsal to the teeth. The nasal process is only partly preserved, rising dorsally in this region.</paragraph>
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<subSection id="E2E32F3A430FFFEFFF38F96E4C68FC5D" pageId="11" pageNumber="352" type="multiple">
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<paragraph id="8BD336D1430FFFEFFF38F96E48ABF8DE" blockId="11.[132,231,1778,1804]" box="[132,231,1778,1804]" pageId="11" pageNumber="352">
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<heading id="D09B81BD430FFFEFFF38F96E48ABF8DE" box="[132,231,1778,1804]" fontSize="11" level="2" pageId="11" pageNumber="352" reason="8">
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<emphasis id="B918EAC3430FFFEFFF38F96E48ABF8DE" box="[132,231,1778,1804]" italics="true" pageId="11" pageNumber="352">Quadrate</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD336D1430FFFEFFF38F88E4B4AF899" blockId="11.[130,776,1810,1868]" pageId="11" pageNumber="352">The left quadrate is completely preserved, being robustly built. In lateral aspect, its dorsal portion is anteroposteriorly expanded.</paragraph>
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<paragraph id="8BD336D1430FFFEFFC96FF4B4C68FC5D" blockId="11.[810,1457,215,911]" pageId="11" pageNumber="352">
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The dorsal margin of this portion is irregularly rounded.In dorsal aspect, this region is clearly divided by a central constriction into two large areas with roughened surfaces. The posteriorly located one is formed by a large cephalic condyle, having a face for the squamosal on the lateral side. The anterior portion is formed by a massive dorsal tuber (
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<emphasis id="B918EAC3430FFFEFFB34FEEB4CF7FE42" box="[1160,1211,375,400]" italics="true" pageId="11" pageNumber="352">sensu</emphasis>
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<bibRefCitation id="EFFD4B20430FFFEFFB03FEEB4DD3FE43" author="KLEMBARA J. & DOBIAsOVA K. & HAIN M. & YARYHIN O." box="[1215,1439,375,401]" pageId="11" pageNumber="352" pagination="460 - 502" refId="ref25649" refString="KLEMBARA J., DOBIAsOVA K., HAIN M. & YARYHIN O. 2017 a. - Skull anatomy and ontogeny of legless lizard Pseudopus apodus (Pallas, 1775): heterochronic influences on form. The Anatomical Record 300 (3): 460 - 502. https: // doi. org / 10.1002 / ar. 23532" type="journal article" year="2017">
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Klembara
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<emphasis id="B918EAC3430FFFEFFA96FEEB4D16FE43" box="[1322,1370,375,401]" italics="true" pageId="11" pageNumber="352">et al.</emphasis>
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2017a
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</bibRefCitation>
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). Here, the quadrate reaches the highest level dorsally. From the posterior region of this anterior portion, a sharp tympanic crest (
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<emphasis id="B918EAC3430FFFEFFCD7FE4B4BD2FE22" box="[875,926,471,496]" italics="true" pageId="11" pageNumber="352">sensu</emphasis>
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<bibRefCitation id="EFFD4B20430FFFEFFC18FE4A4C7EFE23" author="OELRICH T. M." box="[932,1074,470,497]" pageId="11" pageNumber="352" pagination="1 - 122" refId="ref26175" refString="OELRICH T. M. 1956. - The anatomy of the head of Ctenosaura pectinata (Iguanidae). Miscellaneous Publications, Museum of Zoology, University of Michigan 94: 1 - 122. http: // hdl. handle. net / 2027.42 / 56338" type="journal article" year="1956">Oelrich 1956</bibRefCitation>
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) runs along the entire central region ventrally, being laterally expanded. In the dorsal region, a short anterolateral crest (
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<emphasis id="B918EAC3430FFFEFFC4CFD8B4C6FFDE2" box="[1008,1059,535,560]" italics="true" pageId="11" pageNumber="352">sensu</emphasis>
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<bibRefCitation id="EFFD4B20430FFFEFFB9BFD8A4D44FDE2" author="KLEMBARA J. & DOBIAsOVA K. & HAIN M. & YARYHIN O." box="[1063,1288,534,560]" pageId="11" pageNumber="352" pagination="460 - 502" refId="ref25649" refString="KLEMBARA J., DOBIAsOVA K., HAIN M. & YARYHIN O. 2017 a. - Skull anatomy and ontogeny of legless lizard Pseudopus apodus (Pallas, 1775): heterochronic influences on form. The Anatomical Record 300 (3): 460 - 502. https: // doi. org / 10.1002 / ar. 23532" type="journal article" year="2017">
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Klembara
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<emphasis id="B918EAC3430FFFEFFB2EFD8B4C8FFDE2" box="[1170,1219,534,560]" italics="true" pageId="11" pageNumber="352">et al.</emphasis>
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2017a
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</bibRefCitation>
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) is located anterior to the tympanic crest. A small depression divides these two structures. The posterior region of quadrate is bordered by a markedly rounded (concave) posterior crest. In anterior view, the mandibular condyle is mediolaterally broad.Its ventral side is shallow, W-shaped with blunt peaks. On the medial surface of the quadrate, there is a prominent medial crest (
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|
<emphasis id="B918EAC3430FFFEFFA95FD4A4D17FD3D" box="[1321,1371,726,751]" italics="true" pageId="11" pageNumber="352">sensu</emphasis>
|
|||
|
|
<bibRefCitation id="EFFD4B20430FFFEFFADCFD494B2FFCC2" author="OELRICH T. M." pageId="11" pageNumber="352" pagination="1 - 122" refId="ref26175" refString="OELRICH T. M. 1956. - The anatomy of the head of Ctenosaura pectinata (Iguanidae). Miscellaneous Publications, Museum of Zoology, University of Michigan 94: 1 - 122. http: // hdl. handle. net / 2027.42 / 56338" type="journal article" year="1956">Oelrich 1956</bibRefCitation>
|
|||
|
|
). Its ventral portion protrudes medially slightly more than the dorsal portion, as a result of bearing a distinct facet for the quadrate process of the pterygoid. A small, rounded quadrate foramen pierces the quadrate at approximately the mid-length of the posterior surface.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSection>
|
|||
|
|
<subSection id="E2E32F3A430FFFEFFC90FC294DE8FA9F" pageId="11" pageNumber="352" type="multiple">
|
|||
|
|
<paragraph id="8BD336D1430FFFEFFC90FC294BCDFC1C" blockId="11.[812,897,949,974]" box="[812,897,949,974]" pageId="11" pageNumber="352">
|
|||
|
|
<heading id="D09B81BD430FFFEFFC90FC294BCDFC1C" box="[812,897,949,974]" fontSize="11" level="2" pageId="11" pageNumber="352" reason="8">
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFC90FC294BCDFC1C" box="[812,897,949,974]" italics="true" pageId="11" pageNumber="352">Dentary</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD336D1430FFFEFFC91FC484DE8FA9F" blockId="11.[810,1458,980,1357]" pageId="11" pageNumber="352">
|
|||
|
|
The body of the right dentary (Pal. 1403) is slightly concave. However, this element is badly preserved, especially on the medial side.Here, the almost entire region of the subdental shelf (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFC89FBA94B24FB9C" box="[821,872,1077,1102]" italics="true" pageId="11" pageNumber="352">sensu</emphasis>
|
|||
|
|
<bibRefCitation id="EFFD4B20430FFFEFFCCDFBA94C0DFB9C" author="RAGE J. - C. & AUGE M. L." box="[881,1089,1076,1103]" pageId="11" pageNumber="352" pagination="253 - 268" refId="ref26761" refString="RAGE J. - C. & AUGE M. L. 2010. - Squamate reptiles from the middle Eocene of Lissieu (France). A landmark in the middle Eocene of Europe. Geobios 43 (2): 253 - 268. https: // doi. org / 10.1016 / j. geobios. 2009.08.002" type="journal article" year="2010">Rage & Augé 2010</bibRefCitation>
|
|||
|
|
) and the intramandibular septum are broken off and missing. For this reason, the alveolar canal is exposed. In the posterior region, only the splenial articulation and the coronoid articulation are partly preserved. They are separated from each other by a small gap formed by a bony ridge. As preserved,the dentary possesses twelve tooth positions, but only one tooth is still attached (the actual tooth number of the complete tooth row was probably slightly higher). The lateral surface of the bone is pierced by four labial foramina.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSection>
|
|||
|
|
<subSection id="E2E32F3A430FFFEFFC90FAE84C64F91E" pageId="11" pageNumber="352" type="multiple">
|
|||
|
|
<paragraph id="8BD336D1430FFFEFFC90FAE84BDDFA5F" blockId="11.[812,913,1396,1421]" box="[812,913,1396,1421]" pageId="11" pageNumber="352">
|
|||
|
|
<heading id="D09B81BD430FFFEFFC90FAE84BDDFA5F" box="[812,913,1396,1421]" fontSize="11" level="2" pageId="11" pageNumber="352" reason="8">
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFC90FAE84BDDFA5F" box="[812,913,1396,1421]" italics="true" pageId="11" pageNumber="352">Dentition</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD336D1430FFFEFFC91FA0F4C64F91E" blockId="11.[811,1457,1427,1740]" pageId="11" pageNumber="352">A pleurodont tooth implantation is present.Teeth are amblyodont, being enlarged posteriorly to form blunt robust cylinders. No striations can be observed on the tooth crowns (this could be due to poor preservation or wearing of the occlusal surfaces). The penultimate maxillary tooth has a large tooth crown with a distinctly constricted neck. Thus, this tooth is mushroom-like shape. The tooth bases of maxillary teeth are ankylosed to their sockets and surrounded by a spongy (porous) bony tissue. The tooth crown of the dentary tooth is extremely weathered.</paragraph>
|
|||
|
|
</subSection>
|
|||
|
|
<subSection id="E2E32F3A430FFFE9FC91F96E4ABDFC5D" lastPageId="13" lastPageNumber="354" pageId="11" pageNumber="352" type="multiple">
|
|||
|
|
<paragraph id="8BD336D1430FFFEFFC91F96E4B9EF8DE" blockId="11.[813,978,1778,1804]" box="[813,978,1778,1804]" pageId="11" pageNumber="352">
|
|||
|
|
<heading id="D09B81BD430FFFEFFC91F96E4B9EF8DE" box="[813,978,1778,1804]" fontSize="11" level="2" pageId="11" pageNumber="352" reason="8">
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFC91F96E4B9EF8DE" box="[813,978,1778,1804]" italics="true" pageId="11" pageNumber="352">Trunk vertebrae</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD336D1430FFFEFFC91F88E4DE3F899" blockId="11.[813,1456,1810,1868]" pageId="11" pageNumber="352">Several trunk vertebrae are present. They are robustly built. The neural spine starts to rise dorsally approximately at the</paragraph>
|
|||
|
|
<caption id="DF136659430FFFEFFF39F8E14DE3F83B" ID-DOI="http://doi.org/10.5281/zenodo.4487033" ID-Zenodo-Dep="4487033" httpUri="https://zenodo.org/record/4487033/files/figure.png" pageId="11" pageNumber="352" targetBox="[151,1436,215,1878]" targetPageId="12">
|
|||
|
|
<paragraph id="8BD336D1430FFFEFFF39F8E14DE3F83B" blockId="11.[132,1455,1916,2025]" pageId="11" pageNumber="352">
|
|||
|
|
FIG. 5. —
|
|||
|
|
<taxonomicName id="4C6C4D52430FFFEFFF6BF8E1497BF85D" authorityName="Merrem" authorityYear="1820" box="[215,311,1917,1935]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFF6BF8E1497BF85D" box="[215,311,1917,1935]" italics="true" pageId="11" pageNumber="352">Pseudopus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from the early Miocene (MN 4) of MWQ.
|
|||
|
|
<taxonomicName id="4C6C4D52430FFFEFFD29F8E14B0AF85D" baseAuthorityName="Lartet" baseAuthorityYear="1851" box="[661,838,1917,1935]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="laurillardi">
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFD29F8E14B0AF85D" box="[661,838,1917,1935]" italics="true" pageId="11" pageNumber="352">Pseudopus laurillardi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
: parietal Pal. 1574 (2/2003 Reptile Joint) in (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFB7DF8E14C82F85D" bold="true" box="[1217,1230,1917,1935]" pageId="11" pageNumber="352">A</emphasis>
|
|||
|
|
) dorsal and (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFA81F8E14D06F85D" bold="true" box="[1341,1354,1917,1935]" pageId="11" pageNumber="352">B</emphasis>
|
|||
|
|
) ventral aspects; parietal Pal. 1402 (1/2001 Turtle Joint) in (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFD99F80F4A7FF877" bold="true" box="[549,563,1939,1957]" pageId="11" pageNumber="352">C</emphasis>
|
|||
|
|
) dorsal and (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFD1FF80F4AFDF877" bold="true" box="[675,689,1939,1957]" pageId="11" pageNumber="352">D</emphasis>
|
|||
|
|
) ventral aspects.
|
|||
|
|
<taxonomicName id="4C6C4D52430FFFEFFCF6F80F4BE6F877" authorityName="Merrem" authorityYear="1820" box="[842,938,1939,1957]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="352" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFCF6F80F4BE6F877" box="[842,938,1939,1957]" italics="true" pageId="11" pageNumber="352">Pseudopus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp.: left maxilla Pal. 1575 (2/2003 Reptile Joint) in (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFAD8F80F4D3CF877" bold="true" box="[1380,1392,1939,1957]" pageId="11" pageNumber="352">E</emphasis>
|
|||
|
|
) lateral and (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFF13F83648F6F86E" bold="true" box="[175,186,1962,1980]" pageId="11" pageNumber="352">F</emphasis>
|
|||
|
|
) medial aspects. Left quadrate Pal. 1576 (2/2003 Reptile Joint) in (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFD46F8364B44F86E" bold="true" box="[762,776,1962,1980]" pageId="11" pageNumber="352">G</emphasis>
|
|||
|
|
) lateral and (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFCC5F8364BCBF86E" bold="true" box="[889,903,1962,1980]" pageId="11" pageNumber="352">H</emphasis>
|
|||
|
|
) medial aspects. Right dentary Pal. 1403 (1/2001 Turtle Joint) in (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFF35F85D48C3F801" bold="true" box="[137,143,1985,2003]" pageId="11" pageNumber="352">I</emphasis>
|
|||
|
|
) lateral, (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFF62F85D48A4F801" bold="true" box="[222,232,1985,2003]" pageId="11" pageNumber="352">J</emphasis>
|
|||
|
|
) medial and (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFEE1F85D4926F801" bold="true" box="[349,362,1985,2003]" pageId="11" pageNumber="352">K</emphasis>
|
|||
|
|
) dorsal aspects. Trunk vertebra Pal. 1404 (1/2001 Turtle Joint) in (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFC1DF85D4BE0F801" bold="true" box="[929,940,1985,2003]" pageId="11" pageNumber="352">L</emphasis>
|
|||
|
|
) lateral, (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFC47F85D4C40F801" bold="true" box="[1019,1036,1985,2003]" pageId="11" pageNumber="352">M</emphasis>
|
|||
|
|
) dorsal, (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFBE0F85D4C26F801" bold="true" box="[1116,1130,1985,2003]" pageId="11" pageNumber="352">N</emphasis>
|
|||
|
|
) ventral, (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFB03F85C4C82F800" bold="true" box="[1215,1230,1984,2002]" pageId="11" pageNumber="352">O</emphasis>
|
|||
|
|
) anterior and (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFAF7F85D4D1BF801" bold="true" box="[1355,1367,1985,2003]" pageId="11" pageNumber="352">P</emphasis>
|
|||
|
|
) posterior aspects. Trunk vertebra 1405 (1/2001 Turtle Joint) in (
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFDECF84B4A13F83B" bold="true" box="[592,607,2007,2025]" pageId="11" pageNumber="352">Q</emphasis>
|
|||
|
|
) anterior aspect. Abbreviations:
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFCC4F84B4BD9F83B" bold="true" box="[888,917,2007,2025]" pageId="11" pageNumber="352">are</emphasis>
|
|||
|
|
, arcuate edge;
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFBA0F84B4C7EF83B" bold="true" box="[1052,1074,2007,2025]" pageId="11" pageNumber="352">cc</emphasis>
|
|||
|
|
, cephalic condyle;
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFB66F84B4CBCF83B" bold="true" box="[1242,1264,2007,2025]" pageId="11" pageNumber="352">cd</emphasis>
|
|||
|
|
, condyle;
|
|||
|
|
<emphasis id="B918EAC3430FFFEFFAF7F84B4D10F83B" bold="true" box="[1355,1372,2007,2025]" pageId="11" pageNumber="352">cf</emphasis>
|
|||
|
|
, facet for
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BD336D14308FFE8FF38F8E14A4CF83B" blockId="12.[132,1456,1917,2025]" pageId="12" pageNumber="353">
|
|||
|
|
coronoid;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FF61F8E148B7F85D" bold="true" box="[221,251,1917,1935]" pageId="12" pageNumber="353">cpr</emphasis>
|
|||
|
|
, cranial parietal ridge;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FE77F8E14991F85D" bold="true" box="[459,477,1917,1935]" pageId="12" pageNumber="353">ct</emphasis>
|
|||
|
|
, cotyle;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FD95F8E14A77F85D" bold="true" box="[553,571,1917,1935]" pageId="12" pageNumber="353">dt</emphasis>
|
|||
|
|
, dorsal tuber;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FD01F8E14A83F85D" bold="true" box="[701,719,1917,1935]" pageId="12" pageNumber="353">fo</emphasis>
|
|||
|
|
, foramen;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FC92F8E14B09F85D" bold="true" box="[814,837,1917,1935]" pageId="12" pageNumber="353">fst</emphasis>
|
|||
|
|
, facet for supratemporal;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FB96F8E14C79F85D" bold="true" box="[1066,1077,1917,1935]" pageId="12" pageNumber="353">lf</emphasis>
|
|||
|
|
, labial foramina;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FB70F8E14CBFF85D" bold="true" box="[1228,1267,1917,1935]" pageId="12" pageNumber="353">mac</emphasis>
|
|||
|
|
, mandibular condyle;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FF39F80F48EDF877" bold="true" box="[133,161,1939,1957]" pageId="12" pageNumber="353">mc</emphasis>
|
|||
|
|
, medial crest;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FE9CF80F4977F877" bold="true" box="[288,315,1939,1957]" pageId="12" pageNumber="353">ms</emphasis>
|
|||
|
|
, muscular surface;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FE59F80F49B7F877" bold="true" box="[485,507,1939,1957]" pageId="12" pageNumber="353">nc</emphasis>
|
|||
|
|
, neural canal;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FDC6F80F4AC3F877" bold="true" box="[634,655,1939,1957]" pageId="12" pageNumber="353">ns</emphasis>
|
|||
|
|
, neural spine;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FCB2F80F4B69F877" bold="true" box="[782,805,1939,1957]" pageId="12" pageNumber="353">np</emphasis>
|
|||
|
|
, nasal process;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FC0EF80F4B9FF877" bold="true" box="[946,979,1939,1957]" pageId="12" pageNumber="353">ocs</emphasis>
|
|||
|
|
, occipital shield;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FBD6F80F4CC7F877" bold="true" box="[1130,1163,1939,1957]" pageId="12" pageNumber="353">pas</emphasis>
|
|||
|
|
, parietal shield;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FAA5F80F4D67F877" bold="true" box="[1305,1323,1939,1957]" pageId="12" pageNumber="353">pf</emphasis>
|
|||
|
|
, parietal fossa;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FF39F83648EEF86E" bold="true" box="[133,162,1962,1980]" pageId="12" pageNumber="353">pfc</emphasis>
|
|||
|
|
, postfoveal crest;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FEF9F8364912F86E" bold="true" box="[325,350,1962,1980]" pageId="12" pageNumber="353">prf</emphasis>
|
|||
|
|
, prezygapophyseal articular facet;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FD2EF8364AF8F86E" bold="true" box="[658,692,1962,1980]" pageId="12" pageNumber="353">poc</emphasis>
|
|||
|
|
, posterior crest;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FCF7F8364B25F86E" bold="true" box="[843,873,1962,1980]" pageId="12" pageNumber="353">pof</emphasis>
|
|||
|
|
, postzygapophyseal articular facet;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FB1BF8364CF0F86E" bold="true" box="[1191,1212,1962,1980]" pageId="12" pageNumber="353">sa</emphasis>
|
|||
|
|
, smooth area;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FAFDF8364D11F86E" bold="true" box="[1345,1373,1962,1980]" pageId="12" pageNumber="353">saf</emphasis>
|
|||
|
|
, superior alveolar foramen;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FEA3F85D497CF801" bold="true" box="[287,304,1985,2003]" pageId="12" pageNumber="353">sf</emphasis>
|
|||
|
|
, facet for splenial;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FE6AF85D49BEF801" bold="true" box="[470,498,1985,2003]" pageId="12" pageNumber="353">srs</emphasis>
|
|||
|
|
, supradental shelf;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FD22F85D4AF7F801" bold="true" box="[670,699,1985,2003]" pageId="12" pageNumber="353">stp</emphasis>
|
|||
|
|
, supratemporal process;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FC24F85D4BFBF801" bold="true" box="[920,951,1985,2003]" pageId="12" pageNumber="353">syn</emphasis>
|
|||
|
|
, synapophysis;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FBF8F85D4C1AF801" bold="true" box="[1092,1110,1985,2003]" pageId="12" pageNumber="353">tc</emphasis>
|
|||
|
|
, tympanic crest;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FB50F85D4D4EF801" bold="true" box="[1260,1282,1985,2003]" pageId="12" pageNumber="353">vlr</emphasis>
|
|||
|
|
, ventrolateral ridge;
|
|||
|
|
<emphasis id="B918EAC34308FFE8FF39F84B48D2F83B" bold="true" box="[133,158,2007,2025]" pageId="12" pageNumber="353">vls</emphasis>
|
|||
|
|
, ventrolateral surface. Scale bars:
|
|||
|
|
<quantity id="4C949B344308FFE8FE75F84B49B0F83B" box="[457,508,2007,2025]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="353" unit="mm" value="2.0">2 mm</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BD336D14309FFE9FF38FF4B4ABDFC5D" blockId="13.[130,778,215,911]" pageId="13" pageNumber="354">
|
|||
|
|
mid-length of the neural arch. However, this structure is damaged in all vertebrae, thus its exact shape is unknown. In dorsal view, the neural spine widens gradually slightly in a posterior direction and, based on the preserved portion, it probably did not exceed the condyle posteriorly. Anteriorly, a low medial ridge runs to the end of the neural arch, where it forms a small spike. The oval neural canal is small. Pre- and postzygapophyses are well expanded laterally. The interzygapophyseal constriction is well developed and deep. The prezygapophyses are dorsally inclined in an angle of around 30°. The articular surfaces are large, elliptical and oblong laterally. In anterior aspect, a pair of foramina is located ventral to the prezygapophyses, close to the neural canal (
|
|||
|
|
<figureCitation id="13572A544309FFE9FD1EFDCA4ABAFDA2" box="[674,758,598,625]" captionStart="FIG" captionStartId="11.[133,144,1917,1935]" captionTargetId="figure-20@12.[151,1436,215,1878]" captionTargetPageId="12" captionText="FIG. 5. — Pseudopus from the early Miocene (MN 4) of MWQ. Pseudopus laurillardi: parietal Pal. 1574 (2/2003 Reptile Joint) in (A) dorsal and (B) ventral aspects; parietal Pal. 1402 (1/2001 Turtle Joint) in (C) dorsal and (D) ventral aspects. Pseudopus sp.: left maxilla Pal. 1575 (2/2003 Reptile Joint) in (E) lateral and (F) medial aspects. Left quadrate Pal. 1576 (2/2003 Reptile Joint) in (G) lateral and (H) medial aspects. Right dentary Pal. 1403 (1/2001 Turtle Joint) in (I) lateral, (J) medial and (K) dorsal aspects. Trunk vertebra Pal. 1404 (1/2001 Turtle Joint) in (L) lateral, (M) dorsal, (N) ventral, (O) anterior and (P) posterior aspects. Trunk vertebra 1405 (1/2001 Turtle Joint) in (Q) anterior aspect. Abbreviations: are, arcuate edge; cc, cephalic condyle; cd, condyle; cf, facet for" figureDoi="http://doi.org/10.5281/zenodo.4487033" httpUri="https://zenodo.org/record/4487033/files/figure.png" pageId="13" pageNumber="354">Fig. 5O</figureCitation>
|
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|
|
). In lateral aspect, the postzygapophyses reach posteriorly approximately to the level of the mid-region of the condyle. Both condyle and cotyle are markedly depressed. The cotyle has an anteroventral orientation, whereas the condyle is posterodorsally oriented and they lack a precondylar constriction. The synapophyses are well-developed in the anterior region, being slightly posterolaterally oriented. The ventral region of the centrum is flat and the subcentral ridges are straight and gradually converge from anterior to posterior region.
|
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</paragraph>
|
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|
</subSection>
|
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</subSubSection>
|
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<subSubSection id="C376655A4309FFE9FF38FC2949C5F91E" pageId="13" pageNumber="354" type="discussion">
|
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|
|
<paragraph id="8BD336D14309FFE9FF38FC2948A5FC1F" blockId="13.[130,777,949,1740]" box="[132,233,949,975]" pageId="13" pageNumber="354">REMARKS</paragraph>
|
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<paragraph id="8BD336D14309FFE9FF38FC4949C5F91E" blockId="13.[130,777,949,1740]" pageId="13" pageNumber="354">
|
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|
|
In the posterior region of the dentary (Pal. 1403), splenial articulations is present, located anterior to the coronoid articulation. In modern
|
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|
|
<taxonomicName id="4C6C4D524309FFE9FEC6FB894998FBFC" baseAuthorityName="Pallas" baseAuthorityYear="1775" box="[378,468,1044,1070]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="apodus">
|
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|
|
<emphasis id="B918EAC34309FFE9FEC6FB894998FBFC" box="[378,468,1044,1070]" italics="true" pageId="13" pageNumber="354">P.apodus</emphasis>
|
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|
|
</taxonomicName>
|
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|
|
, the coronoid extends further anteriorly and there is no facet for the splenial in this region (e.g.,
|
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|
|
<bibRefCitation id="EFFD4B204309FFE9FF02FBC849D9FBBC" author="KLEMBARA J. & BOHME M. & RUMMEL M." box="[190,405,1108,1134]" pageId="13" pageNumber="354" pagination="159 - 196" refId="ref25523" refString="KLEMBARA J., BOHME M. & RUMMEL M. 2010. - Revision of the anguine lizard Pseudopus laurillardi (Squamata, Anguidae) from the Miocene of Europe, with comments on paleoecology. Journal of Paleontology 84 (2): 159 - 196. https: // doi. org / 10.1666 / 09 - 033 R 1.1" type="journal article" year="2010">
|
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|
|
Klembara
|
|||
|
|
<emphasis id="B918EAC34309FFE9FE95FBC94916FBBC" box="[297,346,1108,1134]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
|
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|
|
2010
|
|||
|
|
</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFFD4B204309FFE9FE1DFBC84994FBBC" author="KLEMBARA J. & HAIN M. & DOBIAsOVA K." box="[417,472,1108,1134]" pageId="13" pageNumber="354" pagination="516 - 544" refId="ref25586" refString="KLEMBARA J., HAIN M. & DOBIAsOVA K. 2014. - Comparative anatomy of the lower jaw and dentition of Pseudopus apodus and the interrelationships of species of subfamily Anguinae (Anguimorpha, Anguidae). The Anatomical Record 297 (3): 516 - 544. https: // doi. org / 10.1002 / ar. 22854" type="journal article" year="2014">2014</bibRefCitation>
|
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|
). However, the same dentary features seen in the dentary from Mokrá, can be observed in
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FF3FFB09496BFB7C" authorityName="Klembara" authorityYear="2012" box="[131,295,1172,1198]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="ahnikoviensis">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FF3FFB09496BFB7C" box="[131,295,1172,1198]" italics="true" pageId="13" pageNumber="354">P. ahnikoviensis</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FEE0FB0949ADFB7C" baseAuthorityName="Lartet" baseAuthorityYear="1851" box="[348,481,1172,1198]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="laurillardi">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FEE0FB0949ADFB7C" box="[348,481,1172,1198]" italics="true" pageId="13" pageNumber="354">P. laurillardi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Unfortunately, the medial side of the dentary is badly preserved and lacks most of the diagnostic features.
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FEDFFB484A44FB3C" authorityName="Klembara" authorityYear="2012" box="[355,520,1236,1262]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="ahnikoviensis">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FEDFFB484A44FB3C" box="[355,520,1236,1262]" italics="true" pageId="13" pageNumber="354">P. ahnikoviensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is characterized by the presence of a surangular spine, whereas an autapomorphic feature of
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FF40FA8849CBFAFF" baseAuthorityName="Lartet" baseAuthorityYear="1851" box="[252,391,1299,1325]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="laurillardi">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FF40FA8849CBFAFF" box="[252,391,1299,1325]" italics="true" pageId="13" pageNumber="354">P. laurillardi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is a medially expanded subdental shelf (
|
|||
|
|
<bibRefCitation id="EFFD4B204309FFE9FF7BFAAF49E6FA9F" author="KLEMBARA J. & BOHME M. & RUMMEL M." box="[199,426,1331,1357]" pageId="13" pageNumber="354" pagination="159 - 196" refId="ref25523" refString="KLEMBARA J., BOHME M. & RUMMEL M. 2010. - Revision of the anguine lizard Pseudopus laurillardi (Squamata, Anguidae) from the Miocene of Europe, with comments on paleoecology. Journal of Paleontology 84 (2): 159 - 196. https: // doi. org / 10.1666 / 09 - 033 R 1.1" type="journal article" year="2010">
|
|||
|
|
Klembara
|
|||
|
|
<emphasis id="B918EAC34309FFE9FE8BFAA84927FA9F" box="[311,363,1331,1357]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
|
|||
|
|
2010
|
|||
|
|
</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFFD4B204309FFE9FE04FAAF4AD0FA9F" author="KLEMBARA J. & HAIN M. & DOBIAsOVA K." box="[440,668,1331,1357]" pageId="13" pageNumber="354" pagination="516 - 544" refId="ref25586" refString="KLEMBARA J., HAIN M. & DOBIAsOVA K. 2014. - Comparative anatomy of the lower jaw and dentition of Pseudopus apodus and the interrelationships of species of subfamily Anguinae (Anguimorpha, Anguidae). The Anatomical Record 297 (3): 516 - 544. https: // doi. org / 10.1002 / ar. 22854" type="journal article" year="2014">
|
|||
|
|
Klembara
|
|||
|
|
<emphasis id="B918EAC34309FFE9FD95FAA84A11FA9F" box="[553,605,1331,1357]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
|
|||
|
|
2014
|
|||
|
|
</bibRefCitation>
|
|||
|
|
). Neither of those features can be demonstrated on the dentary from Mokrá and we decided to allocate it only to
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FDCBFAEF4AACFA5F" authorityName="Merrem" authorityYear="1820" box="[631,736,1395,1421]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FDCBFAEF4AACFA5F" box="[631,736,1395,1421]" italics="true" pageId="13" pageNumber="354">Pseudopus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. The morphology of the quadrate strongly corresponds to that of
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FF65FA2F490EFA1F" authorityName="Merrem" authorityYear="1820" box="[217,322,1459,1485]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FF65FA2F490EFA1F" box="[217,322,1459,1485]" italics="true" pageId="13" pageNumber="354">Pseudopus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(for
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FEC7FA284990FA1F" baseAuthorityName="Pallas" baseAuthorityYear="1775" box="[379,476,1459,1485]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="apodus">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FEC7FA284990FA1F" box="[379,476,1459,1485]" italics="true" pageId="13" pageNumber="354">P. apodus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
see
|
|||
|
|
<bibRefCitation id="EFFD4B204309FFE9FDB2FA2F4B4CFA1F" author="KLEMBARA J. & DOBIAsOVA K. & HAIN M. & YARYHIN O." box="[526,768,1459,1485]" pageId="13" pageNumber="354" pagination="460 - 502" refId="ref25649" refString="KLEMBARA J., DOBIAsOVA K., HAIN M. & YARYHIN O. 2017 a. - Skull anatomy and ontogeny of legless lizard Pseudopus apodus (Pallas, 1775): heterochronic influences on form. The Anatomical Record 300 (3): 460 - 502. https: // doi. org / 10.1002 / ar. 23532" type="journal article" year="2017">
|
|||
|
|
Klembara
|
|||
|
|
<emphasis id="B918EAC34309FFE9FDC2FA284AF8FA1F" box="[638,692,1459,1485]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
|
|||
|
|
2017a
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: fig. 27) and this specimen most likely belongs to this taxon. The trunk vertebrae described in this section possess two features characteristic for
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FE28F98E4B4EF9FF" authority="(Cernansky et al. 2019)" baseAuthorityName="Cernansky" baseAuthorityYear="2019" box="[404,770,1554,1584]" class="Reptilia" family="Anguidae" genus="Pseudopus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FE28F98E49B1F9FE" box="[404,509,1554,1580]" italics="true" pageId="13" pageNumber="354">Pseudopus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFFD4B204309FFE9FDB1F9884ABAF9FF" author="CERNANSKY A. & YARYHIN O. & CICEKOVA J. & WERNEBURG I. & HAIN M. & KLEMBARA J." box="[525,758,1554,1584]" pageId="13" pageNumber="354" pagination="232 - 257" refId="ref23394" refString="CERNANSKY A., YARYHIN O., CICEKOVA J., WERNEBURG I., HAIN M. & KLEMBARA J. 2019. - Vertebral Comparative Anatomy and Morphological Differencesin Anguine Lizards with a Special Reference to Pseudopus apodus. The Anatomical Record 302 (2): 232 - 257. https: // doi. org / 10.1002 / ar. 23944" type="journal article" year="2019">
|
|||
|
|
Čerňanský
|
|||
|
|
<emphasis id="B918EAC34309FFE9FD3BF98F4AF5F9FE" box="[647,697,1554,1580]" italics="true" pageId="13" pageNumber="354">et al.</emphasis>
|
|||
|
|
2019
|
|||
|
|
</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
: 1) the lateral margins of the centrum (subcentral ridges) are straight rather than concave (
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FE6CF9CE4A05F9BE" authorityName="Daudin" authorityYear="1803" box="[464,585,1618,1644]" class="Reptilia" family="Anguidae" genus="Ophisaurus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FE6CF9CE4A05F9BE" box="[464,585,1618,1644]" italics="true" pageId="13" pageNumber="354">Ophisaurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) or parallel with their anterior-most sections being expanded laterally (
|
|||
|
|
<taxonomicName id="4C6C4D524309FFE9FD0FF9EF4AB4F95F" box="[691,760,1651,1677]" class="Reptilia" family="Anguidae" genus="Anguis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="B918EAC34309FFE9FD0FF9EF4AB4F95F" box="[691,760,1651,1677]" italics="true" pageId="13" pageNumber="354">Anguis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
); and 2) the height of the neural canal is distinctly less than the height of the cotyle.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|