231 lines
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231 lines
33 KiB
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<document id="AA27B8E9BECCE10BE079840DCC08D3C5" ID-DOI="10.11646/zootaxa.5501.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13628165" ID-ZooBank="6A91B5D0-0251-4F5C-AA32-D70CAAE55F90" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725278913235" checkinUser="plazi" docAuthor="Telnov, Dmitry" docDate="2024" docId="03C887C4361E34180D84D529B834BD86" docLanguage="en" docName="zootaxa.5501.3.1.pdf" docOrigin="Zootaxa 5501 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5501.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copobaenus erratus Telnov, 2024, sp. nov." docType="treatment" docUuid="E1B53636-2013-4E43-A1DA-94AB97864C89" docUuidSource="ZooBank" docVersion="2" lastPageNumber="410" masterDocId="FFF1FFBC361934110D13D70BBC3AB943" masterDocTitle="On the taxonomy of some South American and Australo-Pacific Anthicidae Latreille, 1819 (Coleoptera) with new subfamily and tribe-rank synonymies" masterLastPageNumber="424" masterPageNumber="401" pageNumber="408" updateTime="1725279392153" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="009A99E87D48BC854E4AC769835D2C9E">On the taxonomy of some South American and Australo-Pacific Anthicidae Latreille, 1819 (Coleoptera) with new subfamily and tribe-rank synonymies</mods:title>
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<mods:name id="99F7C9838A2B9F940CD126580E9A982D" type="personal">
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<mods:namePart id="AC68B43F60EC90085A1C4C3098233F62">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="E3670A2FE85311F50D034DAC3EAF185D">Department of Life Sciences, Natural History Museum, SW 7 5 BD London, United Kingdom & Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401 Daugavpils, Latvia & Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004 Rīga, Latvia</mods:affiliation>
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<mods:typeOfResource id="CD8E1CDB58CA49AE56654C4475A9F3DB">text</mods:typeOfResource>
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<mods:title id="4F8EB6F649B7524F40AC57A0093B296A">Zootaxa</mods:title>
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<mods:part id="8B7FDA54DC44CE84C9172A4AFDD6DFED">
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<mods:date id="44D7919559D9BC44300C0967078BA614">2024</mods:date>
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<mods:number id="450A98BD759A44F95CEC59B2AE25978E">2024-09-02</mods:number>
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<mods:number id="2FCFB5C87AA3185960ADBA3AC397CEF3">5501</mods:number>
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<mods:url id="CE2202F00023677F1ACE2045064C6EF1">http://dx.doi.org/10.11646/zootaxa.5501.3.1</mods:url>
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<mods:classification id="C815E79DA94D7FB0C2884951809F7AA1">journal article</mods:classification>
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<mods:identifier id="9D90146302AC1417E7D57A9F424652F4" type="Zenodo-Dep">13628165</mods:identifier>
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<treatment id="03C887C4361E34180D84D529B834BD86" ID-DOI="http://doi.org/10.5281/zenodo.13628161" ID-Zenodo-Dep="13628161" LSID="urn:lsid:zoobank.org:act:E1B53636-2013-4E43-A1DA-94AB97864C89" httpUri="http://treatment.plazi.org/id/03C887C4361E34180D84D529B834BD86" lastPageId="9" lastPageNumber="410" pageId="7" pageNumber="408">
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<subSubSection id="C37B6559361E34160D84D529BFF8BB22" pageId="7" pageNumber="408" type="nomenclature">
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<paragraph id="8BDE36D2361E34160D84D529BE6BBB7F" blockId="7.[151,962,546,609]" box="[151,593,546,573]" pageId="7" pageNumber="408">
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<heading id="D09681BE361E34160D84D529BE6BBB7F" bold="true" box="[151,593,546,573]" fontSize="11" level="1" pageId="7" pageNumber="408" reason="1">
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<emphasis id="B915EAC0361E34160D84D529BE6BBB7F" bold="true" box="[151,593,546,573]" pageId="7" pageNumber="408">
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<taxonomicName id="4C614D51361E34160D84D529BDB0BB7E" authorityName="Telnov" authorityYear="2024" box="[151,394,546,573]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="erratus" status="sp. nov.">
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<emphasis id="B915EAC0361E34160D84D529BDB0BB7E" bold="true" box="[151,394,546,573]" italics="true" pageId="7" pageNumber="408">Copobaenus erratus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22657BB361E34160C81D528BDCABB7E" box="[402,496,547,573]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="135A2A57361E34160F13D529BE72BB7F" box="[512,584,546,572]" captionStart="FIGURE 3" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[236,1353,190,1873]" captionTargetId="figure-40@8.[225,1363,181,1884]" captionTargetPageId="8" captionText="FIGURE 3. Copobaenus erratus sp. nov., holotype ♂. A—Habitus, dorsal view; B—Forebody, dorsal view; C—Tergite VII, dorsal view; D—Morphological sternite VII, ventral view; E—Morphological sternites VIII and IX; F—Aedeagus, ventral view; G—ditto, lateral view [not to scale]. Image courtesy A. Degiovanni, Bubano, Italy published in Telnov & Degiovanni (2021)." figureDoi="http://doi.org/10.5281/zenodo.13628173" httpUri="https://zenodo.org/record/13628173/files/figure.png" pageId="7" pageNumber="408">Fig. 3</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDE36D2361E34160D84D54DBFF8BB22" blockId="7.[151,962,546,609]" box="[151,962,582,609]" pageId="7" pageNumber="408">
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<uri id="FFF03AD0361E34160D84D54DBFF8BB22" box="[151,962,582,609]" pageId="7" pageNumber="408">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC70C07361E34160CB8D54DBFF8BB22" box="[427,962,582,609]" pageId="7" pageNumber="408">E1B53636-2013-4E43-A1DA-94AB97864C89</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6559361E34160D84D583B978BADD" pageId="7" pageNumber="408" type="description">
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<paragraph id="8BDE36D2361E34160D84D583B978BADD" blockId="7.[151,1437,648,2043]" pageId="7" pageNumber="408">
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Telnov & Degiovanni (2021: 122, 145) erroneously interpreted the only known male of this new species as
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<taxonomicName id="4C614D51361E3416082ED583B9A6BBE1" authorityName="Fairm & Germ Det. J. E. Barriga Tunon" authorityYear="2008" box="[1341,1436,648,674]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B915EAC0361E3416082ED583B9A6BBE1" box="[1341,1436,648,674]" italics="true" pageId="7" pageNumber="408">C. tristis</emphasis>
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</taxonomicName>
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and figured the specimen (Telnov & Degiovanni (2021), figs 46–47, 50–53; also see above). In fact, the aedeagus of
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<taxonomicName id="4C614D51361E34160DA4D5DBBD20BBA9" authorityName="Fairm & Germ Det. J. E. Barriga Tunon" authorityYear="2008" box="[183,282,720,746]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B915EAC0361E34160DA4D5DBBD20BBA9" box="[183,282,720,746]" italics="true" pageId="7" pageNumber="408">C. tristis</emphasis>
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</taxonomicName>
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, as described and figured by
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<bibRefCitation id="EFF04B23361E34160F7ED5DBBFAEBBA9" author="Abdullah, M." box="[621,916,720,746]" pageId="7" pageNumber="408" pagination="323 - 366" refId="ref11957" refString="Abdullah, M. (1969) The natural classification of the family Anthicidae with some ecological and ethological observations (Coleoptera). Deutsche Entomologische Zeitschrift, 16 (iv / v), 323 - 366. https: // doi. org / 10.1002 / mmnd. 4810160404" type="journal article" year="1969">Abdullah (1969: 137–138</bibRefCitation>
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, figs 12–16) and proven herein is completely different and resembles that of
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<taxonomicName id="4C614D51361E34160F16D5FFB82DBA4D" authority="(Telnov & Degiovanni, 2021)" authorityName="Telnov" authorityYear="2024" baseAuthorityName="Telnov & Degiovanni" baseAuthorityYear="2021" box="[517,1047,756,782]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="inexpectatus">
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<emphasis id="B915EAC0361E34160F16D5FFBE8FBA4D" box="[517,693,756,782]" italics="true" pageId="7" pageNumber="408">C. inexpectatus</emphasis>
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(Telnov & Degiovanni, 2021)
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</taxonomicName>
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(see Telnov & Degiovanni 2021, figs 101–102). Additionally, the dorsal colouration of
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<taxonomicName id="4C614D51361E34160FEBD413BF63BA71" authorityName="Fairm & Germ Det. J. E. Barriga Tunon" authorityYear="2008" box="[760,857,792,818]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B915EAC0361E34160FEBD413BF63BA71" box="[760,857,792,818]" italics="true" pageId="7" pageNumber="408">C. tristis</emphasis>
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</taxonomicName>
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, in particular that of elytra, is different (see images in Telnov & Degiovanni (2021), figs 48–49). Therefore, all but
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<specimenCount id="9D67FD5B361E34160E6CD436B826BA15" box="[895,1052,828,855]" count="1" pageId="7" pageNumber="408" type="generic">one specimen</specimenCount>
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listed as ‘
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<taxonomicName id="4C614D51361E3416098BD437B8C6BA15" authorityName="Fairm & Germ Det. J. E. Barriga Tunon" authorityYear="2008" box="[1176,1276,828,854]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B915EAC0361E3416098BD437B8C6BA15" box="[1176,1276,828,854]" italics="true" pageId="7" pageNumber="408">C. tristis</emphasis>
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</taxonomicName>
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’ in Telnov & Degiovanni 2021 from Nahuelbuta National park in
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<collectingCountry id="F3767642361E34160FF4D46BBF1EBA39" box="[743,804,864,890]" name="Chile" pageId="7" pageNumber="408">Chile</collectingCountry>
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are
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<specimenCount id="9D67FD5B361E34160E46D46BBFC8BA39" box="[853,1010,864,890]" count="6" pageId="7" pageNumber="408" type="generic">six specimens</specimenCount>
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of
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<taxonomicName id="4C614D51361E3416090BD46BB840BA39" authorityName="Fairm & Germ Det. J. E. Barriga Tunon" authorityYear="2008" box="[1048,1146,864,890]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B915EAC0361E3416090BD46BB840BA39" box="[1048,1146,864,890]" italics="true" pageId="7" pageNumber="408">C. tristis</emphasis>
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</taxonomicName>
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, and the single male with blackish elytra (as figured by the authors, figs 46–47, 50–54), belongs to the new species described herein.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6559361E34160DD4D4A3BE84BD11" pageId="7" pageNumber="408" type="materials_examined">
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<paragraph id="8BDE36D2361E34160DD4D4A3BE84BD11" blockId="7.[151,1437,648,2043]" pageId="7" pageNumber="408">
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<emphasis id="B915EAC0361E34160DD4D4A3BDC7BA81" bold="true" box="[199,509,936,962]" pageId="7" pageNumber="408">Type material designated.</emphasis>
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<typeStatus id="54DA8870361E34160F17D4A3BE56BA81" box="[516,620,936,962]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D67FD5B361E34160F67D4A2BEB3BA80" box="[628,649,937,963]" count="1" pageId="7" pageNumber="408" type="male">♂</specimenCount>
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<collectionCode id="ED70AE17361E34160F82D4A2BED9BA80" box="[657,739,937,963]" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="7" pageNumber="408">MZUF</collectionCode>
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:
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<collectingCountry id="F3767642361E34160FFDD4A3BF79BA81" box="[750,835,936,962]" name="Chile" pageId="7" pageNumber="408">CHILE</collectingCountry>
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, prov.
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<collectingRegion id="49A5F830361E34160E9DD4A3BFD9BA81" box="[910,995,936,962]" country="Chile" name="Araucania" pageId="7" pageNumber="408">Arauco</collectingRegion>
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, P.N. Nahuelbuta, Pichinahual,
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<quantity id="4C999B37361E34160854D4A3B9A2BA81" box="[1351,1432,936,962]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="408" unit="m" value="1200.0">1200m</quantity>
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,
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<date id="FFDF1012361E34160D84D4C7BD0BBAA5" box="[151,305,972,998]" pageId="7" pageNumber="408" value="2004-11-22">22-Nov-2004</date>
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, leg. J.E. Barriga T., fogging, s/
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<taxonomicName id="4C614D51361E34160FB0D4C6BF16BAA5" box="[675,812,972,999]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="408" phylum="Tracheophyta" rank="species" species="dombeyi">N. dombeyi</taxonomicName>
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,
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<quantity id="4C999B37361E34160E2BD4C7BFABBAA4" box="[824,913,972,999]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="408" unit="m" value="1200.0">1200 m</quantity>
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,
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<geoCoordinate id="EE555015361E34160E8DD4C7BFC3BAA5" box="[926,1017,972,998]" degrees="37" direction="south" minutes="47" orientation="latitude" pageId="7" pageNumber="408" precision="925" value="-37.783333">37°47’S</geoCoordinate>
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<geoCoordinate id="EE555015361E34160911D4C7B854BAA4" box="[1026,1134,972,999]" degrees="73" direction="west" minutes="00" orientation="longitude" pageId="7" pageNumber="408" precision="925" value="-73.0">73°00’ W</geoCoordinate>
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[printed] // Coleccion J.E. BARRIGA
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<collectingCountry id="F3767642361E34160C0DD4FBBD4ABD49" box="[286,368,1008,1034]" name="Chile" pageId="7" pageNumber="408">CHILE</collectingCountry>
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137484 [printed] //
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<taxonomicName id="4C614D51361E34160F71D4FBBFDEBD49" authority="Fairm & Germ" authorityName="Fairmaire & Germain" authorityYear="1863" box="[610,996,1008,1034]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tristis">Copobaenus tristis Fairm& Germ</taxonomicName>
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[handwritten] Det.J.E. Barriga Tunon 2008 [printed] //
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<taxonomicName id="4C614D51361E34160C45D31FBF1ABD6D" authority="Fairmaire & Germain" authorityName="Fairmaire & Germain" authorityYear="1863" box="[342,800,1044,1070]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tristis">Copobaenus tristis Fairmaire & Germain</taxonomicName>
|
|||
|
|
[printed] // Coll. ANGELINI Num.
|
|||
|
|
<date id="FFDF1012361E341609A5D31EB916BD6C" box="[1206,1324,1044,1071]" pageId="7" pageNumber="408" value="2957-05">mag. 2957</date>
|
|||
|
|
[printed]. Antennomeres 10–11 of the left antenna missing.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B6559361E34160DD4D357B953BD35" box="[199,1385,1116,1143]" pageId="7" pageNumber="408" type="etymology">
|
|||
|
|
<paragraph id="8BDE36D2361E34160DD4D357B953BD35" blockId="7.[151,1437,648,2043]" box="[199,1385,1116,1143]" pageId="7" pageNumber="408">
|
|||
|
|
<emphasis id="B915EAC0361E34160DD4D357BD9EBD35" bold="true" box="[199,420,1116,1142]" pageId="7" pageNumber="408">Derivatio nominis.</emphasis>
|
|||
|
|
From the Latin “errātio” (delusion, mistake) to point out a previous misidentification.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B6559361E34180DD4D38BBE9BBD52" lastPageId="9" lastPageNumber="410" pageId="7" pageNumber="408" type="description">
|
|||
|
|
<paragraph id="8BDE36D2361E34160DD4D38BBF4DBDA0" blockId="7.[151,1437,648,2043]" pageId="7" pageNumber="408">
|
|||
|
|
<emphasis id="B915EAC0361E34160DD4D38BBD44BDD9" bold="true" box="[199,382,1152,1178]" pageId="7" pageNumber="408">Measurements.</emphasis>
|
|||
|
|
<typeStatus id="54DA8870361E34160C97D38BBDD5BDD9" box="[388,495,1152,1178]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
|
|||
|
|
, total body length
|
|||
|
|
<quantity id="4C999B37361E34160FADD38BBF2DBDD8" box="[702,791,1152,1179]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="7" pageNumber="408" unit="mm" value="4.1">4.1 mm</quantity>
|
|||
|
|
; head length excluding cranial ‘neck’
|
|||
|
|
<quantity id="4C999B37361E341609ADD38BB92CBDD8" box="[1214,1302,1152,1179]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="408" unit="mm" value="0.7">0.7 mm</quantity>
|
|||
|
|
, head width across eyes
|
|||
|
|
<quantity id="4C999B37361E34160C0FD3AFBDBBBDFC" box="[284,385,1188,1215]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="408" unit="mm" value="0.75">0.75 mm</quantity>
|
|||
|
|
, pronotal length
|
|||
|
|
<quantity id="4C999B37361E34160F2FD3AFBEAEBDFC" box="[572,660,1188,1215]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="408" unit="mm" value="0.7">0.7 mm</quantity>
|
|||
|
|
, maximum pronotal width
|
|||
|
|
<quantity id="4C999B37361E34160ED2D3AFB822BDFC" box="[961,1048,1188,1215]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="408" unit="mm" value="0.8">0.8 mm</quantity>
|
|||
|
|
, minimum pronotal width
|
|||
|
|
<quantity id="4C999B37361E34160853D3AFB9A2BDFC" box="[1344,1432,1188,1215]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="408" unit="mm" value="0.6">0.6 mm</quantity>
|
|||
|
|
, elytral length
|
|||
|
|
<quantity id="4C999B37361E34160C20D3C3BDB1BDA0" box="[307,395,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="408" unit="mm" value="2.7">2.7 mm</quantity>
|
|||
|
|
, combined maximum elytral width
|
|||
|
|
<quantity id="4C999B37361E34160E09D3C3BF49BDA0" box="[794,883,1224,1251]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="408" unit="mm" value="1.1">1.1 mm</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDE36D2361E34180DD4D3E7BE45BAAE" blockId="7.[151,1437,648,2043]" lastBlockId="9.[151,1437,151,1221]" lastPageId="9" lastPageNumber="410" pageId="7" pageNumber="408">
|
|||
|
|
<emphasis id="B915EAC0361E34160DD4D3E7BD63BC45" bold="true" box="[199,345,1260,1286]" pageId="7" pageNumber="408">Description.</emphasis>
|
|||
|
|
<typeStatus id="54DA8870361E34160C77D3E7BDF6BC45" box="[356,460,1260,1286]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
|
|||
|
|
male (
|
|||
|
|
<figureCitation id="135A2A57361E34160F30D3E7BE56BC45" box="[547,620,1260,1286]" captionStart="FIGURE 3" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[236,1353,190,1873]" captionTargetId="figure-40@8.[225,1363,181,1884]" captionTargetPageId="8" captionText="FIGURE 3. Copobaenus erratus sp. nov., holotype ♂. A—Habitus, dorsal view; B—Forebody, dorsal view; C—Tergite VII, dorsal view; D—Morphological sternite VII, ventral view; E—Morphological sternites VIII and IX; F—Aedeagus, ventral view; G—ditto, lateral view [not to scale]. Image courtesy A. Degiovanni, Bubano, Italy published in Telnov & Degiovanni (2021)." figureDoi="http://doi.org/10.5281/zenodo.13628173" httpUri="https://zenodo.org/record/13628173/files/figure.png" pageId="7" pageNumber="408">Fig. 3</figureCitation>
|
|||
|
|
). Dorsal head yellowish rufous in anterior, rufous-brown in posterior half, pronotum yellowish rufous, elytral apex narrowly yellow. Basal antennomere yellowish rufous, remaining antennomeres brown. Maxillary palpomeres 2–4 pale brown, terminal palpomere contrastingly pale yellow at apical margin. Legs including trochanters pale yellow, tarsomeres distally darker. Ventral head and prothorax yellowish rufous, pterothorax and abdomen brown. Male tergite and morphological sternite VII yellowish brown. Head dorsally somewhat flattened, glossy, with moderate, laterally strongly prominent, slightly elliptical compound eyes. Interfacetal setae not observed. Labrum subtruncate, clypeus shallowly emarginate at anterior margin. Frontoclypeal suture present, slightly arched (concave towards clypeus). Tempus slightly converging towards head base, about 0.7–0.8× as long as dorsal eye length. Posterior temporal angles broadly obtuse, head base truncate to inconspicuously emarginate. Frons very broad, about 2.2× as wide as dorsal eye length. Head dorsal punctures minute, variably sparse; intervening spaces glossy and glabrous, generally much wider than punctures. Head dorsal setation yellowish, variably long, rather sparse, suberect. Few much longer erect to suberect tactile setae on dorsal head, in particular, on tempora. Male antenna extending nearly to median third of elytra when directed posteriad, rather densely covered by suberect setae and each of antennomeres 1–9 distally with several long erect tactile setae. Basal antennomere slightly thickened, about 1.7–1.8× as long as antennomere two. Antennomere three about 1.5× as long as antennomere two, about 0.8–0.9× as long as antennomere four. Antennomeres 6–10 about same length as antennomere four; antennomere five about same long as antennomere three, slightly shorter than antennomeres 4, 6–10. Terminal antennomere slightly fusiform, hardly asymmetrical (one of margins indistinctly sinuous), apically rounded, about 1.6× as long as penultimate antennomere. Maxillary palpomeres 2–3 subtriangular, mesal margin angulate-lobate. Terminal maxillary palpomere securiform. Terminal labial palpomere subellipsoidal, flattened on inner, rounded on outer margin. Cranial ‘neck’ nearly half head-width across compound eyes, densely and coarsely punctate, subopaque. Pronotum dorsally somewhat flattened, slightly glossy, slightly wider than head, broadly rounded to subtruncate anteriorly, truncate at base, strongly rounded laterally and moderately deeply constricted thereafter prebasally. Transverse constriction not continuing dorsally across pronotal disc, dorsally and laterally with same large and coarse punctures as pronotal disc. Poorly defined longitudinal sulcus broad and shallow, extending almost length of pronotal disc (
|
|||
|
|
<figureCitation id="135A2A57361034180C49D608BDEDB85E" box="[346,471,259,285]" captionStart="FIGURE 3" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[236,1353,190,1873]" captionTargetId="figure-40@8.[225,1363,181,1884]" captionTargetPageId="8" captionText="FIGURE 3. Copobaenus erratus sp. nov., holotype ♂. A—Habitus, dorsal view; B—Forebody, dorsal view; C—Tergite VII, dorsal view; D—Morphological sternite VII, ventral view; E—Morphological sternites VIII and IX; F—Aedeagus, ventral view; G—ditto, lateral view [not to scale]. Image courtesy A. Degiovanni, Bubano, Italy published in Telnov & Degiovanni (2021)." figureDoi="http://doi.org/10.5281/zenodo.13628173" httpUri="https://zenodo.org/record/13628173/files/figure.png" pageId="9" pageNumber="410">Fig. 3A‒B</figureCitation>
|
|||
|
|
). Anterior rim marginal, vaguely indicated dorsally, very narrow ventrally. Antebasal sulcus vaguely indicated dorsally, moderately broad laterally in constriction area. Pronotal disc with a shallow median longitudinal furrow, surrounded area with poorly distinct tubercles. Punctures on pronotal constriction comparatively larger and denser than those on head, some punctures appear on dorsal tubercles. Intervening spaces generally as wide as to wider than punctures, glabrous. Pronotal setae yellowish, appressed to suberect, moderately dense, several much longer erect tactile setae on lateral margins and base. Scutellar shield small, apically rounded, corrugate punctured and setose. Elytra elongate, somewhat flattened dorsally, slightly glossy. Humerus broadly rounded. Postbasal transverse impression not indicated. Elytral apices broadly rounded separately. Elytral punctures large, dense, and coarse specifically on anterior two thirds, becoming smaller on apical third. Intervening spaces glossy, narrower than punctures and somewhat elevated on anterior two thirds, about as wide as punctures on apical third. Elytral setae yellowish, suberect, moderately long and dense, directed generally posteriad. Much longer erect tactile setae scattered across elytral disc. Sutural stria not indicated. Epipleuron narrow, punctate, and setose, developed in about three fourths of elytral length. Metathoracic wings fully developed (functional). Legs moderately long, rather densely setose. Femora not clavate. Tibia without modifications, about as long as corresponding femora. Terminal tibial spurs paired, nearly equally long, straight, apically acute angulate, laterally smooth. Penultimate tarsomere shortened, bilobate. Basal metatarsomere slightly shorter than combined length of remaining metatarsomeres. Pretarsal claws simple, basally indistinctly subdentate. Male tergite VII broadly rounded at posterior margin (
|
|||
|
|
<figureCitation id="135A2A57361034180C96D46CBDE5BAC2" box="[389,479,871,897]" captionStart="FIGURE 3" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[236,1353,190,1873]" captionTargetId="figure-40@8.[225,1363,181,1884]" captionTargetPageId="8" captionText="FIGURE 3. Copobaenus erratus sp. nov., holotype ♂. A—Habitus, dorsal view; B—Forebody, dorsal view; C—Tergite VII, dorsal view; D—Morphological sternite VII, ventral view; E—Morphological sternites VIII and IX; F—Aedeagus, ventral view; G—ditto, lateral view [not to scale]. Image courtesy A. Degiovanni, Bubano, Italy published in Telnov & Degiovanni (2021)." figureDoi="http://doi.org/10.5281/zenodo.13628173" httpUri="https://zenodo.org/record/13628173/files/figure.png" pageId="9" pageNumber="410">Fig. 3C</figureCitation>
|
|||
|
|
), male morphological sternite VII subtruncate to shallowly emarginate medially at posterior margin (
|
|||
|
|
<figureCitation id="135A2A57361034180C7BD480BDFDBAE6" box="[360,455,907,933]" captionStart="FIGURE 3" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[236,1353,190,1873]" captionTargetId="figure-40@8.[225,1363,181,1884]" captionTargetPageId="8" captionText="FIGURE 3. Copobaenus erratus sp. nov., holotype ♂. A—Habitus, dorsal view; B—Forebody, dorsal view; C—Tergite VII, dorsal view; D—Morphological sternite VII, ventral view; E—Morphological sternites VIII and IX; F—Aedeagus, ventral view; G—ditto, lateral view [not to scale]. Image courtesy A. Degiovanni, Bubano, Italy published in Telnov & Degiovanni (2021)." figureDoi="http://doi.org/10.5281/zenodo.13628173" httpUri="https://zenodo.org/record/13628173/files/figure.png" pageId="9" pageNumber="410">Fig. 3D</figureCitation>
|
|||
|
|
). Male tergite and morphological sternite VIII broadly rounded at posterior margin. Morphological sternite IX Y-shaped, basal stalk long and arms short (
|
|||
|
|
<figureCitation id="135A2A57361034180E8AD4A4BFCABA8A" box="[921,1008,943,969]" captionStart="FIGURE 3" captionStartId="8.[151,250,1908,1932]" captionTargetBox="[236,1353,190,1873]" captionTargetId="figure-40@8.[225,1363,181,1884]" captionTargetPageId="8" captionText="FIGURE 3. Copobaenus erratus sp. nov., holotype ♂. A—Habitus, dorsal view; B—Forebody, dorsal view; C—Tergite VII, dorsal view; D—Morphological sternite VII, ventral view; E—Morphological sternites VIII and IX; F—Aedeagus, ventral view; G—ditto, lateral view [not to scale]. Image courtesy A. Degiovanni, Bubano, Italy published in Telnov & Degiovanni (2021)." figureDoi="http://doi.org/10.5281/zenodo.13628173" httpUri="https://zenodo.org/record/13628173/files/figure.png" pageId="9" pageNumber="410">Fig. 3E</figureCitation>
|
|||
|
|
). Aedeagus as in fig. 3F‒G, penis free, longer than tegmen, phallobase not present.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF1E665A361134190D84D07FBCD8BEBA" ID-DOI="http://doi.org/10.5281/zenodo.13628173" ID-Zenodo-Dep="13628173" httpUri="https://zenodo.org/record/13628173/files/figure.png" pageId="8" pageNumber="409" startId="8.[151,250,1908,1932]" targetBox="[236,1353,190,1873]" targetPageId="8" targetType="figure">
|
|||
|
|
<paragraph id="8BDE36D2361134190D84D07FBCD8BEBA" blockId="8.[151,1436,1908,2041]" pageId="8" pageNumber="409">
|
|||
|
|
<emphasis id="B915EAC0361134190D84D07FBD2EBECF" bold="true" box="[151,276,1908,1932]" pageId="8" pageNumber="409">FIGURE 3.</emphasis>
|
|||
|
|
<taxonomicName id="4C614D51361134190C08D07FBDD1BECF" authorityName="Telnov" authorityYear="2024" box="[283,491,1908,1932]" class="Insecta" family="Anthicidae" genus="Copobaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="erratus" status="sp. nov.">
|
|||
|
|
<emphasis id="B915EAC0361134190C08D07FBDD1BECF" box="[283,491,1908,1932]" italics="true" pageId="8" pageNumber="409">Copobaenus erratus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B915EAC0361134190CE1D07EBE7EBECE" bold="true" box="[498,580,1909,1933]" pageId="8" pageNumber="409">
|
|||
|
|
<taxonomicNameLabel id="A22657BB361134190CE1D07EBE7EBECE" box="[498,580,1909,1933]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, holotype ♂. A—Habitus, dorsal view; B—Forebody, dorsal view; C—Tergite VII, dorsal view; D—Morphological sternite VII, ventral view; E—Morphological sternites VIII and IX; F—Aedeagus, ventral view; G—ditto, lateral view [not to scale]. Image courtesy A. Degiovanni, Bubano, Italy published in Telnov & Degiovanni (2021).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BDE36D2361034180DD4D4FDBE9BBD52" blockId="9.[151,1437,151,1221]" box="[199,673,1014,1041]" pageId="9" pageNumber="410">
|
|||
|
|
<emphasis id="B915EAC0361034180DD4D4FDBD8DBD53" bold="true" box="[199,439,1014,1040]" pageId="9" pageNumber="410">Sexual dimorphism.</emphasis>
|
|||
|
|
Female is unknown.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B6559361034180DD4D311BFFABD1A" pageId="9" pageNumber="410" type="diagnosis">
|
|||
|
|
<paragraph id="8BDE36D2361034180DD4D311BFFABD1A" blockId="9.[151,1437,151,1221]" pageId="9" pageNumber="410">
|
|||
|
|
<emphasis id="B915EAC0361034180DD4D311BDEABD77" bold="true" box="[199,464,1050,1076]" pageId="9" pageNumber="410">Differential diagnosis.</emphasis>
|
|||
|
|
This species is different from its congeners primarily in the contrastingly bicoloured dorsum with non-metallic elytra and the specifically long, narrow penis.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B6559361034180DD4D368BF04BDE2" pageId="9" pageNumber="410" type="biology_ecology">
|
|||
|
|
<paragraph id="8BDE36D2361034180DD4D368BF04BDE2" blockId="9.[151,1437,151,1221]" pageId="9" pageNumber="410">
|
|||
|
|
<emphasis id="B915EAC0361034180DD4D368BD16BD3E" bold="true" box="[199,300,1123,1149]" pageId="9" pageNumber="410">Ecology.</emphasis>
|
|||
|
|
Sampled from the canopy of
|
|||
|
|
<taxonomicName id="4C614D51361034180F62D368BF6DBD3E" box="[625,855,1123,1149]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="410" phylum="Tracheophyta" rank="species" species="dombeyi">
|
|||
|
|
<emphasis id="B915EAC0361034180F62D368BF6DBD3E" box="[625,855,1123,1149]" italics="true" pageId="9" pageNumber="410">Nothofagus dombeyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<taxonomicName id="4C614D51361034180E77D368B828BD3E" authorityName="Kuprianova" authorityYear="1962" box="[868,1042,1123,1149]" class="Magnoliopsida" family="Nothofagaceae" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="410" phylum="Tracheophyta" rank="family">Nothofagaceae</taxonomicName>
|
|||
|
|
), a tree species native to central and southern
|
|||
|
|
<collectingCountry id="F3767642361034180DECD38CBD06BDE2" box="[255,316,1159,1185]" name="Chile" pageId="9" pageNumber="410">Chile</collectingCountry>
|
|||
|
|
and the Andean parts of
|
|||
|
|
<collectingCountry id="F3767642361034180F41D38CBEFBBDE2" box="[594,705,1159,1185]" name="Argentina" pageId="9" pageNumber="410">Argentine</collectingCountry>
|
|||
|
|
Patagonia.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B6559361034180DD4D3A1B834BD86" box="[199,1038,1194,1221]" pageId="9" pageNumber="410" type="distribution">
|
|||
|
|
<paragraph id="8BDE36D2361034180DD4D3A1B834BD86" blockId="9.[151,1437,151,1221]" box="[199,1038,1194,1221]" pageId="9" pageNumber="410">
|
|||
|
|
<emphasis id="B915EAC0361034180DD4D3A1BD5BBD87" bold="true" box="[199,353,1194,1220]" pageId="9" pageNumber="410">Distribution.</emphasis>
|
|||
|
|
So far known from
|
|||
|
|
<collectingRegion id="49A5F830361034180F50D3A0BECCBD86" box="[579,758,1195,1221]" country="Chile" name="Biobio" pageId="9" pageNumber="410">Bío Bío Region</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="49A5F830361034180FECD3A0BF84BD86" box="[767,958,1195,1221]" country="Chile" name="Araucania" pageId="9" pageNumber="410">Arauco Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F3767642361034180ED9D3A0B830BD86" box="[970,1034,1195,1221]" name="Chile" pageId="9" pageNumber="410">Chile</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|