<documentid="B2BEAD5834FD6DF68CBE465E230A4902"ID-DOI="10.5852/ejt.2023.865.2097"ID-GBIF-Dataset="3a63a183-743e-4185-8fdc-1080072ada0a"ID-ISSN="2118-9773"ID-Zenodo-Dep="7867448"ID-ZooBank="845BAE1A-0E6E-484B-BE6D-17F866777B51"IM.bibliography_approvedBy="diego"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="diego"checkinTime="1682512407979"checkinUser="felipe"docAuthor="Rivera-Quiroz, F. Andrés & Álvarez-Padilla, Fernando"docDate="2023"docId="03C8E927FFA3FF8BFDFEFC7BFB6C67CE"docLanguage="en"docName="ejt-865_1-94.pdf"docOrigin="European Journal of Taxonomy 865"docSource="http://dx.doi.org/10.5852/ejt.2023.865.2097"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Anyphaena urieli Rivera-Quiroz & Álvarez-Padilla 2023, sp. nov."docType="treatment"docVersion="11"lastPageNumber="28"masterDocId="FFF1915FFFB4FF90FF81FFF8FF856160"masterDocTitle="Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico"masterLastPageNumber="94"masterPageNumber="1"pageNumber="24"updateTime="1732489834501"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="7FA0A19CA2F567C6F72EC1902A343D0D">Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico</mods:title>
<mods:affiliationid="FCDB2FF3A3613D4B75A0AB47F3056955">Understanding Evolution Research Group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands.</mods:affiliation>
<mods:affiliationid="686F944A680DECAFF20B311BF5412C10">Laboratorio de Aracnología, Departamento de Biología Comparada, Facultad de Ciencias, Universidad Nacional Autónoma de México, Distrito Federal, México Ciudad Universitaria, México D. F. Del. Coyoacán, C. P. 04510, Mexico.</mods:affiliation>
<taxonomicNameid="4C6123B2FFA3FF87FDFEFC7BFCD762FE"ID-CoL="C66LX"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[639,850,899,926]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="urieli"status="sp. nov.">
<figureCitationid="135A44B4FFA3FF87FD1AFC32FC926284"box="[667,791,970,996]"captionStart="Fig"captionStartId="6.[189,232,1926,1952]"captionTargetBox="[197,1396,585,1886]"captionTargetId="figure-103@6.[189,1399,585,1887]"captionTargetPageId="6"captionText="Fig. 1. Anyphaena spp., expanded pedipalps. A–C. A. natachae sp. nov., ♂ (PXF302). A. Ventral view. B. Retrolateral view. C. Prolateral view. – D–F. A. urieli sp. nov., allotype, ♂ (CNAN-T01521). D. Ventral view. E. Retrolateral view. F. Prolateral view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867455"httpUri="https://zenodo.org/record/7867455/files/figure.png"pageId="23"pageNumber="24">Figs 1D–F</figureCitation>
,
<figureCitationid="135A44B4FFA3FF87FCA5FC31FCEA6284"box="[804,879,969,996]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStartId-0="24.[189,232,1891,1917]"captionStartId-1="25.[191,234,1925,1951]"captionStartId-2="26.[189,232,1891,1917]"captionTargetBox-0="[190,1398,739,1847]"captionTargetBox-1="[190,1394,764,1883]"captionTargetBox-2="[232,1361,278,1841]"captionTargetId-0="figure-282@24.[189,1398,737,1847]"captionTargetId-1="figure-195@25.[189,1398,750,1883]"captionTargetId-2="figure-82@26.[225,1370,265,1860]"captionTargetPageId-0="24"captionTargetPageId-1="25"captionTargetPageId-2="26"captionText-0="Fig. 13. Anyphaena urieli sp. nov.A–D. Holotype, ♀ (CNAN-T01542). E. Paratype, ♂ (CNAN-T01582). F. Paratype, ♀ (CNAN-T01581). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C= 1.0 mm; D–E= 0.5 mm; F =0.2 mm."captionText-1="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."captionText-2="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi-0="http://doi.org/10.5281/zenodo.7867497"figureDoi-1="http://doi.org/10.5281/zenodo.7867501"figureDoi-2="http://doi.org/10.5281/zenodo.7867503"httpUri-0="https://zenodo.org/record/7867497/files/figure.png"httpUri-1="https://zenodo.org/record/7867501/files/figure.png"httpUri-2="https://zenodo.org/record/7867503/files/figure.png"pageId="23"pageNumber="24">13–15</figureCitation>
,
<figureCitationid="135A44B4FFA3FF87FCFDFC31FC1C6284"box="[892,921,969,996]"captionStart="Fig"captionStartId="82.[189,232,1018,1044]"captionTargetBox="[193,1392,269,964]"captionTargetId="figure-209@82.[189,1398,265,976]"captionTargetPageId="82"captionText="Fig. 52. Localities for the species of Anyphaena. Black square=Atotonilco Plots I and II, separated by 400 m, nine species registered: A. bifurcata sp. nov., A. bromelicola Platnick, 1977 (ON619634- 38 Supp. file 1), A. catalina Platnick, 1974 (ON619673-75), A. dulceae sp. nov., A. epicardia sp. nov., A. fernandae sp. nov., A. porta sp. nov., A. salgueiroi sp. nov. and A. sofiae sp. nov. White square=Xamaticpac Plots I and II, plots contiguous, eleven species registered: A. adnani sp. nov., A. alachua Platnick, 1974 (ON619665), A. fraterna (Banks, 1896) (ON619671), A. megamedia sp. nov., A. miniducta sp. nov., A. natachae sp. nov., A. porta sp. nov., A. quadrata sp. nov., A. rebecae sp. nov., A. stigma sp. nov. and A. tonoi sp. nov. Black circle =Xilitla, four species registered: A. franciscoi sp. nov., A. ibarrai sp. nov., A. noctua sp. nov. and A. triangularis sp. nov. White circle =Los Tuxtlas, two species registered: A. jimenezi sp. nov. and A. urieli sp. nov."figureDoi="http://doi.org/10.5281/zenodo.7867604"httpUri="https://zenodo.org/record/7867604/files/figure.png"pageId="23"pageNumber="24">52</figureCitation>
<taxonomicNameid="4C6123B2FFA3FF87FE4FFBC2FDB66533"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[462,563,1081,1108]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="urieli"status="sp. nov.">
is broad, more than two thirds of epigastric furrow length, globular, anteriorly projected and ventrally compressed (
<figureCitationid="135A44B4FFA3FF87FCE1FBA4FC756517"box="[864,1008,1116,1143]"captionStart="Fig"captionStartId="24.[189,232,1891,1917]"captionTargetBox="[190,1398,739,1847]"captionTargetId="figure-282@24.[189,1398,737,1847]"captionTargetPageId="24"captionText="Fig. 13. Anyphaena urieli sp. nov.A–D. Holotype, ♀ (CNAN-T01542). E. Paratype, ♂ (CNAN-T01582). F. Paratype, ♀ (CNAN-T01581). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C= 1.0 mm; D–E= 0.5 mm; F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867497"httpUri="https://zenodo.org/record/7867497/files/figure.png"pageId="23"pageNumber="24">Figs 13C, F</figureCitation>
,
<figureCitationid="135A44B4FFA3FF87FC7CFBA5FBD56517"box="[1021,1104,1117,1143]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="23"pageNumber="24">14E–F</figureCitation>
), epigynum anterior half with two deep pits (
<figureCitationid="135A44B4FFA3FF87FE28FB87FD9165FA"box="[425,532,1151,1178]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="23"pageNumber="24">Figs 14E</figureCitation>
(arrow), F, 15G (arrow), H). Similar epigyna are found in
<taxonomicNameid="4C6123B2FFA3FF87FB4BFB78FAF265F9"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[1226,1399,1151,1178]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="triangularis"status="sp. nov.">
<taxonomicNameid="4C6123B2FFA3FF87FDC1FB5BFD1C65DD"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[576,665,1187,1213]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="urieli"status="sp. nov.">
by their anterior projection being ventrally distended, the shape of the anterior pits, and the path of the copulatory ducts extremely short (
<figureCitationid="135A44B4FFA3FF87FB14FB3EFAAC6580"box="[1173,1321,1222,1248]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="23"pageNumber="24">Figs 14A–B</figureCitation>
,
<figureCitationid="135A44B4FFA3FF87FAB4FB3EFF536463"captionStart="Fig"captionStartId="26.[189,232,1891,1917]"captionTargetBox="[232,1361,278,1841]"captionTargetId="figure-82@26.[225,1370,265,1860]"captionTargetPageId="26"captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.7867503"httpUri="https://zenodo.org/record/7867503/files/figure.png"pageId="23"pageNumber="24">15G– H</figureCitation>
). Males of
<taxonomicNameid="4C6123B2FFA3FF87FEE0FB11FE466463"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[353,451,1257,1283]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="urieli"status="sp. nov.">
are differentiated from those of related species by their ventral tegular projection base being bifurcated with a process shaped like a bird head (
<figureCitationid="135A44B4FFA3FF87FB98FAF4FB0E6446"box="[1049,1163,1292,1318]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="23"pageNumber="24">Figs 14A</figureCitation>
,
<figureCitationid="135A44B4FFA3FF87FB19FAF4FB4D6447"box="[1176,1224,1292,1319]"captionStart="Fig"captionStartId="26.[189,232,1891,1917]"captionTargetBox="[232,1361,278,1841]"captionTargetId="figure-82@26.[225,1370,265,1860]"captionTargetPageId="26"captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.7867503"httpUri="https://zenodo.org/record/7867503/files/figure.png"pageId="23"pageNumber="24">15A</figureCitation>
(arrows)), and from
<taxonomicNameid="4C6123B2FFA3FF87FF7AFAD7FE236429"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[251,422,1327,1353]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="23"pageNumber="24"phylum="Arthropoda"rank="species"species="triangularis"status="sp. nov.">
<paragraphid="8BDE5831FFA3FF87FF3CFA1DFE576742"blockId="23.[188,1398,1509,1570]"pageId="23"pageNumber="24">The species epithet is dedicated to Uriel Garcilazo-Cruz, Mexican arachnologist, and former member of the second author’s lab.</paragraph>
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<taxonomicNameid="4C6123B2FFACFF88FE9AF89BFE61661D"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[283,484,1891,1917]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="24"pageNumber="25"phylum="Arthropoda"rank="species"species="urieli"status="sp. nov.">
Total length 5.0. Carapace yellow, ocular area slightly darker, pattern with two darker longitudinal brown parallel bands around cephalic area and fovea, clypeus yellow (
<figureCitationid="135A44B4FFADFF89FB8BFDFEFB1A6340"box="[1034,1183,518,544]"captionStart="Fig"captionStartId="24.[189,232,1891,1917]"captionTargetBox="[190,1398,739,1847]"captionTargetId="figure-282@24.[189,1398,737,1847]"captionTargetPageId="24"captionText="Fig. 13. Anyphaena urieli sp. nov.A–D. Holotype, ♀ (CNAN-T01542). E. Paratype, ♂ (CNAN-T01582). F. Paratype, ♀ (CNAN-T01581). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C= 1.0 mm; D–E= 0.5 mm; F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867497"httpUri="https://zenodo.org/record/7867497/files/figure.png"pageId="25"pageNumber="26">Fig. 13A, D</figureCitation>
). Sternum surface white, intercoxal triangles present on all legs. Labium brown, white at tip, longer than wide. Endites yellow, rectangular, slightly broader at tip (
<figureCitationid="135A44B4FFADFF89FD5AFDB4FCC96306"box="[731,844,588,614]"captionStart="Fig"captionStartId="24.[189,232,1891,1917]"captionTargetBox="[190,1398,739,1847]"captionTargetId="figure-282@24.[189,1398,737,1847]"captionTargetPageId="24"captionText="Fig. 13. Anyphaena urieli sp. nov.A–D. Holotype, ♀ (CNAN-T01542). E. Paratype, ♂ (CNAN-T01582). F. Paratype, ♀ (CNAN-T01581). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C= 1.0 mm; D–E= 0.5 mm; F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867497"httpUri="https://zenodo.org/record/7867497/files/figure.png"pageId="25"pageNumber="26">Fig. 13C</figureCitation>
). Chelicerae yellow, paturon dorsum without pattern (
<figureCitationid="135A44B4FFADFF89FE9CFD97FE2C63EA"box="[285,425,623,650]"captionStart="Fig"captionStartId="24.[189,232,1891,1917]"captionTargetBox="[190,1398,739,1847]"captionTargetId="figure-282@24.[189,1398,737,1847]"captionTargetPageId="24"captionText="Fig. 13. Anyphaena urieli sp. nov.A–D. Holotype, ♀ (CNAN-T01542). E. Paratype, ♂ (CNAN-T01582). F. Paratype, ♀ (CNAN-T01581). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C= 1.0 mm; D–E= 0.5 mm; F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867497"httpUri="https://zenodo.org/record/7867497/files/figure.png"pageId="25"pageNumber="26">Fig. 13B, D</figureCitation>
), promargin with five teeth, retromargin with eight to nine denticles. Leg coloration: yellow, darker from distal section of femora to tarsi. Abdomen yellow, hirsute, dorsal pattern brown, surface delineated with wide band and several chevrons over posterior central half, anterior central half without this pattern, brown pattern of lateral surfaces only at top, lateral sides and ventral surface white covered with faint darker spots, tracheal spiracle at middle (
<figureCitationid="135A44B4FFAFFF8BFC59FEB7FBEF6009"box="[984,1130,335,361]"captionStart="Fig"captionStartId="24.[189,232,1891,1917]"captionTargetBox="[190,1398,739,1847]"captionTargetId="figure-282@24.[189,1398,737,1847]"captionTargetPageId="24"captionText="Fig. 13. Anyphaena urieli sp. nov.A–D. Holotype, ♀ (CNAN-T01542). E. Paratype, ♂ (CNAN-T01582). F. Paratype, ♀ (CNAN-T01581). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C= 1.0 mm; D–E= 0.5 mm; F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867497"httpUri="https://zenodo.org/record/7867497/files/figure.png"pageId="27"pageNumber="28">Fig. 13A–C</figureCitation>
). Copulatory openings under straight hood slightly projected at center. Copulatory ducts sclerotized, short, slightly curved, entering spermathecae on lateral surfaces. Seminal receptacles in anterior third in copulatory ducts inconspicuous. Fertilization ducts short, straight, and emerging from lateral surfaces of spermathecae (
<figureCitationid="135A44B4FFAFFF8BFF46FE24FED06096"box="[199,341,476,502]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="27"pageNumber="28">Figs 14E–F</figureCitation>
,
<figureCitationid="135A44B4FFAFFF8BFEE4FE23FE456096"box="[357,448,475,502]"captionStart="Fig"captionStartId="26.[189,232,1891,1917]"captionTargetBox="[232,1361,278,1841]"captionTargetId="figure-82@26.[225,1370,265,1860]"captionTargetPageId="26"captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.7867503"httpUri="https://zenodo.org/record/7867503/files/figure.png"pageId="27"pageNumber="28">15G–H</figureCitation>
). Cephalothorax length 2.24, thoracic width 1.8, cephalic width 0.97. Clypeus height 0.07. Eye diameters: AME 0.09, ALE 0.12, PME 0.12, PLE 0.12. Eye interdistances: AME– AME 0.05, AME–ALE 0.02, ALE–PLE 0.04, PME–PME 0.12, PME–PLE 0.09. Femur lengths: I 2.58, II 2.42, III 1.81, IV 2.65. Leg spination: femur I d1-1-1, p0-2-1, r0-1-1. Tibia I v2-2-0, p1-1-1, r1-1-1. Metatarsus I v2-2-0, p1-1-1, r1-1-1. Femur II d1-1-1, p0-1-1, r0-1-1. Tibia II v2-2-0, p1-1-1, r1-1-1. Metatarsus II v2-2-0, p1-1-1, r1-1-1. Femur III d1-1-1, p0-1-1, r0-1-1. Tibia III v1-1-2, p0-1-1, r0-1-1. Metatarsus III v2-2-2, p1-1-2, r1-1-2. Femur IV d1-1-1, p0-0-1, r0-0-1. Tibia IV v2-2-2, p0-1-1, r0-1-1. Metatarsus IV v2-2-2, p1-1-2, r1-1-2.
<taxonomicNameid="4C6123B2FFADFF89FEA9F87DFE7266FF"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[296,503,1925,1951]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="25"pageNumber="26"phylum="Arthropoda"rank="species"species="urieli"status="sp. nov.">
<taxonomicNameid="4C6123B2FFAEFF8AFEBBF89BFD97661D"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[314,530,1891,1917]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="26"pageNumber="27"phylum="Arthropoda"rank="species"species="urieli"status="sp. nov.">
Total length 5.8. Cephalothorax and abdomen coloration as in female (
<figureCitationid="135A44B4FFAFFF8BFC6CFCCEFBD06230"box="[1005,1109,822,848]"captionStart="Fig"captionStartId="24.[189,232,1891,1917]"captionTargetBox="[190,1398,739,1847]"captionTargetId="figure-282@24.[189,1398,737,1847]"captionTargetPageId="24"captionText="Fig. 13. Anyphaena urieli sp. nov.A–D. Holotype, ♀ (CNAN-T01542). E. Paratype, ♂ (CNAN-T01582). F. Paratype, ♀ (CNAN-T01581). A. Dorsal habitus. B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Prosoma, anterior view. F. Epigynum, lateral view. Scale bars: A–C= 1.0 mm; D–E= 0.5 mm; F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867497"httpUri="https://zenodo.org/record/7867497/files/figure.png"pageId="27"pageNumber="28">Fig. 13E</figureCitation>
). Chelicerae retromargin with eleven to twelve denticles. Coxa III tubercle roughly rectangular with posterior apical spur. Coxa IV with a long, curled spur curved at tip (
<figureCitationid="135A44B4FFAFFF8BFD26FC84FCB462F7"box="[679,817,892,919]"captionStart="Fig"captionStartId="26.[189,232,1891,1917]"captionTargetBox="[232,1361,278,1841]"captionTargetId="figure-82@26.[225,1370,265,1860]"captionTargetPageId="26"captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.7867503"httpUri="https://zenodo.org/record/7867503/files/figure.png"pageId="27"pageNumber="28">Fig. 15E–F</figureCitation>
). Median apophysis hook-shaped in ventral view. Embolus long, translucent and filiform (
<figureCitationid="135A44B4FFAFFF8BFD1AFC67FC8862DA"box="[667,781,927,954]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="27"pageNumber="28">Figs 14A</figureCitation>
,
<figureCitationid="135A44B4FFAFFF8BFC9BFC67FCD462DA"box="[794,849,927,954]"captionStart="Fig"captionStartId="26.[189,232,1891,1917]"captionTargetBox="[232,1361,278,1841]"captionTargetId="figure-82@26.[225,1370,265,1860]"captionTargetPageId="26"captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.7867503"httpUri="https://zenodo.org/record/7867503/files/figure.png"pageId="27"pageNumber="28">15A</figureCitation>
). RTA anterior branch curved with sclerotized margin, apical edge of posterior branch of RTA with two triangular tips (
<figureCitationid="135A44B4FFAFFF8BFBA2FC3AFB5B62BD"box="[1059,1246,962,989]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="27"pageNumber="28">Figs 14A, C–D</figureCitation>
,
<figureCitationid="135A44B4FFAFFF8BFB6AFC3AFAEF62BD"box="[1259,1386,962,989]"captionStart="Fig"captionStartId="26.[189,232,1891,1917]"captionTargetBox="[232,1361,278,1841]"captionTargetId="figure-82@26.[225,1370,265,1860]"captionTargetPageId="26"captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.7867503"httpUri="https://zenodo.org/record/7867503/files/figure.png"pageId="27"pageNumber="28">15A, C–D</figureCitation>
<figureCitationid="135A44B4FFAFFF8BFD6FFC1EFCFB6560"box="[750,894,998,1024]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="27"pageNumber="28">Figs 14B–C</figureCitation>
,
<figureCitationid="135A44B4FFAFFF8BFC0BFC1DFC676560"box="[906,994,997,1024]"captionStart="Fig"captionStartId="26.[189,232,1891,1917]"captionTargetBox="[232,1361,278,1841]"captionTargetId="figure-82@26.[225,1370,265,1860]"captionTargetPageId="26"captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.7867503"httpUri="https://zenodo.org/record/7867503/files/figure.png"pageId="27"pageNumber="28">15B–C</figureCitation>
). Pedipalp tibia proportion slightly longer than wide, stridulatory ridges extending from ventral branch of median tegular apophysis to tibial distal margin (
<figureCitationid="135A44B4FFAFFF8BFE3BFBD4FDF96526"box="[442,636,1068,1094]"captionStart="Fig"captionStartId="25.[191,234,1925,1951]"captionTargetBox="[190,1394,764,1883]"captionTargetId="figure-195@25.[189,1398,750,1883]"captionTargetPageId="25"captionText="Fig. 14. Anyphaena urieli sp. nov. A–D. Paratype, ♂ (CNAN-T01582). E–F. Paratype, ♀ (CNAN- T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867501"httpUri="https://zenodo.org/record/7867501/files/figure.png"pageId="27"pageNumber="28">Figs 14A–B, D</figureCitation>
,
<figureCitationid="135A44B4FFAFFF8BFD0CFBD4FC8A6526"box="[653,783,1068,1095]"captionStart="Fig"captionStartId="26.[189,232,1891,1917]"captionTargetBox="[232,1361,278,1841]"captionTargetId="figure-82@26.[225,1370,265,1860]"captionTargetPageId="26"captionText="Fig. 15. Anyphaena urieli sp. nov. A–F. Paratype, ♂ (CNAN-T01582). G–H. Paratype, ♀ (CNAN-T01581). A. Pedipalp, ventral view. B. Pedipalp, prolateral view. C. Pedipalp, dorsal view. D. Pedipalp, retrolateral view. E. Coxae, ventral view. F. Coxae, lateral view. G. Epigynum, ventral view. H. Epigynum, dorsal view. Scale bars: A–D, G–H =0.2 mm; E–F =1.0 mm."figureDoi="http://doi.org/10.5281/zenodo.7867503"httpUri="https://zenodo.org/record/7867503/files/figure.png"pageId="27"pageNumber="28">15A–B, D</figureCitation>
). Cephalothorax length 2.54, thoracic width 2.05, cephalic width 0.9. Clypeus height 0.06. Eye diameters: AME 0.07, ALE 0.17, PME 0.15, PLE 0.15. Eye interdistances: AME–AME 0.05, AME–ALE 0.02, ALE–PLE 0.05, PME–PME 0.11, PME–PLE 0.09. Femur lengths: I 3.24, II 2.9, III 2.19, IV 3.1. Leg spination as in female except: femur I p0-1-2. Femur II p0-1-2. Metatarsus II r1-1-0. Femur III r0-0-1. Tibia III v1-2-2, p1-1-0, r1-1-0. Femur IV r0- 1-1. Tibia IV p1-1-1, r1-1-1.
This species is found in primary tropical wet forest (
<figureCitationid="135A44B4FFAFFF8BFCA2F9DDFCFE675F"box="[803,891,1572,1599]"captionStart="Fig"captionStartId="82.[189,232,1018,1044]"captionTargetBox="[193,1392,269,964]"captionTargetId="figure-209@82.[189,1398,265,976]"captionTargetPageId="82"captionText="Fig. 52. Localities for the species of Anyphaena. Black square=Atotonilco Plots I and II, separated by 400 m, nine species registered: A. bifurcata sp. nov., A. bromelicola Platnick, 1977 (ON619634- 38 Supp. file 1), A. catalina Platnick, 1974 (ON619673-75), A. dulceae sp. nov., A. epicardia sp. nov., A. fernandae sp. nov., A. porta sp. nov., A. salgueiroi sp. nov. and A. sofiae sp. nov. White square=Xamaticpac Plots I and II, plots contiguous, eleven species registered: A. adnani sp. nov., A. alachua Platnick, 1974 (ON619665), A. fraterna (Banks, 1896) (ON619671), A. megamedia sp. nov., A. miniducta sp. nov., A. natachae sp. nov., A. porta sp. nov., A. quadrata sp. nov., A. rebecae sp. nov., A. stigma sp. nov. and A. tonoi sp. nov. Black circle =Xilitla, four species registered: A. franciscoi sp. nov., A. ibarrai sp. nov., A. noctua sp. nov. and A. triangularis sp. nov. White circle =Los Tuxtlas, two species registered: A. jimenezi sp. nov. and A. urieli sp. nov."figureDoi="http://doi.org/10.5281/zenodo.7867604"httpUri="https://zenodo.org/record/7867604/files/figure.png"pageId="27"pageNumber="28">Fig. 52</figureCitation>
<paragraphid="8BDE5831FFAFFF8BFF3CF96DFB6C67CE"blockId="27.[189,1257,1684,1711]"box="[189,1257,1684,1711]"pageId="27"pageNumber="28">Most specimens were collected over vegetation by direct searching and beating vegetation.</paragraph>