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304 lines
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<document id="940EAEDA221006DBA27CDA52E1B4C494" ID-DOI="10.11646/phytotaxa.436.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13874448" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727817160391" checkinUser="felipe" docAuthor="Bruyns, Peter V., Klak, Cornelia & Hanáček, Pavel" docDate="2020" docId="03A6CB7E02388E28DC9DFE6EFA126C1F" docLanguage="en" docName="phytotaxa.436.3.1.pdf" docOrigin="Phytotaxa 436 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.436.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Euphorbia schinzii , Pax 1898" docType="treatment" docVersion="2" lastPageNumber="213" masterDocId="FF9FB30602338E24DC15FFD9FFB86F27" masterDocTitle="A review of the Euphorbia schinzii-complex (Euphorbiaceae) in southern Africa" masterLastPageNumber="221" masterPageNumber="201" pageNumber="212" updateTime="1727819793323" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="4A5168682C3C8C4C892A259178AED1E7">
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<mods:title id="36E4A1FDCA658C6D80490F7890B583F6">A review of the Euphorbia schinzii-complex (Euphorbiaceae) in southern Africa</mods:title>
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<mods:name id="E50ECB2F4759CAE227C4D110AB07D2B0" type="personal">
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<mods:roleTerm id="C64D4A5165D299B4FBDDF2CCE39483A5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="499F2F13EBFCA9ED354161091A5E9DF9">Bruyns, Peter V.</mods:namePart>
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<mods:affiliation id="CC868B282B86EF34934D02F886E6C183">Bolus Herbarium, University of Cape Town, 7701 Rondebosch, South Africa</mods:affiliation>
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<mods:nameIdentifier id="89398759ABEA2A139F5D15910075C84D" type="email">peter.bruyns@uct.ac.za</mods:nameIdentifier>
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<mods:name id="B5EF02DE52842EE8552CFF544AE76691" type="personal">
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</mods:role>
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<mods:namePart id="534D5B522569F52D06A969DBA345FFEE">Klak, Cornelia</mods:namePart>
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<mods:affiliation id="2E3241963A448A56EB7E67D2DB1DBC92">Bolus Herbarium, University of Cape Town, 7701 Rondebosch, South Africa</mods:affiliation>
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</mods:name>
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<mods:name id="F9AD4B569ECE0284C769E760DF9B7A73" type="personal">
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<mods:roleTerm id="011C65A67BC649E642A7BC305CA7C33C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C712CA4AC9D2C756F5FF67DFF2318B8F">Hanáček, Pavel</mods:namePart>
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<mods:affiliation id="6A840FA3231FB1C83B060F1098223A48">Department of Plant Biology, Mendel University in Brno, Zemědělská 1, 613 00 Brno, Czechia</mods:affiliation>
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<mods:typeOfResource id="21F5C06FBD4323730108C4E0ED248B2F">text</mods:typeOfResource>
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<mods:title id="A6CF13A92197836B8B58825BB8410FA6">Phytotaxa</mods:title>
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<mods:part id="EAF5DCBE9435EB53D9FFFD1F03A0E48D">
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<mods:date id="B2CE2FCF2EDECA0375E38E6447384CEA">2020</mods:date>
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<mods:detail id="00A518F3CA22A8C061A04E1F03FA76B8" type="pubDate">
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<mods:number id="6C077B7FD2D49AC14F399EE417000DE0">2020-03-19</mods:number>
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<mods:detail id="998EE9C9D57332342D0731FA46DB319A" type="volume">
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<mods:number id="47C7705D7ECBD5E19EC1F039FF741223">436</mods:number>
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<mods:detail id="6E4AC14C121AAD235ED7152AA108CE1D" type="issue">
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<mods:number id="BC5C578A3F75953C189C68CA1506F841">3</mods:number>
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<mods:start id="43CB9D0BD1BB6AC6CEF23996D45F926B">201</mods:start>
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<mods:end id="7814E33DA48E6BFCE04EF98E7E2C4925">221</mods:end>
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<mods:url id="606D1C7829C899A207633756409A4CAB">http://dx.doi.org/10.11646/phytotaxa.436.3.1</mods:url>
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</mods:location>
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<mods:classification id="CB5EF633C7CD99CAE6B297013C4A6893">journal article</mods:classification>
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<mods:identifier id="F7E3DDD488BE74C9D29FECC53D50B9BD" type="DOI">10.11646/phytotaxa.436.3.1</mods:identifier>
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<mods:identifier id="791393568BC96A49C06AA15548C376BC" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="D14619B03389156A3584116FAD26F030" type="Zenodo-Dep">13874448</mods:identifier>
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</mods:mods>
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<treatment id="03A6CB7E02388E28DC9DFE6EFA126C1F" ID-DOI="http://doi.org/10.5281/zenodo.13874478" ID-Zenodo-Dep="13874478" LSID="urn:lsid:plazi:treatment:03A6CB7E02388E28DC9DFE6EFA126C1F" httpUri="http://treatment.plazi.org/id/03A6CB7E02388E28DC9DFE6EFA126C1F" lastPageId="12" lastPageNumber="213" pageId="11" pageNumber="212">
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<subSubSection id="C31529E302388E2FDC9DFE6EFDFB6EF5" box="[136,579,439,466]" pageId="11" pageNumber="212" type="nomenclature">
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<paragraph id="8BB07A6802388E2FDC9DFE6EFDFB6EF5" blockId="11.[136,579,439,466]" box="[136,579,439,466]" pageId="11" pageNumber="212">
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<heading id="D0F8CD0402388E2FDC9DFE6EFDFB6EF5" box="[136,579,439,466]" fontSize="11" level="2" pageId="11" pageNumber="212" reason="4">
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<emphasis id="B97BA67A02388E2FDC9DFE6EFF256EF6" bold="true" box="[136,157,439,465]" pageId="11" pageNumber="212">8.</emphasis>
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<taxonomicName id="4C0F01EB02388E2FDCB1FE6EFD876EF5" ID-CoL="3CQNP" authority="Pax (1898: 739)" authorityName=", Pax" authorityPageNumber="739" authorityYear="1898" box="[164,575,439,466]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="schinzii">
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<emphasis id="B97BA67A02388E2FDCB1FE6EFE396EF5" bold="true" box="[164,385,439,466]" italics="true" pageId="11" pageNumber="212">Euphorbia schinzii</emphasis>
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<bibRefCitation id="EF9E079902388E2FDD92FE61FD876EF5" author="Pax, F." box="[391,575,440,466]" pageId="11" pageNumber="212" pagination="729 - 743" refId="ref16919" refString="Pax, F. (1898) Euphorbiaceae. In: Schinz, H. (Ed.) Beitrage zur Kenntnis der Afrikanischen Flora. Bulletin de l'Herbier Boissier 6: 729 - 743." type="journal article" year="1898">Pax (1898: 739)</bibRefCitation>
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</taxonomicName>
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.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31529E302388E2FDC9DFE24FD4E6D11" pageId="11" pageNumber="212" type="materials_examined">
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<paragraph id="8BB07A6802388E2FDC9DFE24FD4E6D11" blockId="11.[136,1452,508,566]" pageId="11" pageNumber="212">
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Type:—
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<collectingCountry id="F3183AF802388E2FDCC0FE24FE306D35" box="[213,392,509,530]" name="South Africa" pageId="11" pageNumber="212">SOUTH AFRICA</collectingCountry>
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. Transvaal (
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<collectingRegion id="49CBB48A02388E2FDDEBFE25FD3B6D35" box="[510,643,508,530]" country="South Africa" name="Mpumalanga" pageId="11" pageNumber="212">Mpumalanga</collectingRegion>
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): Berea Ridge, Barberton, 3100’,
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<date id="FFB15CA802388E2FDFDBFE24FBC66D35" box="[974,1150,508,530]" pageId="11" pageNumber="212" value="1891-02-13">13 February 1891</date>
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,
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<emphasis id="B97BA67A02388E2FD89FFE25FABC6D35" box="[1162,1284,508,530]" italics="true" pageId="11" pageNumber="212">Galpin 1297</emphasis>
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(
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<typeStatus id="54B4C4CA02388E2FD907FE25FAD46D35" box="[1298,1388,508,530]" pageId="11" pageNumber="212" type="lectotype">lectotype</typeStatus>
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<collectionCode id="ED1EE2AD02388E2FD966FE24FA106D35" box="[1395,1448,509,530]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15865" lsid="urn:lsid:biocol.org:col:15865" name="University of Cape Town" pageId="11" pageNumber="212" type="Herbarium">BOL</collectionCode>
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! designated by
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<bibRefCitation id="EF9E079902388E2FDD5DFDF8FDBE6D11" author="Bruyns, P. V." box="[328,518,545,566]" pageId="11" pageNumber="212" pagination="217 - 245" refId="ref16021" refString="Bruyns, P. V. (2012) Nomenclature and typification of southern African species of Euphorbia. Bothalia 42: 217 - 245. https: // doi. org / 10.4102 / abc. v 42 i 2.23" type="journal article" year="2012">Bruyns (2012: 228)</bibRefCitation>
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,
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<typeStatus id="54B4C4CA02388E2FDE05FDF9FD3C6D11" box="[528,644,544,566]" pageId="11" pageNumber="212" type="isolectotype">isolectotype</typeStatus>
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<collectionCode id="ED1EE2AD02388E2FDE9FFDF8FD196D11" box="[650,673,545,566]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="11" pageNumber="212" type="Herbarium">K</collectionCode>
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!).
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<figureCitation id="133466ED02388E2FDEA2FDF9FD496D11" box="[695,753,544,566]" captionStart="FIGURE 6" captionStartId="11.[136,229,2023,2045]" captionTargetBox="[151,1436,1017,1998]" captionTargetId="figure-362@11.[151,1436,1017,1998]" captionTargetPageId="11" captionText="FIGURE 6. Distribution of Euphorbia schinzii (inverted triangle = subsp. schinzii, circle = subsp. bechuanica, upright triangle = subsp. schinzioides)." figureDoi="http://doi.org/10.5281/zenodo.13874464" httpUri="https://zenodo.org/record/13874464/files/figure.png" pageId="11" pageNumber="212">Fig. 6</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31529E302388E28DC9DFD87FCD66D4F" lastPageId="12" lastPageNumber="213" pageId="11" pageNumber="212" type="discussion">
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<paragraph id="8BB07A6802388E2FDC9DFD87FD5A6CC6" blockId="11.[136,1452,606,993]" pageId="11" pageNumber="212">
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<emphasis id="B97BA67A02388E2FDC9DFD87FEA86D5F" bold="true" box="[136,272,606,632]" pageId="11" pageNumber="212">Discussion:</emphasis>
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—In the protologue of
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<taxonomicName id="4C0F01EB02388E2FDE00FD86FCAE6D5F" authority=", Pax (1898)" authorityName=", Pax" authorityYear="1898" box="[533,790,606,633]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="schinzii">
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<emphasis id="B97BA67A02388E2FDE00FD86FD336D5F" box="[533,651,606,632]" italics="true" pageId="11" pageNumber="212">E. schinzii</emphasis>
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,
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<bibRefCitation id="EF9E079902388E2FDE8FFD86FCAE6D5F" author="Pax, F." box="[666,790,606,633]" pageId="11" pageNumber="212" pagination="729 - 743" refId="ref16919" refString="Pax, F. (1898) Euphorbiaceae. In: Schinz, H. (Ed.) Beitrage zur Kenntnis der Afrikanischen Flora. Bulletin de l'Herbier Boissier 6: 729 - 743." type="journal article" year="1898">Pax (1898)</bibRefCitation>
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</taxonomicName>
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cited
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<specimenCount id="9D09B1E102388E2FDF48FD86FBBF6D5F" box="[861,1031,606,633]" count="2" pageId="11" pageNumber="212" type="generic">two specimens</specimenCount>
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,
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<emphasis id="B97BA67A02388E2FD807FD87FB046D5F" box="[1042,1212,606,632]" italics="true" pageId="11" pageNumber="212">Rehmann 4347</emphasis>
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from Pretoria,
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<collectingCountry id="F3183AF802388E2FD97FFD87FF686DBB" name="South Africa" pageId="11" pageNumber="212">South Africa</collectingCountry>
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(missing and not seen by N.E.
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<bibRefCitation id="EF9E079902388E2FDE2EFD5AFCAD6DBB" author="Brown, N. E." box="[571,789,642,669]" pageId="11" pageNumber="212" pagination="222 - 375" refId="ref15979" refString="Brown, N. E. (1915) Euphorbia. In: Thiselton-Dyer, W. T., Flora Capensis 5 (2). L. Reeve & Co., London, pp. 222 - 375." type="book chapter" year="1915">Brown (1915: 364)</bibRefCitation>
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either) and
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<emphasis id="B97BA67A02388E2FDFB4FD5BFB976DBB" box="[929,1071,642,668]" italics="true" pageId="11" pageNumber="212">Galpin 1297</emphasis>
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(BOL!, K!) from Berea Ridge at Barberton,
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<collectingCountry id="F3183AF802388E2FDD11FD7FFE2A6DE7" box="[260,402,678,704]" name="South Africa" pageId="11" pageNumber="212">South Africa</collectingCountry>
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(±
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<quantity id="4CF7D78D02388E2FDDA0FD7FFDB06DE7" box="[437,520,678,704]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="212" unit="km" value="300.0">300 km</quantity>
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east of Pretoria). Pax gave the branches as 3-angled, but that is certainly an error and the fourth angle must have been entirely hidden in the pressed specimens he saw. He also mentioned ‘
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<emphasis id="B97BA67A02388E2FD910FD13FF5D6C2F" italics="true" pageId="11" pageNumber="212">angulis lobatis podariis</emphasis>
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’, i.e. branches angled with the tubercles projecting somewhat (‘lobed’). Since
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<emphasis id="B97BA67A02388E2FD86FFD37FA9E6C2F" box="[1146,1318,750,776]" italics="true" pageId="11" pageNumber="212">Rehmann 4347</emphasis>
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is missing,
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<emphasis id="B97BA67A02388E2FDC9DFCCBFEAC6C0B" box="[136,276,786,812]" italics="true" pageId="11" pageNumber="212">Galpin 1297</emphasis>
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was selected as the
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<typeStatus id="54B4C4CA02388E2FDDE5FCCBFDEF6C0B" box="[496,599,786,812]" pageId="11" pageNumber="212" type="lectotype">lectotype</typeStatus>
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for
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<taxonomicName id="4C0F01EB02388E2FDE96FCCAFD4E6C0B" authorityName=", Pax" authorityYear="1898" box="[643,758,786,812]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="schinzii">
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<emphasis id="B97BA67A02388E2FDE96FCCAFD4E6C0B" box="[643,758,786,812]" italics="true" pageId="11" pageNumber="212">E. schinzii</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="EF9E079902388E2FDF0BFCCAFC7A6C0B" author="Bruyns, P. V." box="[798,962,786,813]" pageId="11" pageNumber="212" pagination="217 - 245" refId="ref16021" refString="Bruyns, P. V. (2012) Nomenclature and typification of southern African species of Euphorbia. Bothalia 42: 217 - 245. https: // doi. org / 10.4102 / abc. v 42 i 2.23" type="journal article" year="2012">Bruyns (2012)</bibRefCitation>
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. In
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<emphasis id="B97BA67A02388E2FDFFCFCCBFBCC6C0B" box="[1001,1140,786,812]" italics="true" pageId="11" pageNumber="212">Galpin 1297</emphasis>
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the branches were
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<quantity id="4CF7D78D02388E2FD95DFCCBFF0C6C76" metricMagnitude="-1" metricUnit="m" metricValue="1.625" metricValueMax="1.75" metricValueMin="1.5" pageId="11" pageNumber="212" unit="mm" value="162.5" valueMax="175.0" valueMin="150.0">150−175 mm</quantity>
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tall (‘6−8 inches’), the tubercles are fused into angles, the spines are subtended by stipular prickles ±
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<quantity id="4CF7D78D02388E2FD924FCEFFACB6C76" box="[1329,1395,822,849]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="212" unit="mm" value="2.0">2 mm</quantity>
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long and the specimen was collected in flower in February. The relatively tall branches, continuous angles with decurrent but not continuous spine-shields, fairly prominent stipular prickles and sessile cyathia with ± sessile ovaries make it clear that
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<emphasis id="B97BA67A02388E2FDCEDFC7BFE3E6C9B" box="[248,390,930,956]" italics="true" pageId="11" pageNumber="212">Galpin 1297</emphasis>
|
|||
|
|
is the same species as the type of
|
|||
|
|
<taxonomicName id="4C0F01EB02388E2FDF18FC7AFC206C9B" authorityName=", Dyer" authorityYear="1937" box="[781,920,930,956]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="complexa">
|
|||
|
|
<emphasis id="B97BA67A02388E2FDF18FC7AFC206C9B" box="[781,920,930,956]" italics="true" pageId="11" pageNumber="212">E. complexa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Other recent collections from Barberton (e.g.,
|
|||
|
|
<emphasis id="B97BA67A02388E2FDC9DFC1FFEB26CC7" box="[136,266,966,992]" italics="true" pageId="11" pageNumber="212">Kluge 2281</emphasis>
|
|||
|
|
(NBG)) are also identical to
|
|||
|
|
<taxonomicName id="4C0F01EB02388E2FDE45FC1EFD636CC7" authorityName=", Dyer" authorityYear="1937" box="[592,731,966,992]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="complexa">
|
|||
|
|
<emphasis id="B97BA67A02388E2FDE45FC1EFD636CC7" box="[592,731,966,992]" italics="true" pageId="11" pageNumber="212">E. complexa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF702AE002388E2FDC9DF83EFEB36707" ID-DOI="http://doi.org/10.5281/zenodo.13874464" ID-Zenodo-Dep="13874464" httpUri="https://zenodo.org/record/13874464/files/figure.png" pageId="11" pageNumber="212" startId="11.[136,229,2023,2045]" targetBox="[151,1436,1017,1998]" targetPageId="11" targetType="figure">
|
|||
|
|
<paragraph id="8BB07A6802388E2FDC9DF83EFEB36707" blockId="11.[136,1452,2023,2080]" pageId="11" pageNumber="212">
|
|||
|
|
<emphasis id="B97BA67A02388E2FDC9DF83EFF4668DB" bold="true" box="[136,254,2023,2045]" pageId="11" pageNumber="212">FIGURE 6.</emphasis>
|
|||
|
|
Distribution of
|
|||
|
|
<taxonomicName id="4C0F01EB02388E2FDD8FF83EFDE868DB" authorityName=", Pax" authorityYear="1898" box="[410,592,2023,2044]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="species" species="schinzii">
|
|||
|
|
<emphasis id="B97BA67A02388E2FDD8FF83EFDE868DB" box="[410,592,2023,2044]" italics="true" pageId="11" pageNumber="212">Euphorbia schinzii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(inverted triangle
|
|||
|
|
<taxonomicName id="4C0F01EB02388E2FDF11F831FC1F68DB" box="[772,935,2023,2045]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="subSpecies" species="schinzii" subSpecies="schinzii">
|
|||
|
|
= subsp.
|
|||
|
|
<emphasis id="B97BA67A02388E2FDF4EF83EFC1F68DB" box="[859,935,2023,2044]" italics="true" pageId="11" pageNumber="212">schinzii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B97BA67A02388E2FDFB2F83EFC1368DB" box="[935,939,2023,2044]" italics="true" pageId="11" pageNumber="212">,</emphasis>
|
|||
|
|
circle
|
|||
|
|
<taxonomicName id="4C0F01EB02388E2FDFF9F831FB0B68DB" authorityName=", Bruyns" authorityYear="2020" baseAuthorityName=", Bruyns" baseAuthorityYear="2020" box="[1004,1203,2023,2045]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="subSpecies" species="schinzii" subSpecies="bechuanica">
|
|||
|
|
= subsp.
|
|||
|
|
<emphasis id="B97BA67A02388E2FD851F83EFB0B68DB" box="[1092,1203,2023,2044]" italics="true" pageId="11" pageNumber="212">bechuanica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, upright triangle
|
|||
|
|
<taxonomicName id="4C0F01EB02388E2FD94EF831FF456707" authorityName=", Bruyns" authorityYear="2020" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="212" phylum="Tracheophyta" rank="subSpecies" species="schinzii" subSpecies="schinzioides">
|
|||
|
|
= subsp.
|
|||
|
|
<emphasis id="B97BA67A02388E2FDC9DF7D2FF456707" box="[136,253,2059,2080]" italics="true" pageId="11" pageNumber="212">schinzioides</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BB07A68023F8E28DCA8FF47FB496EB7" blockId="12.[136,1452,158,545]" pageId="12" pageNumber="213">
|
|||
|
|
Traditionally, small (mostly <
|
|||
|
|
<quantity id="4CF7D78D023F8E28DE07FF47FDD76F9E" box="[530,623,158,185]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="213" unit="mm" value="150.0">150 mm</quantity>
|
|||
|
|
tall), clump-forming members of this complex from around Pretoria (also westwards and northwards) have been referred to as
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28DED9FF1BFC276FFB" authorityName=", Pax" authorityYear="1898" box="[716,927,194,220]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="schinzii">
|
|||
|
|
<emphasis id="B97BA67A023F8E28DED9FF1BFC276FFB" box="[716,927,194,220]" italics="true" pageId="12" pageNumber="213">Euphorbia schinzii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. These have been taken as typical of
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28D92DFF1AFEDF6E27" authority="(White et al. 1941)" baseAuthorityName="White" baseAuthorityYear="1941" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="schinzii">
|
|||
|
|
<emphasis id="B97BA67A023F8E28D92DFF1AFA146FFB" box="[1336,1452,194,220]" italics="true" pageId="12" pageNumber="213">E. schinzii</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EF9E0799023F8E28DC9AFF3FFEE66E27" author="White, A. & Dyer, R. A. & Sloane, B. L." box="[143,350,230,257]" pageId="12" pageNumber="213" refId="ref17146" refString="White, A., Dyer, R. A. & Sloane, B. L. (1941) The Succulent Euphorbieae (Southern Africa). Abbey Garden Press, Pasadena, California, 990 pp." type="book" year="1941">
|
|||
|
|
White
|
|||
|
|
<emphasis id="B97BA67A023F8E28DCCAFF3EFEAB6E27" box="[223,275,230,256]" italics="true" pageId="12" pageNumber="213">et al</emphasis>
|
|||
|
|
. 1941
|
|||
|
|
</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, to the extent that
|
|||
|
|
<bibRefCitation id="EF9E0799023F8E28DE54FF3FFCE46E26" author="White, A. & Dyer, R. A. & Sloane, B. L." box="[577,860,230,257]" pageId="12" pageNumber="213" refId="ref17146" refString="White, A., Dyer, R. A. & Sloane, B. L. (1941) The Succulent Euphorbieae (Southern Africa). Abbey Garden Press, Pasadena, California, 990 pp." type="book" year="1941">
|
|||
|
|
White
|
|||
|
|
<emphasis id="B97BA67A023F8E28DE9AFF3EFD7B6E27" box="[655,707,230,256]" italics="true" pageId="12" pageNumber="213">et al</emphasis>
|
|||
|
|
. (1941: 745)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF9E0799023F8E28DF7FFF3FFB956E27" author="Fourie, S. P." box="[874,1069,230,256]" pageId="12" pageNumber="213" pagination="83 - 93" refId="ref16618" refString="Fourie, S. P. (1988) An introduction to the succulent Euphorbias of the Transvaal, Part 3: Dwarf Shrubs I. The Euphorbia Journal 5: 83 - 93." type="journal article" year="1988">Fourie (1988: 91</bibRefCitation>
|
|||
|
|
, 92) and
|
|||
|
|
<bibRefCitation id="EF9E0799023F8E28D888FF3EFA136E27" author="Hargreaves, B. J." box="[1181,1451,230,257]" pageId="12" pageNumber="213" pagination="147 - 153" refId="ref16688" refString="Hargreaves, B. J. (1994) The Euphorbia schinzii complex. The Euphorbia Journal 9: 147 - 153." type="journal article" year="1994">Hargreaves (1994: 147)</bibRefCitation>
|
|||
|
|
all gave the ‘
|
|||
|
|
<typeStatus id="54B4C4CA023F8E28DD0CFED3FEEE6E03" box="[281,342,266,292]" pageId="12" pageNumber="213">Type</typeStatus>
|
|||
|
|
locality’ of
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28DDF7FED2FDE06E03" box="[482,600,266,292]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="schinzii">
|
|||
|
|
<emphasis id="B97BA67A023F8E28DDF7FED2FDE06E03" box="[482,600,266,292]" italics="true" pageId="12" pageNumber="213">E. schinzii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as ‘Pretoria Distr.; hills near Pretoria’. These plants are variably, but often strongly rhizomatous (
|
|||
|
|
<bibRefCitation id="EF9E0799023F8E28DD92FEF7FDEC6E6F" author="White, A. & Dyer, R. A. & Sloane, B. L." box="[391,596,302,329]" pageId="12" pageNumber="213" refId="ref17146" refString="White, A., Dyer, R. A. & Sloane, B. L. (1941) The Succulent Euphorbieae (Southern Africa). Abbey Garden Press, Pasadena, California, 990 pp." type="book" year="1941">
|
|||
|
|
White
|
|||
|
|
<emphasis id="B97BA67A023F8E28DDC3FEF6FDB16E6F" box="[470,521,302,328]" italics="true" pageId="12" pageNumber="213">et al</emphasis>
|
|||
|
|
. 1941
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: Fig. 835, 836, 838,
|
|||
|
|
<bibRefCitation id="EF9E0799023F8E28DF56FEF6FBF96E6E" author="Hargreaves, B. J." box="[835,1089,302,329]" pageId="12" pageNumber="213" pagination="147 - 153" refId="ref16688" refString="Hargreaves, B. J. (1994) The Euphorbia schinzii complex. The Euphorbia Journal 9: 147 - 153." type="journal article" year="1994">Hargreaves 1994: 153</bibRefCitation>
|
|||
|
|
). The tubercles are fused into ± continuous though somewhat sinuate angles along the branches (sinuate when viewed from the side), the spine-shields are decurrent below the spines for
|
|||
|
|
<quantity id="4CF7D78D023F8E28DE1EFEAFFDD46EB6" box="[523,620,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="12" pageNumber="213" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4−6 mm</quantity>
|
|||
|
|
, stipular prickles are minute and the ovaries are ± sessile.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BB07A68023F8E28DCA8FE43FA316D06" blockId="12.[136,1452,158,545]" pageId="12" pageNumber="213">
|
|||
|
|
Since the name
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28DD7AFE42FE5C6E93" box="[367,484,410,436]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="schinzii">
|
|||
|
|
<emphasis id="B97BA67A023F8E28DD7AFE42FE5C6E93" box="[367,484,410,436]" italics="true" pageId="12" pageNumber="213">E. schinzii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
now applies to plants from Barberton which are identical to
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28D89CFE42FAAB6E93" authorityName=", Dyer" authorityYear="1937" box="[1161,1299,410,436]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="complexa">
|
|||
|
|
<emphasis id="B97BA67A023F8E28D89CFE42FAAB6E93" box="[1161,1299,410,436]" italics="true" pageId="12" pageNumber="213">E. complexa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28D934FE42FA136E93" authorityName=", Dyer" authorityYear="1937" box="[1313,1451,410,436]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="complexa">
|
|||
|
|
<emphasis id="B97BA67A023F8E28D934FE42FA136E93" box="[1313,1451,410,436]" italics="true" pageId="12" pageNumber="213">E. complexa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is a synonym of
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28DD53FE66FE046EFF" box="[326,444,446,472]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="schinzii">
|
|||
|
|
<emphasis id="B97BA67A023F8E28DD53FE66FE046EFF" box="[326,444,446,472]" italics="true" pageId="12" pageNumber="213">E. schinzii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The plants from Pretoria (also westwards and northwards) are not identical to these and require a name. However, they are not always easily distinguishable from
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28DFD8FE3BFB256EDB" authorityName=", Bruyns" authorityYear="2020" baseAuthorityName=", Bruyns" baseAuthorityYear="2020" box="[973,1181,482,508]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="subSpecies" species="schinzii" subSpecies="bechuanica">
|
|||
|
|
subsp.
|
|||
|
|
<emphasis id="B97BA67A023F8E28D809FE3BFB256EDB" box="[1052,1181,482,508]" italics="true" pageId="12" pageNumber="213">bechuanica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see below). Since they are separated by only rather weak and variable features, all three are now included as subspecies under
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28D918FDDEFA3A6D07" box="[1293,1410,518,544]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="species" species="schinzii">
|
|||
|
|
<emphasis id="B97BA67A023F8E28D918FDDEFA3A6D07" box="[1293,1410,518,544]" italics="true" pageId="12" pageNumber="213">E. schinzii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BB07A68023F8E28DC9DFD97FCD66D4F" blockId="12.[136,878,590,617]" box="[136,878,590,617]" pageId="12" pageNumber="213">Three subspecies are recognized and may be separated as follows:</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C31529E3023F8E28DC9DFD4AFA126C1F" pageId="12" pageNumber="213" type="key">
|
|||
|
|
<key id="FF9E2D59023F8E28DC9DFD4AFA126C1F" pageId="12" pageNumber="213">
|
|||
|
|
<keyStep id="30FB62CD023F8E28DC9DFD4AFA136DD9" pageId="12" pageNumber="213">
|
|||
|
|
<paragraph id="8BB07A68023F8E28DC9DFD4AFA136DE2" blockId="12.[136,1451,658,825]" pageId="12" pageNumber="213">
|
|||
|
|
<keyLead id="30FED95D023F8E28DC9DFD4AFA136DE2" pageId="12" pageNumber="213">
|
|||
|
|
1. Plant forming a robust shrub 15–50 ×
|
|||
|
|
<quantity id="4CF7D78D023F8E28DE54FD4BFD176D8F" box="[577,687,658,680]" metricMagnitude="0" metricUnit="m" metricValue="1.075" metricValueMax="2.0" metricValueMin="0.15" pageId="12" pageNumber="213" unit="cm" value="107.5" valueMax="200.0" valueMin="15.0">15–200 cm</quantity>
|
|||
|
|
, branches 8–12(–20) mm thick, not rhizomatous, spines
|
|||
|
|
<quantity id="4CF7D78D023F8E28D8DBFD4AFA916D8F" box="[1230,1321,659,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.8" metricValueMin="0.6" pageId="12" pageNumber="213" unit="mm" value="12.0" valueMax="18.0" valueMin="6.0">6–18 mm</quantity>
|
|||
|
|
long............. ...............................................................................................................................................................................
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28D8E1FD76FA136DE2" ID-CoL="5H994" authorityName=", Bruyns" authorityYear="2020" baseAuthorityName=", Bruyns" baseAuthorityYear="2020" box="[1268,1451,687,709]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="subSpecies" species="schinzii" subSpecies="bechuanica">
|
|||
|
|
subsp.
|
|||
|
|
<emphasis id="B97BA67A023F8E28D922FD76FA136DE2" bold="true" box="[1335,1451,687,709]" pageId="12" pageNumber="213">bechuanica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BB07A68023F8E28DC9DFD15FA136DD9" blockId="12.[136,1451,658,825]" pageId="12" pageNumber="213">
|
|||
|
|
<keyLead id="30FED95D023F8E28DC9DFD15FA136DD9" pageId="12" pageNumber="213">
|
|||
|
|
1. Plant forming a small clump 5–20(–30) ×
|
|||
|
|
<quantity id="4CF7D78D023F8E28DE6FFD15FD6B6DC5" box="[634,723,716,738]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" metricValueMax="5.0" metricValueMin="0.8" pageId="12" pageNumber="213" unit="cm" value="29.0" valueMax="50.0" valueMin="8.0">8–50 cm</quantity>
|
|||
|
|
, branches 6–12(–20) mm thick, often rhizomatous, spines 3–12(–17) mm long....................................................................................................................................................................................................2.
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="30FB62CD023F8E28DC9DFCDFFA126C1F" pageId="12" pageNumber="213">
|
|||
|
|
<paragraph id="8BB07A68023F8E28DC9DFCDFFA136C3B" blockId="12.[136,1451,658,825]" box="[136,1451,774,796]" pageId="12" pageNumber="213">
|
|||
|
|
<keyLead id="30FED95D023F8E28DC9DFCDFFA136C3B" box="[136,1451,774,796]" pageId="12" pageNumber="213">
|
|||
|
|
2. Stipular prickles
|
|||
|
|
<quantity id="4CF7D78D023F8E28DD62FCDFFE606C3C" box="[375,472,774,796]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="12" pageNumber="213" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
|
|||
|
|
long, spine-shields broad and usually blackish........................................................
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28D8F8FCDFFA136C3B" authorityName=", Bruyns" authorityYear="2020" box="[1261,1451,774,796]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="subSpecies" species="schinzii" subSpecies="schinzioides">
|
|||
|
|
subsp.
|
|||
|
|
<emphasis id="B97BA67A023F8E28D925FCDFFA136C3B" bold="true" box="[1328,1451,774,796]" pageId="12" pageNumber="213">schinzioides</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BB07A68023F8E28DC9DFCFAFA126C1F" blockId="12.[136,1451,658,825]" box="[136,1450,802,825]" pageId="12" pageNumber="213">
|
|||
|
|
<keyLead id="30FED95D023F8E28DC9DFCFAFA126C1F" box="[136,1450,802,825]" pageId="12" pageNumber="213">
|
|||
|
|
2. Stipular prickles
|
|||
|
|
<quantity id="4CF7D78D023F8E28DD62FCFBFE606C1F" box="[375,472,802,824]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="12" pageNumber="213" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 mm</quantity>
|
|||
|
|
long, spine-shields slender and usually grey....................................................................
|
|||
|
|
<taxonomicName id="4C0F01EB023F8E28D90EFCFAFA126C1F" ID-CoL="8YJFS" box="[1307,1450,802,825]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="12" pageNumber="213" phylum="Tracheophyta" rank="subSpecies" species="schinzii" subSpecies="schinzii">
|
|||
|
|
subsp.
|
|||
|
|
<emphasis id="B97BA67A023F8E28D948FCFBFA126C1F" bold="true" box="[1373,1450,802,824]" pageId="12" pageNumber="213">schinzii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
</key>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|