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<document id="1C3CA097E601CDE006ADD5AD95264F7B" ID-DOI="10.5281/zenodo.281652" ID-GBIF-Dataset="1d6d96e1-5b92-4633-b769-d9e7ab8fa254" ID-ISSN="1175-5326" ID-Zenodo-Dep="281652" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460495559621" checkinUser="plazi" docAuthor="Chan, Benny K. K. &amp; Hayashi, Ryota" docDate="2012" docId="03FC87D4FFD68A16C7D721995219FD75" docLanguage="en" docName="zt03367p048.pdf" docOrigin="Zootaxa 3367" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oxynaspis celata Darwin 1852" docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FFC5FFACFFDF8A1AC74020105772FFD1" masterDocTitle="Epibiotic barnacles (Crustacea: Cirripedia: Thoracica) collected by the Kumejima 2009 Expedition, with descriptions of two new species *" masterLastPageNumber="48" masterPageNumber="21" pageNumber="30" updateTime="1698310858039" updateUser="plazi">
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<mods:title id="A7FE971A672C002DD79601C6CADCDB33">Epibiotic barnacles (Crustacea: Cirripedia: Thoracica) collected by the Kumejima 2009 Expedition, with descriptions of two new species *</mods:title>
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<mods:namePart id="1CE439C0CE12426A18F94E8CF9409EC5">Chan, Benny K. K.</mods:namePart>
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<mods:namePart id="0C2442F6AB5B94DBD7AC2FD9BBE50E67">Hayashi, Ryota</mods:namePart>
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<treatment id="03FC87D4FFD68A16C7D721995219FD75" ID-DOI="http://doi.org/10.5281/zenodo.6174489" ID-GBIF-Taxon="119520509" ID-Zenodo-Dep="6174489" LSID="urn:lsid:plazi:treatment:03FC87D4FFD68A16C7D721995219FD75" httpUri="http://treatment.plazi.org/id/03FC87D4FFD68A16C7D721995219FD75" lastPageId="12" lastPageNumber="33" pageId="9" pageNumber="30">
<subSubSection id="C34F6549FFD68A13C7D721995632FE15" pageId="9" pageNumber="30" type="nomenclature">
<paragraph id="8BEA36C2FFD68A13C7D721995563FE73" blockId="9.[151,529,392,452]" box="[151,529,392,419]" pageId="9" pageNumber="30">
<heading id="D0A281AEFFD68A13C7D721995563FE73" bold="true" box="[151,529,392,419]" fontSize="11" level="1" pageId="9" pageNumber="30" reason="1">
<taxonomicName id="4C554D41FFD68A13C7D721995563FE73" authority="Darwin, 1852" authorityName="Darwin" authorityYear="1852" box="[151,529,392,419]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD68A13C7D721995563FE73" bold="true" box="[151,529,392,419]" pageId="9" pageNumber="30">
<emphasis id="B921EAD0FFD68A13C7D721995611FE72" bold="true" box="[151,355,393,419]" italics="true" pageId="9" pageNumber="30">Oxynaspis celata</emphasis>
<bibRefCitation id="EFC44B33FFD68A13C62A21985563FE73" author="Darwin" box="[362,529,392,418]" pageId="9" pageNumber="30" refString="Darwin, C. (1852) A monograph on the sub-class Cirripedia with figures of all species. The Lepadidae: or pedunculated barnacles. Ray Society, London. 400 pp. (1851, published 1852)" type="book" year="1852">Darwin, 1852</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BEA36C2FFD68A13C7D721BC5632FE15" blockId="9.[151,529,392,452]" box="[151,320,428,452]" pageId="9" pageNumber="30">
(
<figureCitation id="136E2A47FFD68A13C7DF21BC579FFE15" box="[159,237,428,452]" captionStart="FIGURE 2" captionStartId="2.[151,244,1869,1890]" captionTargetBox="[153,1433,917,1847]" captionTargetId="figure@2.[153,1433,917,1848]" captionTargetPageId="2" captionText="FIGURE 2. Kumejima epibiotic barnacles. A, Calantica arcuata n. sp. (Left, holotype, RUMF-ZC- 1554, middle, paratype, RUMF- ZC- 1555, note the tip of carina was broken, right, paratype, NMNS- 6690 - 001); B, Euscalpellum c. f. squamoum Hiro, 1937 (left, RUMF-ZC- 1556, right ASIZCR- 000225); C, Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557); D, Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558); E, Poecilasma obliqua (Hoek, 1907) (RUMF-ZC- 1560); F, Platylepas hexastylos (Fabricius, 1798) (RUMF-ZC- 1561). A-E, lateral view for stalked barnacles; F, anterior view for Platylepas hexastylos. Scale bars in mm to indicate the size of specimens." httpUri="https://zenodo.org/record/281654/files/figure.png" pageId="9" pageNumber="30">Figs. 2</figureCitation>
C, 7, 8)
</paragraph>
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<subSubSection id="C34F6549FFD68A13C7D721FE56ABFDEF" pageId="9" pageNumber="30" type="reference_group">
<paragraph id="8BEA36C2FFD68A13C7D721FE56ABFDEF" blockId="9.[151,1436,494,574]" pageId="9" pageNumber="30">
<treatmentCitationGroup id="AB4511ECFFD68A13C7D721FE56ABFDEF" pageId="9" pageNumber="30">
<taxonomicName id="4C554D41FFD68A13C7D721FE568CFDD2" authority="Darwin, 1852: 134" authorityName="Darwin" authorityPageNumber="134" authorityYear="1852" box="[151,510,494,515]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD68A13C7D721FE564FFDD2" box="[151,317,494,515]" italics="true" pageId="9" pageNumber="30">Oxynaspis celata</emphasis>
<treatmentCitation id="0AF410D3FFD68A13C60421FE568CFDD2" author="Darwin" box="[324,510,494,515]" page="134" pageId="9" pageNumber="30" year="1852">
<bibRefCitation id="EFC44B33FFD68A13C60421FE56BCFDD2" author="Darwin" box="[324,462,494,515]" pageId="9" pageNumber="30" refString="Darwin, C. (1852) A monograph on the sub-class Cirripedia with figures of all species. The Lepadidae: or pedunculated barnacles. Ray Society, London. 400 pp. (1851, published 1852)" type="book" year="1852">Darwin, 1852</bibRefCitation>
: 134
</treatmentCitation>
</taxonomicName>
, pl. 3, fig. 1. —
<treatmentCitation id="0AF410D3FFD68A13C5E521FE5421FDD2" author="Gruvel" box="[677,851,494,515]" page="103" pageId="9" pageNumber="30" year="1905">
<bibRefCitation id="EFC44B33FFD68A13C5E521FE5451FDD2" author="Gruvel" box="[677,803,494,515]" pageId="9" pageNumber="30" refString="Gruvel, A. (1905) Monographie des Cirrhipedes ou Thecostraces. Masson et Cie, Paris, 472 pp." type="book" year="1905">Gruvel 1905</bibRefCitation>
: 103
</treatmentCitation>
, fig. 114. —
<treatmentCitation id="0AF410D3FFD68A13C49721FE53A8FDD2" author="Nilsson-Cantell" box="[983,1242,494,515]" page="226" pageId="9" pageNumber="30" year="1921">
<bibRefCitation id="EFC44B33FFD68A13C49721FE53DBFDD2" author="Nilsson-Cantell" box="[983,1193,494,515]" pageId="9" pageNumber="30" refString="Nilsson-Cantell, C. A. (1921) Cirripeden-Studien. Zur Kenntnis der Biologie, Antomie und Systematic dieser Gruppe. Zoologiska Bidrag Fran Uppsala, 7, 75 - 390." type="journal article" year="1921">Nilsson-Cantell 1921</bibRefCitation>
: 226
</treatmentCitation>
, fig. 37. — Nilsson
<treatmentCitation id="0AF410D3FFD68A13C787221B561BFDF1" author="Cantell" box="[199,361,523,544]" page="94" pageId="9" pageNumber="30" year="1937">
<bibRefCitation id="EFC44B33FFD68A13C787221B5637FDF1" author="Cantell" box="[199,325,523,544]" pageId="9" pageNumber="30" refString="Nilsson-Cantell, C. A. (1937) On a second collection of Indo-Malayan Cirripedes from the Raffles Museum. Bulletin of the Raffles Museum, 13, 93 - 96." type="journal article" year="1937">Cantell 1937</bibRefCitation>
: 94
</treatmentCitation>
. —
<treatmentCitation id="0AF410D3FFD68A13C6D0221B5550FDF1" author="Hiro" box="[400,546,523,544]" page="51" pageId="9" pageNumber="30" year="1937">
<bibRefCitation id="EFC44B33FFD68A13C6D0221B568DFDF1" author="Hiro" box="[400,511,523,544]" pageId="9" pageNumber="30" refString="Hiro, F. (1937 a) Order Thoracica I. (Cirripedia Pedunculata) Subclass Cirripedia (Class Crustacea). Fauna Nipponica, 9, 1 - 116. (In Japanese)" type="journal article" year="1937" yearSuffix="a">Hiro 1937a</bibRefCitation>
: 51
</treatmentCitation>
, fig. 42. —
<treatmentCitation id="0AF410D3FFD68A13C5D4221B5448FDF1" author="Totton" box="[660,826,523,544]" page="473" pageId="9" pageNumber="30" year="1940">
<bibRefCitation id="EFC44B33FFD68A13C5D4221B547EFDF1" author="Totton" box="[660,780,523,544]" pageId="9" pageNumber="30" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
: 473
</treatmentCitation>
, fig. 9. —
<treatmentCitation id="0AF410D3FFD68A13C4E0221B534DFDF1" author="Zevina" box="[928,1087,523,544]" page="35" pageId="9" pageNumber="30" year="1968">
<bibRefCitation id="EFC44B33FFD68A13C4E0221B536EFDF1" author="Zevina" box="[928,1052,523,544]" pageId="9" pageNumber="30" refString="Zevina, G. B. (1968) New species of Lepadomorpha (Cirripedia, Thoracica) from the Bay of Tonkin. Crustaceana, 15, 35 - 40." type="journal article" year="1968">Zevina 1968</bibRefCitation>
: 35
</treatmentCitation>
. —
<treatmentCitation id="0AF410D3FFD68A13C326221B5276FDF1" author="Zevina" box="[1126,1284,523,544]" page="31" pageId="9" pageNumber="30" year="1982">
<bibRefCitation id="EFC44B33FFD68A13C326221B5393FDF1" author="Zevina" box="[1126,1249,523,544]" pageId="9" pageNumber="30" refString="Zevina, G. B. (1982) Barnacles of the suborder Lepadomorpha (Cirripedia, Thoracica) of the world oceans. II. Opredeliteli Faune SSSR, 133, 1 - 221. (in Russian)." type="journal article" year="1982">Zevina 1982</bibRefCitation>
: 31
</treatmentCitation>
, fig. 18. — Van Soyc &amp;
<treatmentCitation id="0AF410D3FFD68A13C655223956A7FDEF" author="Dekelboum" box="[277,469,553,574]" page="5" pageId="9" pageNumber="30" year="2011">
<bibRefCitation id="EFC44B33FFD68A13C655223956CCFDEF" author="Dekelboum" box="[277,446,553,574]" pageId="9" pageNumber="30" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article" year="2011">Dekelboum 2011</bibRefCitation>
: 5
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD68A13C7D722735412FD4E" pageId="9" pageNumber="30" type="materials_examined">
<paragraph id="8BEA36C2FFD68A13C7D722735412FD4E" blockId="9.[151,1437,611,2019]" pageId="9" pageNumber="30">
<emphasis id="B921EAD0FFD68A13C7D7227356F9FDAD" bold="true" box="[151,395,611,636]" pageId="9" pageNumber="30">Material examined.</emphasis>
RUMF-ZC-1557, one specimen (CL
<quantity id="4CAD9B27FFD68A13C426227454B5FDAD" box="[870,967,612,636]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="9" pageNumber="30" unit="mm" value="7.9">7.9 mm</quantity>
) on an antipatharian coral, trawl 31,
<geoCoordinate id="EE615005FFD68A13C7D722975659FD4E" box="[151,299,647,671]" direction="north" orientation="latitude" pageId="9" pageNumber="30" precision="1" value="26.313084">26°18.785´N</geoCoordinate>
,
<geoCoordinate id="EE615005FFD68A13C676229756A4FD4E" box="[310,470,647,671]" direction="east" orientation="longitude" pageId="9" pageNumber="30" precision="1" value="126.88748">126°53.249´E</geoCoordinate>
,
<date id="FFEB1002FFD68A13C6A122975509FD4E" box="[481,635,647,671]" pageId="9" pageNumber="30" value="2009-11-13">13 Nov. 2009</date>
, depth
<quantity id="4CAD9B27FFD68A13C58C22975429FD4E" box="[716,859,647,671]" metricMagnitude="1" metricUnit="m" metricValue="7.284999999999999" metricValueMax="7.529999999999999" metricValueMin="7.040000000000001" pageId="9" pageNumber="30" unit="m" value="72.85" valueMax="75.3" valueMin="70.4">70.4–75.3 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD68A13C78722BA540DFC81" pageId="9" pageNumber="30" type="diagnosis">
<paragraph id="8BEA36C2FFD68A13C78722BA540DFC81" blockId="9.[151,1437,611,2019]" pageId="9" pageNumber="30">
<emphasis id="B921EAD0FFD68A13C78722BA56B5FD12" bold="true" box="[199,455,682,707]" pageId="9" pageNumber="30">Diagnosis (emended).</emphasis>
Capitulum completely or nearly covered by 5 thin, fragile plates; surfaces often overgrown by antipatharian corals. Tergum with apex distally recurved; length of occludent margin half length of scutal margin. Umbo of scutum located in middle portion of occludent margin. Carina bowed; umbo located in proximal one third of carina, length of distal arm of carina about twice length of basal arm. Filamentary appendages absent. Caudal appendages minute, oval shaped, bundles of setae apically.
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD68A13C787234C56BAF9C7" pageId="9" pageNumber="30" type="description">
<paragraph id="8BEA36C2FFD68A13C787234C5612FB62" blockId="9.[151,1437,611,2019]" pageId="9" pageNumber="30">
<emphasis id="B921EAD0FFD68A13C787234C562BFCA4" bold="true" box="[199,345,860,885]" pageId="9" pageNumber="30">Description.</emphasis>
Capitulum covered by 5 fully calcified, thin, fragile plates - carina, paired tergum and scutum (
<figureCitation id="136E2A47FFD68A13C7DF236F5797FC46" box="[159,229,895,919]" captionStart="FIGURE 2" captionStartId="2.[151,244,1869,1890]" captionTargetBox="[153,1433,917,1847]" captionTargetId="figure@2.[153,1433,917,1848]" captionTargetPageId="2" captionText="FIGURE 2. Kumejima epibiotic barnacles. A, Calantica arcuata n. sp. (Left, holotype, RUMF-ZC- 1554, middle, paratype, RUMF- ZC- 1555, note the tip of carina was broken, right, paratype, NMNS- 6690 - 001); B, Euscalpellum c. f. squamoum Hiro, 1937 (left, RUMF-ZC- 1556, right ASIZCR- 000225); C, Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557); D, Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558); E, Poecilasma obliqua (Hoek, 1907) (RUMF-ZC- 1560); F, Platylepas hexastylos (Fabricius, 1798) (RUMF-ZC- 1561). A-E, lateral view for stalked barnacles; F, anterior view for Platylepas hexastylos. Scale bars in mm to indicate the size of specimens." httpUri="https://zenodo.org/record/281654/files/figure.png" pageId="9" pageNumber="30">Fig. 2</figureCitation>
C); surface of plates overgrown by antipatharian corals. Tergum narrow, occludent margin half length of scutal margin, occludent margin slightly convex, apex of tergum recurved distally. Scutum trapezoid, umbo at middle of occludent margin, basal margin perpendicular to occludent margin, carinal margin convex. Carina bowed, umbo located in proximal one third of carinal margin, length of distal arm twice length of basal arm (
<figureCitation id="136E2A47FFD68A13C24423FA5235FBD3" box="[1284,1351,1002,1026]" captionStart="FIGURE 2" captionStartId="2.[151,244,1869,1890]" captionTargetBox="[153,1433,917,1847]" captionTargetId="figure@2.[153,1433,917,1848]" captionTargetPageId="2" captionText="FIGURE 2. Kumejima epibiotic barnacles. A, Calantica arcuata n. sp. (Left, holotype, RUMF-ZC- 1554, middle, paratype, RUMF- ZC- 1555, note the tip of carina was broken, right, paratype, NMNS- 6690 - 001); B, Euscalpellum c. f. squamoum Hiro, 1937 (left, RUMF-ZC- 1556, right ASIZCR- 000225); C, Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557); D, Oxynaspis ryukyuensis n. sp. (holotype, RUMF-ZC- 1558); E, Poecilasma obliqua (Hoek, 1907) (RUMF-ZC- 1560); F, Platylepas hexastylos (Fabricius, 1798) (RUMF-ZC- 1561). A-E, lateral view for stalked barnacles; F, anterior view for Platylepas hexastylos. Scale bars in mm to indicate the size of specimens." httpUri="https://zenodo.org/record/281654/files/figure.png" pageId="9" pageNumber="30">Fig. 2</figureCitation>
C). Maxilla rounded, simple setae on all margins (
<figureCitation id="136E2A47FFD68A13C5E6241D559BFBF7" box="[678,745,1037,1062]" captionStart="FIGURE 7" captionStartId="10.[151,244,1936,1957]" captionTargetBox="[186,1402,156,1908]" captionTargetId="figure@10.[146,1462,119,1929]" captionTargetPageId="10" captionText="FIGURE 7. Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="9" pageNumber="30">Fig. 7</figureCitation>
A, B). Maxillule slightly notched, 2 large simple setae above notch, notch shallow with several fine simple setae, more than 10 longer, simple setae below notch (
<figureCitation id="136E2A47FFD68A13C25124215227FB98" box="[1297,1365,1073,1097]" captionStart="FIGURE 7" captionStartId="10.[151,244,1936,1957]" captionTargetBox="[186,1402,156,1908]" captionTargetId="figure@10.[146,1462,119,1929]" captionTargetPageId="10" captionText="FIGURE 7. Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="9" pageNumber="30">Fig. 7</figureCitation>
C, D). Mandible with 4 teeth, first well separated from remainder (
<figureCitation id="136E2A47FFD68A13C405244454FBFBBD" box="[837,905,1108,1132]" captionStart="FIGURE 7" captionStartId="10.[151,244,1936,1957]" captionTargetBox="[186,1402,156,1908]" captionTargetId="figure@10.[146,1462,119,1929]" captionTargetPageId="10" captionText="FIGURE 7. Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="9" pageNumber="30">Fig. 7</figureCitation>
E), lower margin very short, smooth (
<figureCitation id="136E2A47FFD68A13C2782444520EFBBD" box="[1336,1404,1108,1132]" captionStart="FIGURE 7" captionStartId="10.[151,244,1936,1957]" captionTargetBox="[186,1402,156,1908]" captionTargetId="figure@10.[146,1462,119,1929]" captionTargetPageId="10" captionText="FIGURE 7. Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="9" pageNumber="30">Fig. 7</figureCitation>
F), inferior angle terminating in 1 pectination (
<figureCitation id="136E2A47FFD68A13C53F246855B1FB41" box="[639,707,1144,1168]" captionStart="FIGURE 7" captionStartId="10.[151,244,1936,1957]" captionTargetBox="[186,1402,156,1908]" captionTargetId="figure@10.[146,1462,119,1929]" captionTargetPageId="10" captionText="FIGURE 7. Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="9" pageNumber="30">Fig. 7</figureCitation>
F). Mandibular palps elongated (
<figureCitation id="136E2A47FFD68A13C37324685307FB41" box="[1075,1141,1144,1168]" captionStart="FIGURE 7" captionStartId="10.[151,244,1936,1957]" captionTargetBox="[186,1402,156,1908]" captionTargetId="figure@10.[146,1462,119,1929]" captionTargetPageId="10" captionText="FIGURE 7. Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="9" pageNumber="30">Fig. 7</figureCitation>
G, H); dense, simple setae distally (
<figureCitation id="136E2A47FFD68A13C7B9248B564EFB62" box="[249,316,1179,1203]" captionStart="FIGURE 7" captionStartId="10.[151,244,1936,1957]" captionTargetBox="[186,1402,156,1908]" captionTargetId="figure@10.[146,1462,119,1929]" captionTargetPageId="10" captionText="FIGURE 7. Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="9" pageNumber="30">Fig. 7</figureCitation>
H).
</paragraph>
<paragraph id="8BEA36C2FFD68A13C78724AF56BAF9C7" blockId="9.[151,1437,611,2019]" pageId="9" pageNumber="30">
Cirrus I with anterior and posterior rami subequal, setae simple, anterior and posterior rami 12-and 8-segmented, respectively (
<figureCitation id="136E2A47FFD68A13C6D224F256A7FB2B" box="[402,469,1250,1274]" captionStart="FIGURE 8" captionStartId="11.[151,244,1933,1954]" captionTargetBox="[182,1400,161,1909]" captionTargetId="figure@11.[143,1459,125,1935]" captionTargetPageId="11" captionText="FIGURE 8. Oxynaspis celata (RUMF-ZC- 1557). A, cirrus I; B, simple type setae on cirrus I; C, cirrus II; D, simple type setae on cirrus II; E, simple type setae on cirrus IV; F, intermediate segment of posterior ramus of cirrus VI; G, caudal appendage; H, pedicel of penis. Scale bars in μm." httpUri="https://zenodo.org/record/281660/files/figure.png" pageId="9" pageNumber="30">Fig. 8</figureCitation>
A, B). Cirri II–VI ctenopod. Cirrus II with anterior and posterior rami 9-and 13-segmented, respectively (
<figureCitation id="136E2A47FFD68A13C6CD251656A2FACF" box="[397,464,1286,1310]" captionStart="FIGURE 8" captionStartId="11.[151,244,1933,1954]" captionTargetBox="[182,1400,161,1909]" captionTargetId="figure@11.[143,1459,125,1935]" captionTargetPageId="11" captionText="FIGURE 8. Oxynaspis celata (RUMF-ZC- 1557). A, cirrus I; B, simple type setae on cirrus I; C, cirrus II; D, simple type setae on cirrus II; E, simple type setae on cirrus IV; F, intermediate segment of posterior ramus of cirrus VI; G, caudal appendage; H, pedicel of penis. Scale bars in μm." httpUri="https://zenodo.org/record/281660/files/figure.png" pageId="9" pageNumber="30">Fig. 8</figureCitation>
C), setae simple (
<figureCitation id="136E2A47FFD68A13C5D3251655A6FACF" box="[659,724,1286,1310]" captionStart="FIGURE 8" captionStartId="11.[151,244,1933,1954]" captionTargetBox="[182,1400,161,1909]" captionTargetId="figure@11.[143,1459,125,1935]" captionTargetPageId="11" captionText="FIGURE 8. Oxynaspis celata (RUMF-ZC- 1557). A, cirrus I; B, simple type setae on cirrus I; C, cirrus II; D, simple type setae on cirrus II; E, simple type setae on cirrus IV; F, intermediate segment of posterior ramus of cirrus VI; G, caudal appendage; H, pedicel of penis. Scale bars in μm." httpUri="https://zenodo.org/record/281660/files/figure.png" pageId="9" pageNumber="30">Fig. 8</figureCitation>
D) and serrulate with very fine setules (
<figureCitation id="136E2A47FFD68A13C3D0251653A0FACF" box="[1168,1234,1286,1310]" captionStart="FIGURE 8" captionStartId="11.[151,244,1933,1954]" captionTargetBox="[182,1400,161,1909]" captionTargetId="figure@11.[143,1459,125,1935]" captionTargetPageId="11" captionText="FIGURE 8. Oxynaspis celata (RUMF-ZC- 1557). A, cirrus I; B, simple type setae on cirrus I; C, cirrus II; D, simple type setae on cirrus II; E, simple type setae on cirrus IV; F, intermediate segment of posterior ramus of cirrus VI; G, caudal appendage; H, pedicel of penis. Scale bars in μm." httpUri="https://zenodo.org/record/281660/files/figure.png" pageId="9" pageNumber="30">Fig. 8</figureCitation>
E). Cirrus III with anterior and posterior rami 14- and 15-segmented, respectively, both rami with simple and serrulate setae. Cirrus IV with anterior and posterior rami 17-segmented, setae simple and serrulate. Cirrus V with both rami 16-segmented, setae simple and serrulate. Cirrus VI with anterior and posterior rami 16- and 17-segmented, setae simple and serrulate; intermediate segment of posterior ramus of cirrus VI with 3 pairs of long serrulate setae and 2 pairs of short simple setae (
<figureCitation id="136E2A47FFD68A13C6CB25A756BFFA1E" box="[395,461,1463,1487]" captionStart="FIGURE 8" captionStartId="11.[151,244,1933,1954]" captionTargetBox="[182,1400,161,1909]" captionTargetId="figure@11.[143,1459,125,1935]" captionTargetPageId="11" captionText="FIGURE 8. Oxynaspis celata (RUMF-ZC- 1557). A, cirrus I; B, simple type setae on cirrus I; C, cirrus II; D, simple type setae on cirrus II; E, simple type setae on cirrus IV; F, intermediate segment of posterior ramus of cirrus VI; G, caudal appendage; H, pedicel of penis. Scale bars in μm." httpUri="https://zenodo.org/record/281660/files/figure.png" pageId="9" pageNumber="30">Fig. 8</figureCitation>
F). Caudal appendage oval uniarticulate, short (one third height of proximal segment of pedicel of cirrus VI), 4 simple setae distally (
<figureCitation id="136E2A47FFD68A13C5CF25CA55A3FA23" box="[655,721,1498,1522]" captionStart="FIGURE 8" captionStartId="11.[151,244,1933,1954]" captionTargetBox="[182,1400,161,1909]" captionTargetId="figure@11.[143,1459,125,1935]" captionTargetPageId="11" captionText="FIGURE 8. Oxynaspis celata (RUMF-ZC- 1557). A, cirrus I; B, simple type setae on cirrus I; C, cirrus II; D, simple type setae on cirrus II; E, simple type setae on cirrus IV; F, intermediate segment of posterior ramus of cirrus VI; G, caudal appendage; H, pedicel of penis. Scale bars in μm." httpUri="https://zenodo.org/record/281660/files/figure.png" pageId="9" pageNumber="30">Fig. 8</figureCitation>
G). Penis long, half length of cirrus VI, pedicel of penis without basi-dorsal point (
<figureCitation id="136E2A47FFD68A13C62225EE56D6F9C7" box="[354,420,1534,1558]" captionStart="FIGURE 8" captionStartId="11.[151,244,1933,1954]" captionTargetBox="[182,1400,161,1909]" captionTargetId="figure@11.[143,1459,125,1935]" captionTargetPageId="11" captionText="FIGURE 8. Oxynaspis celata (RUMF-ZC- 1557). A, cirrus I; B, simple type setae on cirrus I; C, cirrus II; D, simple type setae on cirrus II; E, simple type setae on cirrus IV; F, intermediate segment of posterior ramus of cirrus VI; G, caudal appendage; H, pedicel of penis. Scale bars in μm." httpUri="https://zenodo.org/record/281660/files/figure.png" pageId="9" pageNumber="30">Fig. 8</figureCitation>
H).
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD68A13C78726315571F98C" pageId="9" pageNumber="30" type="distribution">
<paragraph id="8BEA36C2FFD68A13C78726315571F98C" blockId="9.[151,1437,611,2019]" pageId="9" pageNumber="30">
<emphasis id="B921EAD0FFD68A13C78726315612F9EB" bold="true" box="[199,352,1569,1594]" pageId="9" pageNumber="30">Distribution.</emphasis>
Atlantic Ocean (Madeira); Indian Ocean (Bay of Bengal); South
<collectingCountry id="F3427652FFD68A13C310263153E1F9E8" box="[1104,1171,1569,1593]" name="China" pageId="9" pageNumber="30">China</collectingCountry>
Sea (Beibu Gulf);
<collectingCountry id="F3427652FFD68A13C22F263157B4F98C" name="Taiwan" pageId="9" pageNumber="30">Taiwan</collectingCountry>
Strait;
<collectingCountry id="F3427652FFD68A13C65726565628F98C" box="[279,346,1606,1629]" name="Japan" pageId="9" pageNumber="30">Japan</collectingCountry>
;
<collectingCountry id="F3427652FFD68A13C6262656568DF98C" box="[358,511,1605,1629]" name="New Zealand" pageId="9" pageNumber="30">New Zealand</collectingCountry>
,
</paragraph>
</subSubSection>
<subSubSection id="C34F6549FFD68A13C78726795543F832" pageId="9" pageNumber="30" type="discussion">
<paragraph id="8BEA36C2FFD68A13C78726795543F832" blockId="9.[151,1437,611,2019]" pageId="9" pageNumber="30">
<emphasis id="B921EAD0FFD68A13C78726795649F950" bold="true" box="[199,315,1641,1665]" pageId="9" pageNumber="30">Remarks.</emphasis>
<bibRefCitation id="EFC44B33FFD68A13C6032678569BF951" author="Darwin" box="[323,489,1640,1664]" pageId="9" pageNumber="30" refString="Darwin, C. (1852) A monograph on the sub-class Cirripedia with figures of all species. The Lepadidae: or pedunculated barnacles. Ray Society, London. 400 pp. (1851, published 1852)" type="book" year="1852">Darwin (1852)</bibRefCitation>
identified
<taxonomicName id="4C554D41FFD68A13C52526795454F951" box="[613,806,1640,1664]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD68A13C52526795454F951" box="[613,806,1640,1664]" italics="true" pageId="9" pageNumber="30">Oxynaspis celata</emphasis>
</taxonomicName>
from an antipatharian coral,
<taxonomicName id="4C554D41FFD68A13C330267852E7F951" box="[1136,1429,1640,1664]" class="Anthozoa" family="Aphanipathidae" genus="Aphanipathes" kingdom="Animalia" order="Antipatharia" pageId="9" pageNumber="30" phylum="Cnidaria" rank="species" species="woollastoni">
<emphasis id="B921EAD0FFD68A13C330267852E7F951" box="[1136,1429,1640,1664]" italics="true" pageId="9" pageNumber="30">Aphanipathes woollastoni</emphasis>
</taxonomicName>
, from Madeira, Atlantic Ocean (
<bibRefCitation id="EFC44B33FFD68A13C6B9269C55FAF975" author="Totton" box="[505,648,1676,1700]" pageId="9" pageNumber="30" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
). Subsequently, it has been widely recorded as four sub-species in the world’s oceans (
<taxonomicName id="4C554D41FFD68A13C61226A055A4F916" authority="Annandale, 1909" authorityName="Annandale" authorityYear="1909" box="[338,726,1711,1735]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD68A13C61226A0557BF916" box="[338,521,1711,1735]" italics="true" pageId="9" pageNumber="30">O. celata indica</emphasis>
<bibRefCitation id="EFC44B33FFD68A13C55226BF55A4F916" author="Annandale" box="[530,726,1711,1735]" pageId="9" pageNumber="30" refString="Annandale, N. (1909) An account of the Indian Cirripedia Pedunculata Part 1. Family Lepadidae (sensu stricto). Records of the Indian Museum, 2 (2), 61 - 137." type="journal article" year="1909">Annandale, 1909</bibRefCitation>
</taxonomicName>
; also see
<bibRefCitation id="EFC44B33FFD68A13C40626A054BDF916" author="Foster" box="[838,975,1711,1735]" pageId="9" pageNumber="30" refString="Foster, B. A. (1978) The marine fauna of New Zealand: barnacles (Cirripedia: Thoracica). New Zealand Oceaographic Institute Memoir, 69, 1 - 160." type="journal article" year="1978">Foster 1978</bibRefCitation>
;
<emphasis id="B921EAD0FFD68A13C49D26A053CDF916" box="[989,1215,1711,1736]" italics="true" pageId="9" pageNumber="30">O. c. novazelandica</emphasis>
<bibRefCitation id="EFC44B33FFD68A13C38826BF522AF916" author="Broch" box="[1224,1368,1711,1735]" pageId="9" pageNumber="30" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch, 1922</bibRefCitation>
;
<taxonomicName id="4C554D41FFD68A13C22426A056E1F93A" authority="Broch, 1922" authorityName="Broch" authorityYear="1922" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD68A13C22426A0578BF93A" italics="true" pageId="9" pageNumber="30">O. c. japonica</emphasis>
<bibRefCitation id="EFC44B33FFD68A13C64226C356E1F93A" author="Broch" box="[258,403,1747,1771]" pageId="9" pageNumber="30" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch, 1922</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C554D41FFD68A13C6E026C455CFF93A" authority="Totton 1940" authorityName="Totton" authorityYear="1940" box="[416,701,1747,1771]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="hirtae">
<emphasis id="B921EAD0FFD68A13C6E026C45557F93A" box="[416,549,1747,1771]" italics="true" pageId="9" pageNumber="30">O. c. hirtae</emphasis>
<bibRefCitation id="EFC44B33FFD68A13C56E26C455CFF93A" author="Totton" box="[558,701,1747,1771]" pageId="9" pageNumber="30" refString="Totton, A. K. (1940) New species of the cirripede genus Oxynaspis commensal with Antipatharia. The Annals and Magazine of Natural History, Series, 11, 6, 465 - 487." type="journal article" year="1940">Totton 1940</bibRefCitation>
</taxonomicName>
).
<bibRefCitation id="EFC44B33FFD68A13C59226C35406F93A" author="Zevina" box="[722,884,1747,1771]" pageId="9" pageNumber="30" refString="Zevina, G. B. (1982) Barnacles of the suborder Lepadomorpha (Cirripedia, Thoracica) of the world oceans. II. Opredeliteli Faune SSSR, 133, 1 - 221. (in Russian)." type="journal article" year="1982">Zevina (1982)</bibRefCitation>
revised the genus
<taxonomicName id="4C554D41FFD68A13C30E26C453B1F93A" box="[1102,1219,1748,1771]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD0FFD68A13C30E26C453B1F93A" box="[1102,1219,1748,1771]" italics="true" pageId="9" pageNumber="30">Oxynaspis</emphasis>
</taxonomicName>
and grouped
<taxonomicName id="4C554D41FFD68A13C22426C4578BF8DF" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD68A13C22426C4578BF8DF" italics="true" pageId="9" pageNumber="30">O. c. japonica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D41FFD68A13C66E26E756DDF8DF" box="[302,431,1782,1806]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD68A13C66E26E756DDF8DF" box="[302,431,1782,1806]" italics="true" pageId="9" pageNumber="30">O. c. indica</emphasis>
</taxonomicName>
as synonyms of
<taxonomicName id="4C554D41FFD68A13C52926E755BDF8DF" box="[617,719,1782,1806]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD68A13C52926E755BDF8DF" box="[617,719,1782,1806]" italics="true" pageId="9" pageNumber="30">O. celata</emphasis>
</taxonomicName>
, due to similarities in their external capitular morphology.
<bibRefCitation id="EFC44B33FFD68A13C21E26E7566BF8E0" author="Newman" pageId="9" pageNumber="30" refString="Newman, W. A. (1972) An Oxynaspid (Cirripedia, Thoracica) from the Eastern Pacific. Crustaceana, 23, 202 - 208" type="journal article" year="1972">Newman (1972)</bibRefCitation>
and
<bibRefCitation id="EFC44B33FFD68A13C610270A55B7F8E0" box="[336,709,1817,1841]" pageId="9" pageNumber="30" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article">Van Syoc and Dekelboum (2011)</bibRefCitation>
retained
<taxonomicName id="4C554D41FFD68A13C46D270A54BEF8E3" box="[813,972,1818,1842]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD68A13C46D270A54BEF8E3" box="[813,972,1818,1842]" italics="true" pageId="9" pageNumber="30">O. c. japonica</emphasis>
</taxonomicName>
,
<emphasis id="B921EAD0FFD68A13C49A270A53CBF8E3" box="[986,1209,1818,1842]" italics="true" pageId="9" pageNumber="30">O. c. novazelandica</emphasis>
,
<taxonomicName id="4C554D41FFD68A13C387270A5238F8E3" box="[1223,1354,1818,1842]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD68A13C387270A5238F8E3" box="[1223,1354,1818,1842]" italics="true" pageId="9" pageNumber="30">O. c. indica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D41FFD68A13C2C1270A5786F884" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="hirtae">
<emphasis id="B921EAD0FFD68A13C2C1270A5786F884" italics="true" pageId="9" pageNumber="30">O. c. hirtae</emphasis>
</taxonomicName>
as distinct sub-species of
<taxonomicName id="4C554D41FFD68A13C555272E5509F884" box="[533,635,1853,1877]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD68A13C555272E5509F884" box="[533,635,1853,1877]" italics="true" pageId="9" pageNumber="30">O. celata</emphasis>
</taxonomicName>
.
<taxonomicName id="4C554D41FFD68A13C5C9272E543AF884" box="[649,840,1853,1877]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD68A13C5C9272E543AF884" box="[649,840,1853,1877]" italics="true" pageId="9" pageNumber="30">Oxynaspis celata</emphasis>
</taxonomicName>
may be a cryptic species complex with high morphological variations in the world oceans and the taxonomy of this cryptic species group is unclear (see
<bibRefCitation id="EFC44B33FFD68A13C246277152FDF8A9" author="Foster" box="[1286,1423,1888,1912]" pageId="9" pageNumber="30" refString="Foster, B. A. (1978) The marine fauna of New Zealand: barnacles (Cirripedia: Thoracica). New Zealand Oceaographic Institute Memoir, 69, 1 - 160." type="journal article" year="1978">Foster 1978</bibRefCitation>
). Further studies should focus on the taxonomic status and population genetics of
<taxonomicName id="4C554D41FFD68A13C35F279553F5F84D" box="[1055,1159,1924,1948]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="9" pageNumber="30" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD68A13C35F279553F5F84D" box="[1055,1159,1924,1948]" italics="true" pageId="9" pageNumber="30">O. celata</emphasis>
</taxonomicName>
from different oceans in the world, using a combination of morphological and molecular approaches. In the present study we tentatively follow
<bibRefCitation id="EFC44B33FFD68A13C78827DC555EF832" box="[200,556,1995,2019]" pageId="9" pageNumber="30" refString="Van Syoc, R. &amp; Dekelboum, A. (2011) Oxynaspididae (Crustacea, Cirripedia): phylogenetic and evolutionary ecology, with description of three new genera and six new species. Zootaxa, 3103, 1 - 32." type="journal article">Van Syoc &amp; Dekelboum (2011)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2A664AFFD58A10C7D7278056CFF80F" httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="10" pageNumber="31" targetBox="[186,1402,156,1908]" targetPageId="10">
<paragraph id="8BEA36C2FFD58A10C7D7278056CFF80F" blockId="10.[151,1436,1936,2016]" pageId="10" pageNumber="31">
<emphasis id="B921EAD0FFD58A10C7D72780567FF874" bold="true" box="[151,269,1936,1957]" pageId="10" pageNumber="31">FIGURE 7.</emphasis>
<taxonomicName id="4C554D41FFD58A10C65327805536F874" authority="Darwin, 1852" authorityName="Darwin" authorityYear="1852" box="[275,580,1936,1957]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD58A10C653278056CAF874" box="[275,440,1936,1957]" italics="true" pageId="10" pageNumber="31">Oxynaspis celata</emphasis>
Darwin, 1852
</taxonomicName>
(RUMF-ZC-1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm.
</paragraph>
</caption>
<caption id="DF2A664AFFD48A11C7D7279D560FF80A" httpUri="https://zenodo.org/record/281660/files/figure.png" pageId="11" pageNumber="32" targetBox="[182,1400,161,1909]" targetPageId="11">
<paragraph id="8BEA36C2FFD48A11C7D7279D560FF80A" blockId="11.[151,1436,1933,2012]" pageId="11" pageNumber="32">
<emphasis id="B921EAD0FFD48A11C7D7279D567EF873" bold="true" box="[151,268,1933,1954]" pageId="11" pageNumber="32">FIGURE 8.</emphasis>
<taxonomicName id="4C554D41FFD48A11C652279D56C5F873" box="[274,439,1933,1954]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD48A11C652279D56C5F873" box="[274,439,1933,1954]" italics="true" pageId="11" pageNumber="32">Oxynaspis celata</emphasis>
</taxonomicName>
(RUMF-ZC-1557). A, cirrus I; B, simple type setae on cirrus I; C, cirrus II; D, simple type setae on cirrus II; E, simple type setae on cirrus IV; F, intermediate segment of posterior ramus of cirrus VI; G, caudal appendage; H, pedicel of penis. Scale bars in μm.
</paragraph>
</caption>
<subSubSection id="C34F6549FFD38A16C78720885219FD75" pageId="12" pageNumber="33" type="reference_group">
<paragraph id="8BEA36C2FFD38A16C78720885219FD75" blockId="12.[151,1436,151,676]" pageId="12" pageNumber="33">
<taxonomicName id="4C554D41FFD38A16C7872088552EFF7E" authority="Broch, 1922" authorityName="Broch" authorityYear="1922" box="[199,604,151,176]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD38A16C787208856B7FF61" box="[199,453,152,176]" italics="true" pageId="12" pageNumber="33">Oxynaspis c. japonica</emphasis>
<bibRefCitation id="EFC44B33FFD38A16C68E2087552EFF7E" author="Broch" box="[462,604,151,175]" pageId="12" pageNumber="33" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch, 1922</bibRefCitation>
</taxonomicName>
was collected from Nagasaki,
<collectingCountry id="F3427652FFD38A16C48320885374FF7E" box="[963,1030,152,175]" name="Japan" pageId="12" pageNumber="33">Japan</collectingCountry>
. The capitular morphology and the arthropodal characters (cirri and mouth parts) of the present specimen fit the descriptions of
<taxonomicName id="4C554D41FFD38A16C3B020AC5671FF26" authority="Darwin,1852" authorityName="Darwin" authorityYear="1852" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD38A16C3B020AC522EFF05" box="[1264,1372,188,212]" italics="true" pageId="12" pageNumber="33">O. celata</emphasis>
<bibRefCitation id="EFC44B33FFD38A16C22920AC5671FF26" author="Darwin" pageId="12" pageNumber="33" refString="Darwin, C. (1852) A monograph on the sub-class Cirripedia with figures of all species. The Lepadidae: or pedunculated barnacles. Ray Society, London. 400 pp. (1851, published 1852)" type="book" year="1852">Darwin,1852</bibRefCitation>
</taxonomicName>
. Comparing the present specimen with
<taxonomicName id="4C554D41FFD38A16C5FB20F0542BFF26" box="[699,857,224,247]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD38A16C5FB20F0542BFF26" box="[699,857,224,247]" italics="true" pageId="12" pageNumber="33">O. c. japonica</emphasis>
</taxonomicName>
collected in Nagasaki,
<collectingCountry id="F3427652FFD38A16C31D20F053EFFF26" box="[1117,1181,224,247]" name="Japan" pageId="12" pageNumber="33">Japan</collectingCountry>
(
<bibRefCitation id="EFC44B33FFD38A16C3EB20CF5246FF26" author="Broch" box="[1195,1332,223,247]" pageId="12" pageNumber="33" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch 1922</bibRefCitation>
), most of the morphology described for
<taxonomicName id="4C554D41FFD38A16C6B4211455E5FECA" box="[500,663,260,283]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD38A16C6B4211455E5FECA" box="[500,663,260,283]" italics="true" pageId="12" pageNumber="33">O. c. japonica</emphasis>
</taxonomicName>
is similar to the present specimen except for morphological differences exhibited by the labrum.
<bibRefCitation id="EFC44B33FFD38A16C6BA213755E0FEEE" author="Broch" box="[506,658,295,319]" pageId="12" pageNumber="33" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch (1922)</bibRefCitation>
described the labrum of
<taxonomicName id="4C554D41FFD38A16C4EE213853F2FEEE" box="[942,1152,295,319]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="japonica">
<emphasis id="B921EAD0FFD38A16C4EE213853F2FEEE" box="[942,1152,295,319]" italics="true" pageId="12" pageNumber="33">O. celata japonica</emphasis>
</taxonomicName>
as follows - “a deep and broad median furrow extends from between the palpi and its extreme top and makes the projecting end of the labrum appear a little cleft”. Such a deep and broad median burrow is absent from the labrum in the present specimen.
<bibRefCitation id="EFC44B33FFD38A16C79521825608FE7B" author="Darwin" box="[213,378,402,426]" pageId="12" pageNumber="33" refString="Darwin, C. (1852) A monograph on the sub-class Cirripedia with figures of all species. The Lepadidae: or pedunculated barnacles. Ray Society, London. 400 pp. (1851, published 1852)" type="book" year="1852">Darwin (1852)</bibRefCitation>
did not mentioned the existence of such a furrow in the labrum of
<taxonomicName id="4C554D41FFD38A16C323218353BAFE7B" box="[1123,1224,402,426]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD38A16C323218353BAFE7B" box="[1123,1224,402,426]" italics="true" pageId="12" pageNumber="33">O. celata</emphasis>
</taxonomicName>
. The present specimen differs from
<taxonomicName id="4C554D41FFD38A16C62121A75691FE1F" box="[353,483,438,462]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD38A16C62121A75691FE1F" box="[353,483,438,462]" italics="true" pageId="12" pageNumber="33">O. c. indica</emphasis>
</taxonomicName>
as the opercular plate margins are smooth whereas those of
<taxonomicName id="4C554D41FFD38A16C3C021A75270FE1F" box="[1152,1282,438,462]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="indica">
<emphasis id="B921EAD0FFD38A16C3C021A75270FE1F" box="[1152,1282,438,462]" italics="true" pageId="12" pageNumber="33">O. c. indica</emphasis>
</taxonomicName>
have serrated margins (Table 1). The present specimen also differs from
<taxonomicName id="4C554D41FFD38A16C47621CA54CBFE20" box="[822,953,473,497]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="celata" subSpecies="hirtae">
<emphasis id="B921EAD0FFD38A16C47621CA54CBFE20" box="[822,953,473,497]" italics="true" pageId="12" pageNumber="33">O. c. hirtae</emphasis>
</taxonomicName>
as the latter has an additional pair of filamentary appendages at the base of cirrus I (Table 1).
<taxonomicName id="4C554D41FFD38A16C5BC21EE54CFFDC4" box="[764,957,509,533]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD38A16C5BC21EE54CFFDC4" box="[764,957,509,533]" italics="true" pageId="12" pageNumber="33">Oxynaspis celata</emphasis>
</taxonomicName>
collected in the present study also differs from
<emphasis id="B921EAD0FFD38A16C797223256CFFDE8" box="[215,445,545,569]" italics="true" pageId="12" pageNumber="33">O. c. novazelandica</emphasis>
, as the maxillule of
<emphasis id="B921EAD0FFD38A16C5F2223254E5FDE8" box="[690,919,545,569]" italics="true" pageId="12" pageNumber="33">O. c. novazelandica</emphasis>
is strongly notched.
<bibRefCitation id="EFC44B33FFD38A16C3CE223152EEFDE8" author="Broch" box="[1166,1436,545,569]" pageId="12" pageNumber="33" refString="Broch, H. (1922) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916, X. Studies on Pacific Cirripedes. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening I Kobenhavn, 73, 215 - 358." type="journal article" year="1922">Broch (1922: 278–279)</bibRefCitation>
described the maxillule of
<emphasis id="B921EAD0FFD38A16C681225555EDFD8C" box="[449,671,581,605]" italics="true" pageId="12" pageNumber="33">O. c. novazelandica</emphasis>
as having “only three spines above the excavation (= notch) and the excavation is very broad, occupying almost half of the cutting edge….”. In the present study, the maxillule of the
<taxonomicName id="4C554D41FFD38A16C7D7229D578FFD75" box="[151,253,652,676]" class="Maxillopoda" family="Oxynaspididae" genus="Oxynaspis" kingdom="Animalia" order="Pedunculata" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="B921EAD0FFD38A16C7D7229D578FFD75" box="[151,253,652,676]" italics="true" pageId="12" pageNumber="33">O. celata</emphasis>
</taxonomicName>
is slightly notched (
<figureCitation id="136E2A47FFD38A16C6A1229C5556FD75" box="[481,548,652,676]" captionStart="FIGURE 7" captionStartId="10.[151,244,1936,1957]" captionTargetBox="[186,1402,156,1908]" captionTargetId="figure@10.[146,1462,119,1929]" captionTargetPageId="10" captionText="FIGURE 7. Oxynaspis celata Darwin, 1852 (RUMF-ZC- 1557). A, maxilla; B, magnified view of setae of maxilla; C, maxillule; D, cutting margin of maxillule; E, mandible; F, lower margin and inferior angle of mandible; G, mandibular palp; H, setae on the of mandibular palp. Scale bars in μm." httpUri="https://zenodo.org/record/281659/files/figure.png" pageId="12" pageNumber="33">Fig. 7</figureCitation>
C), differing from
<emphasis id="B921EAD0FFD38A16C5B3229D54A3FD75" box="[755,977,652,676]" italics="true" pageId="12" pageNumber="33">O. c. novazelandica</emphasis>
which has a broad maxillular notch.
</paragraph>
</subSubSection>
</treatment>
</document>
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