172 lines
18 KiB
XML
172 lines
18 KiB
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<document id="A1F640FF880FF5748759ADF8289E036D" ID-DOI="10.11646/zootaxa.5363.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146851" ID-ZooBank="B8979382-355B-410A-B1C9-20AD910233E6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700196986855" checkinUser="plazi" docAuthor="Riedel, Matthias" docDate="2023" docId="038C87C4FFFDFFD9BDAEF98DFD11672B" docLanguage="en" docName="zootaxa.5363.1.1.pdf" docOrigin="Zootaxa 5363 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5363.1.1/52218" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Achaius sabahensis Riedel 2023, nov. sp." docType="treatment" docVersion="2" lastPageNumber="11" masterDocId="FFB5FFBCFFF5FFD3BD39FF9FFF806516" masterDocTitle="Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia" masterLastPageNumber="94" masterPageNumber="1" pageNumber="9" updateTime="1700197734164" updateUser="ExternalLinkService">
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<mods:title id="D1DAC470114D2428E0D92CF1886BC804">Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) from Maritime Southeast Asia</mods:title>
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<mods:namePart id="F46B910A4E7A6B2D1BCCAA1C80EB36A9">Riedel, Matthias</mods:namePart>
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<mods:typeOfResource id="026619B3A3D3DEBB7F83C3F55162A964">text</mods:typeOfResource>
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<mods:title id="18EE53051CCE3DB7D32B3506FCD6E7ED">Zootaxa</mods:title>
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<mods:part id="6FDA89F1DFA31F447A1188594007F015">
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<mods:date id="14C327A97B54671D8C06FA238A134FEC">2023</mods:date>
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<mods:number id="F38B55F25E25468166507805435276B0">2023-11-06</mods:number>
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<mods:number id="3AAA6DD74E679084F87F507AA15DA068">5363</mods:number>
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<mods:classification id="AD071812D4C653A96DE4EA6159B91AAC">journal article</mods:classification>
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<mods:identifier id="B7F8970619F7E2AFE99BF444478FB511" type="DOI">10.11646/zootaxa.5363.1.1</mods:identifier>
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<mods:identifier id="3E3B2210ECC88C2442C801A508F05816" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="7FC5FC44F56381A2594CEC32D9351C86" type="Zenodo-Dep">10146851</mods:identifier>
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<treatment id="038C87C4FFFDFFD9BDAEF98DFD11672B" LSID="urn:lsid:plazi:treatment:038C87C4FFFDFFD9BDAEF98DFD11672B" httpUri="http://treatment.plazi.org/id/038C87C4FFFDFFD9BDAEF98DFD11672B" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
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<subSubSection id="C33F6559FFFDFFDBBDAEF98DFE6B633B" box="[151,491,1554,1581]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8B9A36D2FFFDFFDBBDAEF98DFE6B633B" blockId="8.[151,491,1554,1617]" box="[151,491,1554,1581]" pageId="8" pageNumber="9">
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<heading id="D0D281BEFFFDFFDBBDAEF98DFE6B633B" bold="true" box="[151,491,1554,1581]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="4">
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<emphasis id="B951EAC0FFFDFFDBBDAEF98DFE6B633B" bold="true" box="[151,491,1554,1581]" pageId="8" pageNumber="9">
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<taxonomicName id="4C254D51FFFDFFDBBDAEF98DFE06633B" authority="Riedel, 2023" authorityName="Riedel" authorityYear="2023" box="[151,390,1554,1581]" class="Insecta" family="Ichneumonidae" genus="Achaius" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
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<emphasis id="B951EAC0FFFDFFDBBDAEF98DFE06633B" bold="true" box="[151,390,1554,1581]" italics="true" pageId="8" pageNumber="9">Achaius sabahensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26257BBFFFDFFDBBCB4F98CFE6B633B" box="[397,491,1555,1581]" pageId="8" pageNumber="9" rank="species">nov. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6559FFFDFFDBBDAEF9A8FEA66347" box="[151,294,1591,1617]" pageId="8" pageNumber="9" type="description">
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<paragraph id="8B9A36D2FFFDFFDBBDAEF9A8FEA66347" blockId="8.[151,491,1554,1617]" box="[151,294,1591,1617]" pageId="8" pageNumber="9">
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(
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<figureCitation id="131E2A57FFFDFFDBBDA6F9A8FE9D6347" box="[159,285,1591,1617]" captionStart="FIGURE 2" captionStartId="9.[151,250,1540,1564]" captionTargetBox="[151,1408,181,1512]" captionTargetId="figure-16@9.[151,1436,181,1516]" captionTargetPageId="9" captionText="FIGURE 2. Achaius sabahensis n. sp. HT ♀. A face, frontal view, B head, dorsal view, C propodeum, dorsal view, D basal tergites, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10146855" httpUri="https://zenodo.org/record/10146855/files/figure.png" pageId="8" pageNumber="9">Figs 2A–D</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6559FFFDFFDBBDAEF9E0FE0063AB" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8B9A36D2FFFDFFDBBDAEF9E0FE0063AB" blockId="8.[151,1437,1663,2013]" pageId="8" pageNumber="9">
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<materialsCitation id="3B4D3C8FFFFDFFDBBDAEF9E0FE0063AB" collectingDate="1987-03-08" collectingDateMax="1987-03-11" collectingDateMin="1987-03-08" collectorName="Kinabalu Park Head Q. & C. van Achterberg" country="Malaysia" elevation="1600" location="Sabah" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
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<emphasis id="B951EAC0FFFDFFDBBDAEF9E0FE84638F" bold="true" box="[151,260,1663,1689]" pageId="8" pageNumber="9">
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<typeStatus id="549E8870FFFDFFDBBDAEF9E0FE84638F" box="[151,260,1663,1689]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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</emphasis>
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. (
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<specimenCount id="9D23FD5BFFFDFFDBBC22F9E0FEAD638F" box="[283,301,1663,1689]" count="1" pageId="8" pageNumber="9" type="female">♀</specimenCount>
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)
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<collectingCountry id="F3327642FFFDFFDBBC07F9E0FE29638F" box="[318,425,1663,1689]" name="Malaysia" pageId="8" pageNumber="9">Malaysia</collectingCountry>
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:
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<collectingRegion id="49E1F830FFFDFFDBBC8BF9E0FE79638F" box="[434,505,1663,1689]" country="Malaysia" name="Sabah" pageId="8" pageNumber="9">Sabah</collectingRegion>
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,
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<collectorName id="26D05304FFFDFFDBBF3AF9E0FC85638F" box="[515,773,1663,1689]" pageId="8" pageNumber="9">Kinabalu Park Head Q.</collectorName>
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,
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<date id="FF9B1012FFFDFFDBBE29F9E0FC26638F" box="[784,934,1663,1689]" pageId="8" pageNumber="9" value="1987-03-08" valueMax="1987-03-11" valueMin="1987-03-08">
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<collectingDate id="EFDFE9FAFFFDFFDBBE29F9E0FC26638F" box="[784,934,1663,1689]" pageId="8" pageNumber="9" value="1987-03-08" valueMax="1987-03-11" valueMin="1987-03-08">8–11.iii.1987</collectingDate>
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</date>
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, c
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<quantity id="4CDD9B37FFFDFFDBBEFAF9E0FB98638F" box="[963,1048,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="m" value="1600.0">
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<elevation id="0008D1E1FFFDFFDBBEFAF9E0FB98638F" box="[963,1048,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="9" unit="m" value="1600.0">1600 m</elevation>
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</quantity>
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,
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<collectingMethod id="52644EC5FFFDFFDBB91BF9E0FB2D638F" box="[1058,1197,1663,1689]" pageId="8" pageNumber="9">Malaise trap</collectingMethod>
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1,
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<collectorName id="26D05304FFFDFFDBB9F5F9E0FA18638F" box="[1228,1432,1663,1689]" pageId="8" pageNumber="9">C. van Achterberg</collectorName>
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, RMNH´87 (Leiden).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33F6559FFFDFFD9BDFEF959FED364C7" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
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<paragraph id="8B9A36D2FFFDFFDBBDFEF959FDFE63F7" blockId="8.[151,1437,1663,2013]" box="[199,638,1734,1761]" pageId="8" pageNumber="9">
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<emphasis id="B951EAC0FFFDFFDBBDFEF959FED163F6" bold="true" box="[199,337,1734,1760]" pageId="8" pageNumber="9">Description</emphasis>
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.
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<specimenCount id="9D23FD5BFFFDFFDBBC59F958FEF663F7" box="[352,374,1735,1761]" count="1" pageId="8" pageNumber="9" type="female">♀</specimenCount>
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. Body length
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<quantity id="4CDD9B37FFFDFFDBBF2AF959FDFA63F7" box="[531,634,1734,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="9" unit="mm" value="10.5">10.5 mm</quantity>
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.
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</paragraph>
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<paragraph id="8B9A36D2FFFDFFDBBDFEF974FF5062CB" blockId="8.[151,1437,1663,2013]" pageId="8" pageNumber="9">
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<emphasis id="B951EAC0FFFDFFDBBDFEF974FE846213" box="[199,260,1771,1797]" italics="true" pageId="8" pageNumber="9">Head</emphasis>
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. Flagellum with 40 flagellomeres, slender, almost filiform; 1
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<superScript id="7C509B9AFFFDFFDBBE91F974FC3363EF" attach="left" box="[936,947,1771,1785]" fontSize="6" pageId="8" pageNumber="9">st</superScript>
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flagellomere 3.7 × longer than wide, widest flagellomeres 1.2 × longer than wider. Temple rather long, distinctly and linearly narrowed behind eye (fig. 2B). OED 1.0 × and OOD 1.1 × ocellar diameter. Frons smooth ventrally, transversely rugose-punctate dorsally. Face densely punctate, shining. Clypeus almost flat, with sparse punctures; apical margin sharp and slightly concave (fig. 2A). Mandible long, with two teeth, ventral tooth smaller than dorsal one. Malar space 1.2 × as long as width of mandibular base. Gena with sparse fine punctures. Genal carina reaching hypostomal carina away from mandibular base.
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</paragraph>
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<caption id="DF5A665AFFFCFFDABDAEF99BFEE96356" ID-DOI="http://doi.org/10.5281/zenodo.10146855" ID-Zenodo-Dep="10146855" httpUri="https://zenodo.org/record/10146855/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1540,1564]" targetBox="[151,1408,181,1512]" targetPageId="9" targetType="figure">
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<paragraph id="8B9A36D2FFFCFFDABDAEF99BFEE96356" blockId="9.[151,1436,1540,1600]" pageId="9" pageNumber="10">
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<emphasis id="B951EAC0FFFCFFDABDAEF99BFE93630A" bold="true" box="[151,275,1540,1564]" pageId="9" pageNumber="10">FIGURE 2.</emphasis>
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<taxonomicName id="4C254D51FFFCFFDABC23F99BFE61630A" authority="Riedel, 2023" authorityName="Riedel" authorityYear="2023" box="[282,481,1540,1564]" class="Insecta" family="Ichneumonidae" genus="Achaius" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
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<emphasis id="B951EAC0FFFCFFDABC23F99BFE61630A" box="[282,481,1540,1564]" italics="true" pageId="9" pageNumber="10">Achaius sabahensis</emphasis>
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</taxonomicName>
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<emphasis id="B951EAC0FFFCFFDABCD1F99BFDA2630A" bold="true" box="[488,546,1540,1564]" pageId="9" pageNumber="10">
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<taxonomicNameLabel id="A26257BBFFFCFFDABCD1F99BFDA2630A" box="[488,546,1540,1564]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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HT ♀. A face, frontal view, B head, dorsal view, C propodeum, dorsal view, D basal tergites, dorsal view.
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</paragraph>
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</caption>
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<paragraph id="8B9A36D2FFFCFFDABDFEF9F0FB156292" blockId="9.[151,1437,1646,2033]" pageId="9" pageNumber="10">
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<taxonomicName id="4C254D51FFFCFFDABDFEF9F0FEBF639E" box="[199,319,1647,1672]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
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<emphasis id="B951EAC0FFFCFFDABDFEF9F0FEBF639E" box="[199,319,1647,1672]" italics="true" pageId="9" pageNumber="10">Mesosoma</emphasis>
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</taxonomicName>
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. Side of pronotum densely punctate dorsally, with irregular rugae ventrally. Notaulus indistinct. Mesoscutum finely and densely punctate, slightly shining. Mesopleuron densely punctate and finely striate ventrally, densely punctate dorsally; speculum smooth. Metapleuron rugose-punctate; juxtacoxal carina present. Scutellum moderately elevated, about as long as wide, with fine sparse punctures; without lateral carina. Propodeum (fig. 2C) rugose, with distinct blunt apophysis; spiracle slit-shaped. Area basalis not carinated, without median tubercle. Area superomedia almost rectangular, 1.4 × longer than wide; costula absent. Hind coxa densely punctate, without scopa. Hind femur slender, densely punctate, 5.1 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 5 × their diameter. Vein 1cu-a postfurcal by 3.0 × its width.
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</paragraph>
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<paragraph id="8B9A36D2FFFCFFD9BDFEF810FD0865EF" blockId="9.[151,1437,1646,2033]" lastBlockId="10.[151,1436,150,573]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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<emphasis id="B951EAC0FFFCFFDABDFEF810FEBC62BE" box="[199,316,1935,1960]" italics="true" pageId="9" pageNumber="10">Metasoma</emphasis>
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. Strongly amblypygous; hypopygium unmodified, with fine setae, reaching metasomal apex. 2
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<superScript id="7C509B9AFFFCFFDAB8B5F811FA1C628A" attach="none" box="[1420,1436,1934,1948]" fontSize="6" pageId="9" pageNumber="10">nd</superScript>
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and 3
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<superScript id="7C509B9AFFFCFFDABDEEF82DFF6562D6" attach="left" box="[215,229,1970,1984]" fontSize="6" pageId="9" pageNumber="10">rd</superScript>
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sternites with median folds. Postpetiole moderately widened, without latero-median carina; median field moderately elevated, finely rugose; lateral field punctate. Gastrocoelus impressed, without ridges. Thyridium very small, oblique, 0.2 × as wide as the interval between thyridia (fig. 2D). 2
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<superScript id="7C509B9AFFFFFFD9BE87FF09FC4E65B2" attach="left" box="[958,974,150,164]" fontSize="6" pageId="10" pageNumber="11">nd</superScript>
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tergite as long as wide, finely rugose and slightly aciculate in basal half, granulate apically, ± dull. Following tergites finely granulate, shining. Ovipositor sheath not reaching behind metasomal apex.
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</paragraph>
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<paragraph id="8B9A36D2FFFFFFD9BDFEFE9CFE6964BB" blockId="10.[151,1436,150,573]" pageId="10" pageNumber="11">
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<emphasis id="B951EAC0FFFFFFD9BDFEFE9CFE96640B" box="[199,278,259,285]" italics="true" pageId="10" pageNumber="11">Colour</emphasis>
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. Black. Flagellum black; flagellomeres 8–12 brownish dorsally. Face and gena with dark reddish suffusion; frontal orbit with reddish-yellow stripe. Hind edge of pronotum, tegula, subtegular ridge, central spot of mesoscutum and scutellum reddish. Tergites dark reddish-brown; 7
|
|||
|
|
<superScript id="7C509B9AFFFFFFD9BE47FED5FC0B644E" attach="left" box="[894,907,330,344]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
|
|||
|
|
tergite with ivory median spot. Legs black; fore and mid femora and tibia reddish-yellow stripes frontally; hind tibia diffusely reddish-brown sub-basally. Wings hyaline; pterostigma blackish.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9A36D2FFFFFFD9BDFEFE27FED364C7" blockId="10.[151,1436,150,573]" box="[199,339,439,467]" pageId="10" pageNumber="11">♁ unknown.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C33F6559FFFFFFD9BDFEFE44FA06670F" pageId="10" pageNumber="11" type="discussion">
|
|||
|
|
<paragraph id="8B9A36D2FFFFFFD9BDFEFE44FA06670F" blockId="10.[151,1436,150,573]" pageId="10" pageNumber="11">
|
|||
|
|
<emphasis id="B951EAC0FFFFFFD9BDFEFE44FEB464E3" bold="true" box="[199,308,475,501]" pageId="10" pageNumber="11">Remarks</emphasis>
|
|||
|
|
. This new species runs to
|
|||
|
|
<taxonomicName id="4C254D51FFFFFFD9BF6AFE44FC5064E3" authority="Heinrich, 1965" authorityName="Heinrich" authorityYear="1965" box="[595,976,474,501]" class="Insecta" family="Ichneumonidae" genus="Achaius" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="flavorufus">
|
|||
|
|
<emphasis id="B951EAC0FFFFFFD9BF6AFE44FCA464E3" box="[595,804,475,501]" italics="true" pageId="10" pageNumber="11">Achaius flavorufus</emphasis>
|
|||
|
|
<bibRefCitation id="EFB44B23FFFFFFD9BE10FE44FC5064E3" author="Heinrich, G. H." box="[809,976,474,501]" pageId="10" pageNumber="11" pagination="74 - 130" refId="ref58375" refString="Heinrich, G. H. (1965) Burmesische Ichneumoninae I. Entomologisk Tidskrift, 86, 74 - 130." type="journal article" year="1965">
|
|||
|
|
Heinrich,
|
|||
|
|
<quantity id="4CDD9B37FFFFFFD9BEA1FE45FC5064E3" box="[920,976,474,501]" metricMagnitude="1" metricUnit="m" metricValue="4.9911" pageId="10" pageNumber="11" unit="in" value="1965.0">1965</quantity>
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in the key of the Oriental
|
|||
|
|
<taxonomicName id="4C254D51FFFFFFD9B9D5FE44FA1C64E3" box="[1260,1436,475,501]" pageId="10" pageNumber="11">
|
|||
|
|
<emphasis id="B951EAC0FFFFFFD9B9D5FE44FAC664E3" box="[1260,1350,475,501]" italics="true" pageId="10" pageNumber="11">Achaius</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFB44B23FFFFFFD9BDA6FE60FE33670F" author="Heinrich, G. H." box="[159,435,510,537]" pageId="10" pageNumber="11" pagination="74 - 130" refId="ref58375" refString="Heinrich, G. H. (1965) Burmesische Ichneumoninae I. Entomologisk Tidskrift, 86, 74 - 130." type="journal article" year="1965">Heinrich 1965: 111–112</bibRefCitation>
|
|||
|
|
), but differs from all known Oriental species by the strong reduction of pale coloration.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C33F6559FFFFFFD9BDFEFDBDFD11672B" box="[199,657,546,573]" pageId="10" pageNumber="11" type="distribution">
|
|||
|
|
<paragraph id="8B9A36D2FFFFFFD9BDFEFDBDFD11672B" blockId="10.[151,1436,150,573]" box="[199,657,546,573]" pageId="10" pageNumber="11">
|
|||
|
|
<emphasis id="B951EAC0FFFFFFD9BDFEFDBDFED9672A" bold="true" box="[199,345,546,572]" pageId="10" pageNumber="11">Distribution</emphasis>
|
|||
|
|
. Only known from Borneo.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|