<documentid="9D2CE68279B711153EFA9597B4F25946"ID-CLB-Dataset="8134"ID-DOI="10.11646/zootaxa.4927.1.2"ID-GBIF-Dataset="f8e774a8-36c4-482c-8d53-adc4612ea281"ID-ISSN="1175-5326"ID-Zenodo-Dep="4533849"ID-ZooBank="E4C28E67-3072-4023-BD47-C2FD9CECAAE5"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="admin"IM.taxonomicNames_approvedBy="admin"checkinTime="1613034223463"checkinUser="plazi"docAuthor="Lyu, Zhi-Tong, Zeng, Zhao-Chi, Wang, Jian, Liu, Zu-Yao, Huang, Ya-Qiong, Li, Wen-Zhou & Wang, Ying-Yong"docDate="2021"docId="03B087AFFFAFFFB4628EFC3EFDCFD163"docLanguage="en"docName="zootaxa.4927.1.2.pdf"docOrigin="Zootaxa 4927 (1)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Panophrys sanmingensis Lyu & Zeng & Wang & Liu & Huang & Li & Wang 2021, sp. nov."docType="treatment"docVersion="10"lastPageNumber="31"masterDocId="FF89FFD7FFBCFFA26219FFBFFF8DD303"masterDocTitle="Four new species of Panophrys (Anura, Megophryidae) from eastern China, with discussion on the recognition of Panophrys as a distinct genus"masterLastPageNumber="40"masterPageNumber="9"pageNumber="28"updateTime="1720383473109"updateUser="admin"zenodo-license-document="CLOSED">
<mods:titleid="7B8E2E77BDC8359493FAC471D4B304BC">Four new species of Panophrys (Anura, Megophryidae) from eastern China, with discussion on the recognition of Panophrys as a distinct genus</mods:title>
<mods:affiliationid="8FD912EDD0E60190189CBC1012E8D9A3">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & lvzht @ foxmail. com; https: // orcid. org / 0000 - 0002 - 4891 - 3859</mods:affiliation>
<mods:affiliationid="51DD64038CF834488BC8C72626D9CBF5">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Shenzhen Shuanghuamu Biological Technology Co., Ltd, Shenzhen 51800, PR China & zhaochizeng @ outlook. com; https: // orcid. org / 0000 - 0003 - 4054 - 8399</mods:affiliation>
<mods:affiliationid="3A68002431D78C96EDA787AED1A63B26">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Shenzhen Shuanghuamu Biological Technology Co., Ltd, Shenzhen 51800, PR China & wangj 397 @ mail. sysu. edu. cn; https: // orcid. org / 0000 - 0003 - 4249 - 7767</mods:affiliation>
<mods:affiliationid="F4B13CD2244BE4130B24A60D2797ED6D">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Institute of Ecology and Evolution, University of Bern, Bern 3012, Switzerland.</mods:affiliation>
<mods:affiliationid="75F1B619B221FBA6992CD24FEAD790ED">Fujian Daiyun Mountain National Nature Reserve, Quanzhou 362500, PR China & Fujian Wildlife Conservation Center, Fujian Provincial Department of Forestry, Fuzhou 350003, PR China</mods:affiliation>
<mods:affiliationid="4D238DD13288737F488FA0917D101FD3">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
<taxonomicNameid="4C194D3AFFAFFFB1628EFC3EFDE0D09F"authority="Lyu & Wang"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[151,621,897,924]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFAFFFB16335FC79FE13D0DD"ID-CoL="5N6Z"authorityName="Kuhl & Van Hasselt"authorityYear="1822"box="[300,414,966,990]"class="Amphibia"family="Megophryidae"genus="Megophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="genus">
<bibRefCitationid="EF884B48FFAFFFB163F5FC79FD0ED0DD"author="Liu, Z. Y. & Chen, G. L. & Zhu, T. Q. & Zeng, Z. C. & Lyu, Z. T. & Wang, J. & Messenger, K. & Greenberg, A. J. & Guo, Z. X. & Yang, Z. H. & Shi, S. H. & Wang, Y. Y."box="[492,643,966,990]"pageId="19"pageNumber="28"pagination="723 - 731"refId="ref18211"refString="Liu, Z. Y., Chen, G. L., Zhu, T. Q., Zeng, Z. C., Lyu, Z. T., Wang, J., Messenger, K., Greenberg, A. J., Guo, Z. X., Yang, Z. H., Shi, S. H. & Wang, Y. Y. (2018) Prevalence of cryptic species in morphologically uniform taxa-Fast speciation and evolutionary radiation in Asian frogs. Molecular Phylogenetics and Evolution, 127, 723 - 731. https: // doi. org / 10.1016 / j. ympev. 2018.06.020"type="journal article"year="2018">
<figureCitationid="13222A3CFFAFFFB163A2FBB3FE71D724"box="[443,508,1036,1063]"captionStart="FIGURE 5"captionStartId="21.[151,250,1866,1890]"captionTargetBox="[151,1436,181,1842]"captionTargetId="figure-20@21.[151,1436,181,1842]"captionTargetPageId="21"captionText="FIGURE 5. (A)–(D) dorsolateral view, ventral view, right hand, and right foot of adult male holotype SYS a002498 of Panophrys sanmingensis sp. nov. in life. (E)–(F) dorsolateral views of adult males paratypes SYS a002493–2494."figureDoi="http://doi.org/10.5281/zenodo.4533860"httpUri="https://zenodo.org/record/4533860/files/figure.png"pageId="19"pageNumber="28">Fig. 5</figureCitation>
<collectingDateid="EFE3E991FFAFFFB1665FFBB2FB6DD724"box="[1094,1248,1036,1063]"pageId="19"pageNumber="28"value="2014-04-18">18 April 2014</collectingDate>
<specimenCodeid="DBBF9EC2FFAFFFB161D8FBEAFBC2D76C"box="[961,1103,1108,1135]"collectionCode="CIB"country="China"httpUri="http://grbio.org/cool/vxkr-andy"name="Chengdu Institute of Biology"pageId="19"pageNumber="28">CIB 116077</specimenCode>
) and one adult female (SYS a002500), collected at the same time from the same locality as the holotype
<taxonomicNameid="4C194D3AFFAFFFB16020FB22FD5DD7B5"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[569,720,1181,1206]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="sanmingensis">
in adult female; (2) tympanum distinct, TD/ED 0.49–0.50; (3) vomerine teeth absent; (4) margin of tongue notched; (5) heels overlapping; (6) tibio-tarsal articulation reaching just posterior to eye; (7) TIB/SVL 0.44–0.48, FTL/SVL 0.61–0.65; (8) narrow lateral fringes on fingers present, one subarticular tubercle present at the base of each finger, relative finger lengths I <II <IV <III; (9) rudimentary web and wide lateral fringes on toes present, one subarticular tubercle present at the base of each toe; (10) dorsal body’s skin texture rough, with densely-distributed granules and scattered raised tubercles, “X” shaped skin ridge on center of dorsum and dorsolateral skin ridges present; (11) flank with enlarged raised conical tubercles; (12) supratympanic fold distinct; (13) dorsal surfaces yellowish brown or reddish brown, a dark incomplete triangular marking with light edge between eyes, a dark “X” shaped marking, with light edge, present on center of dorsum; (14) single subgular vocal sac present in males; (15) nuptial spines present on the finger
<taxonomicNameid="4C194D3AFFAFFFB16072F9CFFC08D589"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[619,901,1648,1674]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFAFFFB16628F9CEFB59D589"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1073,1236,1648,1674]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="daiyunensis">
<taxonomicNameid="4C194D3AFFAFFFB166FCF9CEFAB0D589"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1253,1341,1648,1674]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="daoji">
<taxonomicNameid="4C194D3AFFAFFFB16300F92BFE1FD5AD"authorityName="Rao & Yang"authorityYear="1997"box="[281,402,1684,1710]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="genus">
<tableCitationid="C69B0302FFAFFFB16085F92BFD7CD5AC"box="[668,753,1684,1711]"captionStart="TABLE 3"captionStartId="8.[151,239,468,494]"captionTargetPageId="8"captionText="TABLE 3. Diagnostic characters separating all 51 species of the genus Panophrys."httpUri="http://table.plazi.org/id/DF666631FFB4FFAA628EFE6BFB81D2ED"pageId="19"pageNumber="28"tableUuid="DF666631FFB4FFAA628EFE6BFB81D2ED">Table 3</tableCitation>
<taxonomicNameid="4C194D3AFFAFFFB162DEF907FE52D5D1"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[199,479,1720,1746]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFAFFFB160CDF906FCF9D5D1"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[724,884,1720,1746]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="daiyunensis">
by its vomerine teeth absent (vs present), tongue notched (vs not notched), lateral fringes on toes wide (vs narrow), and nuptial spines present on the finger I in breeding males (vs absent).
<taxonomicNameid="4C194D3AFFAFFFB163CEF8BFFD62D419"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[471,751,1792,1818]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFAFFFB161F2F8BEFBCFD419"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1003,1090,1792,1818]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="daoji">
<taxonomicNameid="4C194D3AFFAFFFB163DEF8D3FDCDD485"authorityName="Rao & Yang"authorityYear="1997"box="[455,576,1900,1926]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="genus">
<taxonomicNameid="4C194D3AFFAFFFB16679F8D2FA9CD485"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1120,1297,1901,1926]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFAFFFB1669FF80BFA11D4CD"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1158,1436,1972,1998]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFAFFFB1639FF866FDD8D4F1"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Fei & Ye"baseAuthorityYear="2005"box="[390,597,2008,2034]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="19"pageNumber="28"phylum="Chordata"rank="species"species="huangshanensis">
in female), heels overlapping (vs just meeting or not meeting), wide lateral fringes and rudimentary web on toes present (vs both absent), and round light patches on the shoulder absent (vs present).
<taxonomicNameid="4C194D3AFFA8FFB6663DFF04FAB1D3D7"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1060,1340,187,213]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFA8FFB660B8FEBCFC37D21F"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[673,954,259,285]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFA8FFB666A1FEBCFAC1D21E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Liu & Jiang"baseAuthorityYear="2017"box="[1208,1356,259,285]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="lishuiensis">
<taxonomicNameid="4C194D3AFFA8FFB661CFFEF4FB60D267"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[982,1261,331,357]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFA8FFB662CFFED0FEF8D28A"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Messenger & Dahn"baseAuthorityYear="2019"box="[214,373,367,393]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="ombrophila">
by the tongue notched (vs not notched), heels overlapping (vs not meeting), and wide lateral fringes and rudimentary web on toes present (vs both absent).
<taxonomicNameid="4C194D3AFFA8FFB66175FE2CFB0BD2AF"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[876,1158,403,429]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFA8FFB66791FE2CFE98D2D3"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Wu, Peng, Shi, Lu & Wu"baseAuthorityYear="2020"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="xianjuensis">
<taxonomicNameid="4C194D3AFFA8FFB6663FFE40FAB0D11B"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1062,1341,511,537]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFA8FFB666A3FD9CFA97D13E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy & Che"baseAuthorityYear="2019"box="[1210,1306,547,573]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="angka">
<taxonomicNameid="4C194D3AFFA8FFB66730FD9CFF50D163"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye, Fei & Xie"baseAuthorityYear="2007"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="baolongensis">
<taxonomicNameid="4C194D3AFFA8FFB662F2FDF8FE2BD162"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"box="[235,422,583,609]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="binchuanensis">
<taxonomicNameid="4C194D3AFFA8FFB663ADFDF8FDDBD162"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Jiang, Fei & Ye"baseAuthorityYear="2009"box="[436,598,583,609]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="binlingensis">
<taxonomicNameid="4C194D3AFFA8FFB6607DFDF8FC85D162"baseAuthorityName="Inger & Romer"baseAuthorityYear="1961"box="[612,776,583,609]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="brachykolos">
<taxonomicNameid="4C194D3AFFA8FFB6610CFDF8FC45D162"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Nguyen, Pham, Nguyen, Luong & Ziegler"baseAuthorityYear="2020"box="[789,968,583,609]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="caobangensis">
<taxonomicNameid="4C194D3AFFA8FFB661CFFDF8FBF3D162"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Shen"baseAuthorityYear="1994"box="[982,1150,583,609]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="caudoprocta">
<taxonomicNameid="4C194D3AFFA8FFB66692FDF8FAB9D162"baseAuthorityName="Rao & Yang"baseAuthorityYear="1997"box="[1163,1332,583,609]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="daweimontis">
<taxonomicNameid="4C194D3AFFA8FFB66758FDF8FF74D187"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley"baseAuthorityYear="2018"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="fansipanensis">
<taxonomicNameid="4C194D3AFFA8FFB66311FDD4FE41D186"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley"baseAuthorityYear="2018"box="[264,460,619,645]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="hoanglienensis">
<taxonomicNameid="4C194D3AFFA8FFB663C2FDD4FDD4D186"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Liu, Lyu, Zeng & Wang"baseAuthorityYear="2017"box="[475,601,619,645]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="insularis">
<taxonomicNameid="4C194D3AFFA8FFB66071FDD4FD48D187"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang & Wu"baseAuthorityYear="2020"box="[616,709,619,644]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="jiangi">
<taxonomicNameid="4C194D3AFFA8FFB660CDFDD4FC00D186"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Fei & Ye"baseAuthorityYear="1983"box="[724,909,619,645]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="jingdongensis">
<taxonomicNameid="4C194D3AFFA8FFB66182FDD4FBDED187"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang"baseAuthorityYear="2012"box="[923,1107,619,644]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="jinggangensis">
<taxonomicNameid="4C194D3AFFA8FFB6667BFDD4FA8AD186"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Li, Xu, Liu, Jiang, Wei & Wang"baseAuthorityYear="2018"box="[1122,1287,619,645]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="leishanensis">
<taxonomicNameid="4C194D3AFFA8FFB6670CFDD4FA1BD186"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang & Zhou"baseAuthorityYear="2017"box="[1301,1430,619,645]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="liboensis">
<taxonomicNameid="4C194D3AFFA8FFB6628EFD30FF57D1AA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Yang"baseAuthorityYear="2014"box="[151,218,655,681]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="lini">
<taxonomicNameid="4C194D3AFFA8FFB66342FD30FE50D1AA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang & Zhao"baseAuthorityYear="2020"box="[347,477,655,681]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="mirabilis">
<taxonomicNameid="4C194D3AFFA8FFB663F7FD30FD28D1AA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Lyu & Wang"baseAuthorityYear="2019"box="[494,677,655,681]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="mufumontana">
<taxonomicNameid="4C194D3AFFA8FFB660ACFD30FCD0D1AA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng & Wang"baseAuthorityYear="2019"box="[693,861,655,681]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="nankunensis">
<taxonomicNameid="4C194D3AFFA8FFB66174FD30FC41D1AA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Li & Zhao"baseAuthorityYear="2014"box="[877,972,655,681]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="obesa">
<taxonomicNameid="4C194D3AFFA8FFB66791FD30FEA8D1CE"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Jiang, Ye & Fei"baseAuthorityYear="2008"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="sangzhiensis">
<taxonomicNameid="4C194D3AFFA8FFB6632BFD0CFE7BD1CE"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tian & Sun"baseAuthorityYear="1995"box="[306,502,691,717]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="shuichengensis">
<taxonomicNameid="4C194D3AFFA8FFB6601AFD0CFD5DD1CE"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Deng, Liu, Wu & Liu"baseAuthorityYear="2019"box="[515,720,691,717]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="shunhuangensis">
<taxonomicNameid="4C194D3AFFA8FFB660C4FD0CFCC4D1CF"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Liu & Hu"baseAuthorityYear="1973"box="[733,841,691,716]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="spinata">
<taxonomicNameid="4C194D3AFFA8FFB6614EFD0CFBBFD1CE"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Shen, Mo & Li"baseAuthorityYear="2010"box="[855,1074,691,717]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="tuberogranulatus">
<taxonomicNameid="4C194D3AFFA8FFB66626FD0CFB66D1CF"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Lyu & Wang"baseAuthorityYear="2019"box="[1087,1259,691,716]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="wugongensis">
<taxonomicNameid="4C194D3AFFA8FFB666EEFD0CFF50D1F3"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="wuliangshanensis">
<taxonomicNameid="4C194D3AFFA8FFB662F2FD68FE18D1F2"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"box="[235,405,727,753]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="wushanensis">
<taxonomicNameid="4C194D3AFFA8FFB663BBFD68FDD3D1F3"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Zeng & Wang"baseAuthorityYear="2020"box="[418,606,727,752]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="xiangnanensis">
<taxonomicNameid="4C194D3AFFA8FFB66082FD68FCD0D1F3"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Zeng & Wang"baseAuthorityYear="2020"box="[667,861,727,752]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="yangmingensis">
<taxonomicNameid="4C194D3AFFA8FFB662DEFD44FE6DD017"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[199,480,763,789]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<taxonomicNameid="4C194D3AFFA8FFB66724FCA0FA18D03A"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Liu"baseAuthorityYear="2014"box="[1341,1429,799,825]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="cheni">
<taxonomicNameid="4C194D3AFFA8FFB6601AFCFCFD4AD05E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Wang"baseAuthorityYear="2019"box="[515,711,835,861]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="dongguanensis">
<taxonomicNameid="4C194D3AFFA8FFB66791FCFCFE95D082"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng, Lyu & Wang"baseAuthorityYear="2019"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="jiulianensis">
<taxonomicNameid="4C194D3AFFA8FFB660BFFCD8FCC2D082"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang, Liu & Wang"baseAuthorityYear="2019"box="[678,847,871,897]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="nanlingensis">
<taxonomicNameid="4C194D3AFFA8FFB6628EFC34FF7CD0A7"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Li & Jin"baseAuthorityYear="2014"box="[151,241,907,932]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="acuta">
<taxonomicNameid="4C194D3AFFA8FFB660BDFC34FCEAD0A6"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Wang"baseAuthorityYear="2019"box="[676,871,907,933]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="dongguanensis">
<taxonomicNameid="4C194D3AFFA8FFB6616CFC34FB9FD0A6"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng, Lyu & Wang"baseAuthorityYear="2019"box="[885,1042,907,933]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="jiulianensis">
<taxonomicNameid="4C194D3AFFA8FFB66639FC34FB47D0A6"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang, Liu & Wang"baseAuthorityYear="2019"box="[1056,1226,907,933]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="nanlingensis">
<taxonomicNameid="4C194D3AFFA8FFB6628EFC10FEB1D0CA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Liu & Wang"baseAuthorityYear="2020"box="[151,316,943,969]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="shimentaina">
<taxonomicNameid="4C194D3AFFA8FFB660C1FC10FCBFD0CB"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Li & Jin"baseAuthorityYear="2014"box="[728,818,943,968]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="acuta">
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<taxonomicNameid="4C194D3AFFA8FFB6602AFC6CFD7AD0EE"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Wang"baseAuthorityYear="2019"box="[563,759,979,1005]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="dongguanensis">
<taxonomicNameid="4C194D3AFFA8FFB66722FC6CFA18D0EF"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Li & Jin"baseAuthorityYear="2014"box="[1339,1429,979,1004]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="acuta">
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<taxonomicNameid="4C194D3AFFA8FFB66007FC48FD49D712"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Liu & Wang"baseAuthorityYear="2020"box="[542,708,1015,1041]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="shimentaina">
<taxonomicNameid="4C194D3AFFA8FFB66178FC48FBA8D712"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Wang"baseAuthorityYear="2019"box="[865,1061,1015,1041]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="dongguanensis">
<taxonomicNameid="4C194D3AFFA8FFB66644FC48FB77D712"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng, Lyu & Wang"baseAuthorityYear="2019"box="[1117,1274,1015,1041]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="20"pageNumber="29"phylum="Chordata"rank="species"species="jiulianensis">
; head length shorter than head width, HDW/HDL 1.02; snout rounded in dorsal view, projecting, sloping posteriorly to mouth in profile, protruding well beyond margin of lower jaw; dorsal surface of head flat; eye large, ED/HDL 0.42; nostril obliquely ovoid; pupil vertical; canthus rostralis well developed, curved above nostril; loreal region sloping; internasal distance larger than interorbital distance; tympanum distinct; choanae large ovoid, situated at base of maxilla; vomerine teeth absent; margin of tongue notched.
</paragraph>
<paragraphid="8BA636B9FFA8FFB662DEFAA8FF72D69E"blockId="20.[151,1437,151,2013]"pageId="20"pageNumber="29">Lower arm length 0.25 of SVL and hand length 0.24 of SVL; relative finger lengths I <II <IV <III; tip of finger rounded, slightly dilated; one subarticular tubercle present at base of each finger; fingers without webs, narrow lateral fringes present on fingers; outer and inner metacarpal tubercles distinct, inner metacarpal tubercle observably enlarged.</paragraph>
<paragraphid="8BA636B9FFA8FFB662DEFA18FD32D552"blockId="20.[151,1437,151,2013]"pageId="20"pageNumber="29">Shank length 0.48 of SVL and foot 0.64 of SVL; tibio-tarsal articulation reaches posterior margin of eye when hindlimb stretched alongside body; heels overlapping when hindlimbs held at right angles to body; relative toe lengths I <II <V <III <IV; tips of toes rounded, slightly dilated; one subarticular tubercle present at base of each toe; rudimentary web and wide lateral fringes present between toes; tarsal folds absent; inner metatarsal tubercle long, ovoid, and outer metatarsal tubercle absent.</paragraph>
<paragraphid="8BA636B9FFA8FFB662DEF9E4FA98D406"blockId="20.[151,1437,151,2013]"pageId="20"pageNumber="29">Dorsal body’s skin texture rough with densely-distributed granules and scattered raised tubercles; “X” shaped skin ridge on center of dorsum; dorsolateral skin ridges present; a small horn-like tubercle present at edge of upper eyelid; supratympanic fold distinct, curving from posterior corner of eye, posteroventrally to above insertion of arm; flank with raised tubercles; raised tubercles on dorsal arms and hindlimbs. Ventral surface smooth with tiny granules; pectoral gland large, closer to axilla; single large femoral gland on posterior surface of thigh.</paragraph>
Dorsal surface brown; a dark incomplete triangular marking with light edge between eyes; a dark “X” shaped marking with light edge on center of dorsum; dark crossbars on dorsal upper arms and hindlimbs; dark stripes below eyes and at lateral tip of snout; iris yellowish brown. Ventral surface pale with densely-distributed white spots; three dark longitudinal stripes on the throat; a pair of longitudinal black stripes with white edge on the lateroventral belly; palms and soles purplish brown, tips of digits greyish white, metacarpal and metatarsal tubercles orange red; pectoral glands and femoral glands white.
<taxonomicNameid="4C194D3AFFA9FFB76747F8F4FED2D485"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="21"pageNumber="30"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
<tableCitationid="C69B0302FFAAFFB46102FF29FCFCD3B3"box="[795,881,150,176]"captionStart="TABLE 6"captionStartId="22.[151,239,656,680]"captionTargetPageId="22"captionText="TABLE 6. Measurements (in mm) of the type series of Panophrys sanmingensis sp. nov., * for the holotype."httpUri="http://table.plazi.org/id/DF666631FFAAFFB4628EFD2FFAAFD1AB"pageId="22"pageNumber="31"tableUuid="DF666631FFAAFFB4628EFD2FFAAFD1AB">Table 6</tableCitation>
. All specimens were similar in morphology. SYS a002493 (
<figureCitationid="13222A3CFFAAFFB46311FF05FED0D3D6"box="[264,349,186,213]"captionStart="FIGURE 5"captionStartId="21.[151,250,1866,1890]"captionTargetBox="[151,1436,181,1842]"captionTargetId="figure-20@21.[151,1436,181,1842]"captionTargetPageId="21"captionText="FIGURE 5. (A)–(D) dorsolateral view, ventral view, right hand, and right foot of adult male holotype SYS a002498 of Panophrys sanmingensis sp. nov. in life. (E)–(F) dorsolateral views of adult males paratypes SYS a002493–2494."figureDoi="http://doi.org/10.5281/zenodo.4533860"httpUri="https://zenodo.org/record/4533860/files/figure.png"pageId="22"pageNumber="31">Fig. 5E</figureCitation>
) has reddish brown dorsal surface, with red spots on the dorsum and flanks. SYS a002494 (
<figureCitationid="13222A3CFFAAFFB46776FF05FF38D3FA"captionStart="FIGURE 5"captionStartId="21.[151,250,1866,1890]"captionTargetBox="[151,1436,181,1842]"captionTargetId="figure-20@21.[151,1436,181,1842]"captionTargetPageId="21"captionText="FIGURE 5. (A)–(D) dorsolateral view, ventral view, right hand, and right foot of adult male holotype SYS a002498 of Panophrys sanmingensis sp. nov. in life. (E)–(F) dorsolateral views of adult males paratypes SYS a002493–2494."figureDoi="http://doi.org/10.5281/zenodo.4533860"httpUri="https://zenodo.org/record/4533860/files/figure.png"pageId="22"pageNumber="31">Fig. 5F</figureCitation>
) and SYS a002495 have black spots on flanks and the posterior of dorsum.
<taxonomicNameid="4C194D3AFFAAFFB46099FEBDFC11D21F"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[640,924,258,284]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="22"pageNumber="31"phylum="Chordata"rank="species"species="sanmingensis"status="sp. nov.">
provinces. This species inhabits streams surrounded by moist subtropical secondary evergreen broadleaved forests. Males call actively from mid-April to early June in Mt Longqi, Mt Emeifeng, Mt Magu, and Mt Junfeng; two individuals were found from Gutian Town in mid-July, and they were calling inactively; only one individual was found from Mt Jinpen in late September, and it was not calling.
