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<document id="82CC28C7E04A154916171BE2C1B87CB4" ID-CLB-Dataset="22593" ID-DOI="10.3897/zookeys.34.309" ID-GBIF-Dataset="09605303-c39a-4100-b5b4-fe409cccf3ab" ID-ISSN="1313–2970" ID-Zenodo-Dep="576611" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1588011467026" checkinUser="plazi" docAuthor="Gasca-Álvarez, Héctor &amp; Amat-García, Germán" docDate="2010" docId="F34FE44E1F24FFF8CD84F9ED566AFC7C" docLanguage="en" docName="ZK_article_2145.pdf" docOrigin="ZooKeys 34 (34)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@577d001f" docStyleName="zookeys.2008.journal_article" docTitle="Agaocephalini Burmeister 1847" docType="key" docVersion="7" lastPageNumber="157" masterDocId="0F769C361F27FFFCCD08FFFF523EFF96" masterDocTitle="Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia" masterLastPageNumber="192" masterPageNumber="153" pageNumber="156" updateTime="1698837288339" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo id="510FAC05D1FA296E58F4B997FDCBAAB1">
<mods:title id="97C65202DF63272F50E752D1D874BCFE">Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia</mods:title>
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<mods:namePart id="202D37943B69AF73DE844F5715A3EBE6">Gasca-Álvarez, Héctor</mods:namePart>
<mods:affiliation id="387D471D04A48E1CE30CED3A4F2C20E9">Researcher Corporación Sentido Natural ,, Colombia</mods:affiliation>
</mods:name>
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<mods:namePart id="63206F37FF4328CF71E02CF08097802D">Amat-García, Germán</mods:namePart>
<mods:affiliation id="6D218B24D14CF831A51A879B2C4D68F5">Universidad Nacional de Colombia, Bogotá ,, Colombia</mods:affiliation>
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<mods:part id="DADC6D1BD562BEC50298363D65C0FF79">
<mods:date id="A88C1EC76CD90717334A3BF7060E72F4">2010</mods:date>
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<mods:number id="263B236E2D8B7A064D752241F7DC543C">2010-01-28</mods:number>
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<mods:number id="52BA1F452892F49EE238400D46A4860C">34</mods:number>
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<mods:identifier id="2E307C11CA2C83CC5BC7D93A1D1E8A0C" type="DOI">10.3897/zookeys.34.309</mods:identifier>
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<mods:identifier id="5E2DE687881580CC19C8496F86F91F70" type="ISSN">1313–2970</mods:identifier>
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<treatment id="F34FE44E1F24FFF8CD84F9ED566AFC7C" ID-DOI="http://doi.org/10.5281/zenodo.3789622" ID-GBIF-Taxon="163794583" ID-Zenodo-Dep="3789622" LSID="urn:lsid:plazi:treatment:F34FE44E1F24FFF8CD84F9ED566AFC7C" httpUri="http://treatment.plazi.org/id/F34FE44E1F24FFF8CD84F9ED566AFC7C" lastPageId="4" lastPageNumber="157" pageId="3" pageNumber="156">
<subSubSection id="33FC06D31F24FFFFCD84F9ED5158F9BB" box="[140,870,1554,1581]" pageId="3" pageNumber="156" type="nomenclature">
<paragraph id="7B5955581F24FFFFCD84F9ED5158F9BB" blockId="3.[140,870,1554,1581]" box="[140,870,1554,1581]" pageId="3" pageNumber="156">
<heading id="2011E2341F24FFFFCD84F9ED5158F9BB" bold="true" box="[140,870,1554,1581]" fontSize="11" level="1" pageId="3" pageNumber="156" reason="1">
<emphasis id="4992894A1F24FFFFCD84F9ED5158F9BB" bold="true" box="[140,870,1554,1581]" pageId="3" pageNumber="156">
Key to the genera of adult
<taxonomicName id="BCE62EDB1F24FFFFCCFCF9ED508BF9BB" authorityName="Burmeister" authorityYear="1847" box="[500,693,1554,1581]" class="Insecta" family="Dynastidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="tribe" tribe="Agaocephalini">Agaocephalini</taxonomicName>
of
<collectingCountry id="03F115C81F24FFFFCFE8F9ED5158F9BB" box="[736,870,1554,1581]" name="Colombia" pageId="3" pageNumber="156">Colombia</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="33FC06D31F24FFF8CD84F9B5566AFC7C" lastPageId="4" lastPageNumber="157" pageId="3" pageNumber="156" type="key">
<paragraph id="7B5955581F24FFFFCD84F9B5566DF911" blockId="3.[140,1107,1609,1672]" pageId="3" pageNumber="156">
<keyLead id="C017F66D1F24FFFFCD84F9B5566DF911" pageId="3" pageNumber="156">
<keyStep id="C0124DFD1F24FFFFCD84F9B5566DF911" pageId="3" pageNumber="156">
1. Body surface tomentose, grayish brown color.............................................
<emphasis id="4992894A1F24FFFFC94CF9B6566DF9F5" bold="true" box="[1092,1107,1609,1635]" pageId="3" pageNumber="156">2</emphasis>
– Body surface glabrous, irregularly punctate ................................................
<emphasis id="4992894A1F24FFFFC94CF992566DF911" bold="true" box="[1092,1107,1645,1671]" pageId="3" pageNumber="156">3</emphasis>
</keyStep>
</keyLead>
</paragraph>
<keyStep id="C0124DFD1F23FFF8CD84FF58566DFEE6" pageId="4" pageNumber="157">
<paragraph id="7B5955581F23FFF8CD84FF58566DFE91" blockId="4.[140,1108,167,1003]" pageId="4" pageNumber="157">
<keyLead id="C017F66D1F23FFF8CD84FF58566DFE91" pageId="4" pageNumber="157">
2. External edge of mandibles with 3 teeth (
<figureCitation id="E3DD49DD1F23FFF8CFD8FF585133FF57" box="[720,781,167,193]" captionStart="Figure 3" captionStartId="5.[140,212,965,987]" captionTargetBox="[499,717,649,885]" captionTargetId="figure@5.[475,772,625,945]" captionTargetPageId="5" captionText="Figure 3. Right mandible of Lycomedes sp. (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770633" httpUri="https://zenodo.org/record/3770633/files/figure.png" pageId="4" pageNumber="157">Fig 3</figureCitation>
). Prosternal process short or moderately long. Pronotum with strong horn, short to large, flat, narrow, usually directed upwards (
<figureCitation id="E3DD49DD1F23FFF8CF13FF125069FE91" box="[539,599,237,263]" captionStart="Figure 4" captionStartId="5.[140,212,1654,1676]" captionTargetBox="[371,827,1029,1631]" captionTargetId="figure@5.[345,903,1015,1634]" captionTargetPageId="5" captionText="Figure 4. Male head and pronotum (lateral view) of Lycomedes sp. (scale bar: 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.3770635" httpUri="https://zenodo.org/record/3770635/files/figure.png" pageId="4" pageNumber="157">Fig 4</figureCitation>
).......................................
<emphasis id="4992894A1F23FFF8CE8BFF13566DFE91" bold="true" box="[899,1107,236,263]" pageId="4" pageNumber="157">
<taxonomicName id="BCE62EDB1F23FFF8CE8BFF1351C1FE91" box="[899,1023,236,263]" class="Insecta" family="Dynastidae" genus="Lycomedes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="4992894A1F23FFF8CE8BFF1351C1FE91" bold="true" box="[899,1023,236,263]" italics="true" pageId="4" pageNumber="157">Lycomedes</emphasis>
</taxonomicName>
<collectingRegion id="B9229BBA1F23FFF8C90EFF13566DFE91" box="[1030,1107,236,263]" country="Germany" name="Bremen" pageId="4" pageNumber="157">Bréme</collectingRegion>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="7B5955581F23FFF8CD84FEEE566DFEE6" blockId="4.[140,1108,167,1003]" pageId="4" pageNumber="157">
<keyLead id="C017F66D1F23FFF8CD84FEEE566DFEE6" pageId="4" pageNumber="157">
– External edge of mandibles with 2 teeth (
<figureCitation id="E3DD49DD1F23FFF8CFDFFEEF5128FEBD" box="[727,790,272,299]" captionStart="Figure 5" captionStartId="6.[140,212,464,486]" captionTargetBox="[515,729,195,424]" captionTargetId="figure@6.[496,751,165,444]" captionTargetPageId="6" captionText="Figure 5. Right mandible of Spodistes sp. (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3770637" httpUri="https://zenodo.org/record/3770637/files/figure.png" pageId="4" pageNumber="157">Fig 5</figureCitation>
). Prosternal process absent. Pronotum with large horn, directed forward, thin and curved (
<figureCitation id="E3DD49DD1F23FFF8CEB1FECC51CBFED8" box="[953,1013,307,334]" captionStart="Figure 6" captionStartId="6.[140,212,1043,1065]" captionTargetBox="[322,937,544,1011]" captionTargetId="figure@6.[305,943,514,1023]" captionTargetPageId="6" captionText="Figure 6. Male head and pronotum (lateral view) of Spodistes sp. (scale bar: 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.3770639" httpUri="https://zenodo.org/record/3770639/files/figure.png" pageId="4" pageNumber="157">Fig 6</figureCitation>
) .......... ...................................................................................
<taxonomicName id="BCE62EDB1F23FFF8CE54FEA9566DFEE6" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[860,1107,342,369]" class="Insecta" family="Dynastidae" genus="Spodistes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="4992894A1F23FFF8CE54FEA9566DFEE6" bold="true" box="[860,1107,342,369]" pageId="4" pageNumber="157">
<emphasis id="4992894A1F23FFF8CE54FEA951F7FEE7" bold="true" box="[860,969,342,369]" italics="true" pageId="4" pageNumber="157">Spodistes</emphasis>
Burmeister
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C0124DFD1F23FFF8CD84FE85566DFE6B" pageId="4" pageNumber="157">
<paragraph id="7B5955581F23FFF8CD84FE85566DFE4C" blockId="4.[140,1108,167,1003]" pageId="4" pageNumber="157">
<keyLead id="C017F66D1F23FFF8CD84FE85566DFE4C" pageId="4" pageNumber="157">
3. Protibia quadridentate, basal tooth short in minor males. Head of males with two large, divergent, forward directed horns; horns curved upward, apices rounded (
<figureCitation id="E3DD49DD1F23FFF8CC6CFE3F539CFE4D" box="[356,418,448,475]" captionStart="Figure 7" captionStartId="6.[150,222,1653,1675]" captionTargetBox="[382,861,1196,1590]" captionTargetId="figure@6.[375,873,1102,1632]" captionTargetPageId="6" captionText="Figure 7. Male head and pronotum (lateral view) of Aegopsis curvicornis Burmeister (scale bar: 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.3770641" httpUri="https://zenodo.org/record/3770641/files/figure.png" pageId="4" pageNumber="157">Fig 7</figureCitation>
). Females lacking horns........................
<taxonomicName id="BCE62EDB1F23FFF8CE6EFE3F566DFE4C" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[870,1107,448,474]" class="Insecta" family="Dynastidae" genus="Aegopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="4992894A1F23FFF8CE6EFE3F566DFE4C" bold="true" box="[870,1107,448,474]" pageId="4" pageNumber="157">
<emphasis id="4992894A1F23FFF8CE6EFE3F51F6FE4C" bold="true" box="[870,968,448,474]" italics="true" pageId="4" pageNumber="157">Aegopsis</emphasis>
Burmeister
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7B5955581F23FFF8CD84FE1B566DFE6B" blockId="4.[140,1108,167,1003]" box="[140,1107,483,510]" pageId="4" pageNumber="157">
<keyLead id="C017F66D1F23FFF8CD84FE1B566DFE6B" box="[140,1107,483,510]" pageId="4" pageNumber="157">
– Protibia tridentate. Head with thick horn, curved upward, apex bifurcated .....
<emphasis id="4992894A1F23FFF8C94CFE1C566DFE6B" bold="true" box="[1092,1107,483,509]" pageId="4" pageNumber="157">4</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C0124DFD1F23FFF8CD84FDF8566DFD47" pageId="4" pageNumber="157">
<paragraph id="7B5955581F23FFF8CD84FDF8566DFD1C" blockId="4.[140,1108,167,1003]" pageId="4" pageNumber="157">
<keyLead id="C017F66D1F23FFF8CD84FDF8566DFD1C" pageId="4" pageNumber="157">
4. Males with frontal horn short and thick, slightly curved. Pronotum with forward directed knob with acute apex, surface smooth. Females lacking horn on head and knob on pronotum. Head and pronotum black, elytra reddish brown ....................................................................
<taxonomicName id="BCE62EDB1F23FFF8CE33FD8F566DFD1C" authority="Thomson" authorityName="Thomson" authorityYear="1859" box="[827,1107,624,651]" class="Insecta" family="Dynastidae" genus="Mitracephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="4992894A1F23FFF8CE33FD8F566DFD1C" bold="true" box="[827,1107,624,651]" pageId="4" pageNumber="157">
<emphasis id="4992894A1F23FFF8CE33FD8F51E2FD1D" bold="true" box="[827,988,624,651]" italics="true" pageId="4" pageNumber="157">Mitracephala</emphasis>
Thomson
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7B5955581F23FFF8CD84FD6B566DFD47" blockId="4.[140,1108,167,1003]" pageId="4" pageNumber="157">
<keyLead id="C017F66D1F23FFF8CD84FD6B566DFD47" pageId="4" pageNumber="157">
– Males with frontal horn large, curved upwards, apex bifurcate. Pronotum convex or with high knob. Body black, brown, or with dark spots .............
<emphasis id="4992894A1F23FFF8C94CFD48566DFD47" bold="true" box="[1092,1107,695,721]" pageId="4" pageNumber="157">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C0124DFD1F23FFF8CD84FD25566AFC7C" pageId="4" pageNumber="157">
<paragraph id="7B5955581F23FFF8CD84FD25566DFCCB" blockId="4.[140,1108,167,1003]" pageId="4" pageNumber="157">
<keyLead id="C017F66D1F23FFF8CD84FD25566DFCCB" pageId="4" pageNumber="157">
5. Apex of frontal horn in males bifurcate. Pronotum with wide, high knob; apex rounded, surface densely punctate. Females lacking frontal horn. Elytra with punctures strongly impressed. Prosternal process high, apex dilated. Body black or brown.................................................
<taxonomicName id="BCE62EDB1F23FFF8CE46FCBC566DFCCB" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[846,1107,834,862]" class="Insecta" family="Dynastidae" genus="Horridocalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="4992894A1F23FFF8CE46FCBC566DFCCB" bold="true" box="[846,1107,834,862]" pageId="4" pageNumber="157">
<emphasis id="4992894A1F23FFF8CE46FCBC51D7FCC8" bold="true" box="[846,1001,835,862]" italics="true" pageId="4" pageNumber="157">Horridocalia</emphasis>
Endrődi
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7B5955581F23FFF8CD84FC98566AFC7C" blockId="4.[140,1108,167,1003]" pageId="4" pageNumber="157">
<keyLead id="C017F66D1F23FFF8CD84FC98566AFC7C" pageId="4" pageNumber="157">
– Apex of frontal horn in males emarginated or bifurcated. Head of females with 2 transversal tubercles. Pronotum convex, smooth. Elytra strongly reticulated and finely punctate. Prosternal process absent. Body yellowish brown with dark spots .............................................................
<taxonomicName id="BCE62EDB1F23FFF8CE66FC2F566AFC7C" authority="Prell" authorityName="Prell" box="[878,1108,976,1003]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="4992894A1F23FFF8CE66FC2F566AFC7C" bold="true" box="[878,1108,976,1003]" pageId="4" pageNumber="157">
<emphasis id="4992894A1F23FFF8CE66FC2F562AFC7D" bold="true" box="[878,1044,976,1003]" italics="true" pageId="4" pageNumber="157">Brachysiderus</emphasis>
Prell
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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