240 lines
25 KiB
XML
240 lines
25 KiB
XML
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1171.104142" ID-Pensoft-Pub="1313-2970-1171-1" ID-Pensoft-UUID="7559C2D42DE85144AEFFC98A16172F97" ID-ZooBank="267D0D4559CA4A18A08034768E652607" ModsDocID="1313-2970-1171-1" checkinTime="1689969829564" checkinUser="pensoft" docAuthor="Short, Andrew Edward Z. & Giron, Jennifer C." docDate="2023" docId="F3445DA102645F5AA4C9FE7CB3229119" docLanguage="en" docName="ZooKeys 1171: 1-112" docOrigin="ZooKeys 1171" docPubDate="2023-07-20" docSource="http://dx.doi.org/10.3897/zookeys.1171.104142" docTitle="Novochares tenedor Short & Girón 2023, sp. nov." docType="treatment" docUuid="B95F2EE8-D28B-4EAE-B161-AA2783A25F57" docUuidSource="ZooBank" docVersion="1" id="7559C2D42DE85144AEFFC98A16172F97" lastPageNumber="1" masterDocId="7559C2D42DE85144AEFFC98A16172F97" masterDocTitle="Revision of the Neotropical water scavenger beetle genus Novochares Giron & Short (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="112" masterPageNumber="1" pageNumber="1" updateTime="1689969829564" updateUser="pensoft">
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<mods:mods id="140E647D3616B1B5875D47C3477ED3FE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DDEAB0DFF8AD427E077602DA9D545B98">
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<mods:title id="CEC081B2F111649106E6BF6F0A242230">Revision of the Neotropical water scavenger beetle genus Novochares Giron & Short (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="7AAFB416BEB678C00EEBC3533F87C139" type="personal">
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<mods:roleTerm id="884768F6D222A72934AB1B49CA51F5A9">Author</mods:roleTerm>
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<mods:namePart id="D8DF879F809D8ADDED4AEBF10169F18B">Short, Andrew Edward Z.</mods:namePart>
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<mods:nameIdentifier id="2F09FF0C3CAB1A87111A525574087B0E" type="ORCID">https://orcid.org/0000-0002-7467-7116</mods:nameIdentifier>
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<mods:affiliation id="9173395CAA78AFA46B9CD7C4B681B23B">Department of Entomology & Nematology, University of Florida, Gainesville, FL, 32611, USA</mods:affiliation>
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<mods:nameIdentifier id="CA6A52D068AE013643556A06408905E7" type="email">aezshort@ku.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="221B9C08CF523833E2EA1081A32A0AC5" type="personal">
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<mods:roleTerm id="93C54D53D5C1D9C9E2407DC25972ACDA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="36D873DC71E4270FEBD1842FB78D662B">Giron, Jennifer C.</mods:namePart>
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<mods:nameIdentifier id="9F8DCB8BC4A969AD0C68EAE290E2B4D7" type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
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<mods:affiliation id="088EA59AEAAAAABC76AB8292870469FC">Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, TX 79409, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="26DD7D14A945939D275574D8C9DD2802">text</mods:typeOfResource>
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<mods:titleInfo id="4896C3BF655358CB23C55E35C64EBF8F">
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<mods:title id="F670C1FA7DA6FA49F18224B94B1B24AD">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="3815DD0F924C805A921A1A40D4B5B962">
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<mods:date id="C24BDE21796DE08B5C31C29A915D5E57">2023</mods:date>
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<mods:detail id="3B449056E6B162603964A03D2A7FA7A0" type="pubDate">
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<mods:number id="944F877F2900142126A5246CD4F00950">2023-07-20</mods:number>
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</mods:detail>
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<mods:detail id="748BEBBFCA6EB66AC59C57D6CAB577F7" type="volume">
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<mods:number id="887DC0BA21691AD6E7A33A72F2F78E8E">1171</mods:number>
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</mods:detail>
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<mods:start id="6D25E27B75E58D7CEB2BA9C31C412048">1</mods:start>
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<mods:end id="1909544FC287E7D6553AAED960F7F99F">112</mods:end>
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<mods:location id="B0A6DE088AD7B69E3E97BDEA11B02A10">
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<mods:url id="FCB229C02C5779967236132BADDED285">http://dx.doi.org/10.3897/zookeys.1171.104142</mods:url>
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</mods:location>
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<mods:classification id="7495AA4DCD4E7C31457147317A68382C">journal article</mods:classification>
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<mods:identifier id="FEF3AA3692B5ABBC7634CBB0E69081A8" type="DOI">http://dx.doi.org/10.3897/zookeys.1171.104142</mods:identifier>
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<mods:identifier id="0B693C469C285D4C6BF9D68570DEAF3D" type="Pensoft-Pub">1313-2970-1171-1</mods:identifier>
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<mods:identifier id="E6B2A8AEC4185E0C6589E20C73DAE1DC" type="ZooBank">267D0D4559CA4A18A08034768E652607</mods:identifier>
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<mods:identifier id="658EABC7EFB88CE6CC63080AA3B2ED1C" type="Pensoft-UUID">7559C2D42DE85144AEFFC98A16172F97</mods:identifier>
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</mods:mods>
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<treatment id="F3445DA102645F5AA4C9FE7CB3229119" LSID="urn:lsid:zoobank.org:act:B95F2EE8-D28B-4EAE-B161-AA2783A25F57" httpUri="http://treatment.plazi.org/id/F3445DA102645F5AA4C9FE7CB3229119" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="4A1DD3B3477C7DE93CDC1068E562CF7A" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="091D8B9F3F7945056B4ABF3E5977F600" pageId="0" pageNumber="1">
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<taxonomicName id="7F226D3432E61B6A539013627B9FA4FC" LSID="https://zoobank.org/B95F2EE8-D28B-4EAE-B161-AA2783A25F57" authority="Short & Girón, 2023" authorityName="Short & Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares tenedor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tenedor" status="sp. nov.">Novochares tenedor</taxonomicName>
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<taxonomicNameLabel id="2C920D25EA3C74B1B3AF4A7BC1B34DA2" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D1E520B9C7AE437D1AA926D4A1BA9D0C" pageId="0" pageNumber="1" type="description">
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<paragraph id="9C482ADA1FA61C0189308AFE774A153A" pageId="0" pageNumber="1">
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<figureCitation id="EE206B3BB3CE8FBF690C495BF33A64CD" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagi of the Novochares garfo species group A-C N. bidens D-F N. furcatus G-J N. garfo K-M N. tenedor A, D, G, K dorsal view B, E, H, L ventral view C, I, M lateral view F, J oblique view." figureDoi="10.3897/zookeys.1171.104142.figure12" httpUri="https://binary.pensoft.net/fig/881179" pageId="0" pageNumber="1">Figs 12K-M</figureCitation>
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<figureCitation id="23384F9EAE79660E51A2CAE4016CD9C2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Novochares spp. A N. aperito (red), N. minor (yellow), N. orchis (blue) B N. bidens (red), N. furcatus (yellow), N. garfo (blue), N. tenedor (green)." figureDoi="10.3897/zookeys.1171.104142.figure13" httpUri="https://binary.pensoft.net/fig/881180" pageId="0" pageNumber="1">, 13B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F9B87DA2EC287731008AED52ECA34F6B" pageId="0" pageNumber="1" type="type material">
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<paragraph id="0DF5F8DDDD69B012D4A74A7E7365F819" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="46E6E74576A1420559701A8B2495A2B4" pageId="0" pageNumber="1">
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<emphasis id="0B724426A981217CA9AC699358BD7C53" bold="true" pageId="0" pageNumber="1">
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<emphasis id="6C559C8104DEE38E38B860F647C8A23C" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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(male)
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</emphasis>
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: "VENEZUELA: Apure State/
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<geoCoordinate id="C4F45C141079D7E433DA079DEDDAE23E" degrees="7" direction="north" minutes="37.289" orientation="latitude" precision="1" value="7.6214833">7°37.289'N</geoCoordinate>
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,
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<geoCoordinate id="F572A51556518CB374752EC1B3D6EC27" degrees="69" direction="west" minutes="3.679" orientation="longitude" precision="1" value="-69.06132">69°3.679'W</geoCoordinate>
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, 83m/ side road ca. 10 km E. Mantecal/ leg. Short,
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<normalizedToken id="2F599153EC61E3B48646BC6D4A2DA697" originalValue="García">Garcia</normalizedToken>
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, & Camacho/ 18.i.2009; marshy area and pool by road; VZ09-0118-02X" (MIZA).
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<emphasis id="0EE23B91A020D7479EE66133C7DAC35F" bold="true" pageId="0" pageNumber="1">
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<emphasis id="B04BB4D8B3F66BE65049AB6A2C64AB85" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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(59 exs.): Guyana: Region 6
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</emphasis>
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: Upper Berbice, Basecamp 1,
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<geoCoordinate id="BEE37D215DC446034467D02BEB533E3C" degrees="4" direction="north" minutes="08.809" orientation="latitude" precision="1" value="4.1468167">4°08.809'N</geoCoordinate>
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,
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<geoCoordinate id="194D06F23C4495346ED8C3340811B8F9" degrees="58" direction="west" minutes="14.232" orientation="longitude" precision="1" value="-58.2372">58°14.232'W</geoCoordinate>
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, 108 m, 22.ix.2014, leg. Short, Salisbury, La Cruz, margin of Berbice River, GY14-0922-02A (36, SEMC, CBDG, TTU-Z, including DNA voucher SLE1219).
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<emphasis id="1721007D3287741CD25C0D35DD79134E" bold="true" pageId="0" pageNumber="1">Region 9</emphasis>
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: Karanambu,
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<geoCoordinate id="95E4AB87B53193BB164D4C760F1237B1" degrees="3" direction="north" minutes="45.1" orientation="latitude" precision="92" value="3.7516665">3°45.1'N</geoCoordinate>
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,
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<geoCoordinate id="6D8507FA6CCC720AF6DD9E963443F06F" degrees="59" direction="west" minutes="18.6" orientation="longitude" precision="92" value="-59.31">59°18.6'W</geoCoordinate>
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, Rupununi River, 2.iv.1994, leg. P.J. Spangler, colln #8 (1, USNM).
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<emphasis id="7E6BD8D7CC53CF8CBCA1C8AE312D6443" bold="true" pageId="0" pageNumber="1">Venezuela: Apure</emphasis>
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: Same data as holotype (1, SEMC, DNA voucher SLE1205).
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<emphasis id="8B2FB2ECF0AEC4E8CDCE46ACCF9F988A" bold="true" pageId="0" pageNumber="1">Barinas</emphasis>
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: SW of Batatuy,
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<geoCoordinate id="80BE3738C810A9FAA1993634B9B4330E" degrees="8" direction="north" minutes="10.259" orientation="latitude" precision="1" value="8.170983">8°10.259'N</geoCoordinate>
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,
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<geoCoordinate id="0E078936209EBDBBE56469B0C02DDC9E" degrees="70" direction="west" minutes="51.866" orientation="longitude" precision="1" value="-70.86443">70°51.866'W</geoCoordinate>
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, 275 m, 25.i.2012, leg. Short, Arias, & Gustafson, sandbar/gravel margin, VZ12-0125-03C (1, SEMC).
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<emphasis id="41B7A775D3A50177E01FFEA46B23BCF6" bold="true" pageId="0" pageNumber="1">
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<normalizedToken id="F2D73F646F1012DE5B1896982855A906" originalValue="Bolívar">Bolivar</normalizedToken>
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</emphasis>
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: Cuchivero, 30 km SE of Caicara, 4.viii.1987, leg. S. & J. Peck, Woodland, UV light, SBP87-108 (6, SEMC).
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<emphasis id="573BB89ACA3E75187EF98E2C03CA5CD0" bold="true" pageId="0" pageNumber="1">Cojedes</emphasis>
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: Rio Tinaco, near, approx. 5 km NE of Tinaco,
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<geoCoordinate id="1BF49965EF22E64F26AD29C12832B238" degrees="9" direction="north" minutes="44.160" orientation="latitude" precision="1" value="9.736">9°44.160'N</geoCoordinate>
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,
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<geoCoordinate id="B68F5E8FB1E197025CD7DBFBB9B43CE0" degrees="68" direction="west" minutes="24.219" orientation="longitude" precision="1" value="-68.40365">68°24.219'W</geoCoordinate>
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, 170 m, 20.i.2012, leg. Short, Arias, & Gustafson, stream margins, VZ12-0120-02A (4, SEMC); El Pao, approx. 7.5 km, Rio Caiman Grande at San Brano,
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<geoCoordinate id="281F3E53A0554585C9306CD187FFAE2A" degrees="9" direction="north" minutes="39.246" orientation="latitude" precision="1" value="9.6541">9°39.246'N</geoCoordinate>
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,
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<geoCoordinate id="470A66EAB002787D40C932B0EEB8CBA0" degrees="68" direction="west" minutes="11.860" orientation="longitude" precision="1" value="-68.19767">68°11.860'W</geoCoordinate>
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, 137 m, 20.i.2012, leg. Short, Arias, & Gustafson, stream margins, VZ12-0120-03A (3, SEMC); Aparicion, at highway, lagoon/pond,
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<geoCoordinate id="CB9AB1F809F1B8B10F98002693DDDD3A" degrees="9" direction="north" minutes="22.268" orientation="latitude" precision="1" value="9.371134">9°22.268'N</geoCoordinate>
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,
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<geoCoordinate id="61CC1B81D379A34810A6CF5E922A2156" degrees="69" direction="west" minutes="23.062" orientation="longitude" precision="1" value="-69.38437">69°23.062'W</geoCoordinate>
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, 213 m, 22.i.2012, leg. Short, Arias, & Gustafson, pond, VZ12-0122-01A (2, SEMC).
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<emphasis id="61FC91FE262DE2FEC96CB06A811E1848" bold="true" pageId="0" pageNumber="1">Portuguesa</emphasis>
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: Aparicion, Rio Are,
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<geoCoordinate id="D47E6856183C05949443C5414E635706" degrees="9" direction="north" minutes="22.900" orientation="latitude" precision="1" value="9.381666">9°22.900'N</geoCoordinate>
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,
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<geoCoordinate id="4095EF84B68B9D781278E706814785A4" degrees="69" direction="west" minutes="23.153" orientation="longitude" precision="1" value="-69.38588">69°23.153'W</geoCoordinate>
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, 220 m, 22.i.2012, leg. Short & Arias, river margin, VZ12-0122-02A (5, SEMC).
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<emphasis id="3A36650992223F1A509666DA4A0206A5" bold="true" pageId="0" pageNumber="1">Zulia</emphasis>
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: Quebrada Riencito,
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<geoCoordinate id="B7F69AA26A92D369752CC121AF4046FE" degrees="10.86041" direction="north" orientation="latitude" precision="1" value="10.86041">10.86041°N</geoCoordinate>
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,
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<geoCoordinate id="1D70190277FB857AFAC56AB59E137F92" degrees="72.32210" direction="west" orientation="longitude" precision="1" value="-72.3221">72.32210°W</geoCoordinate>
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, 95 m, 30.xii.2008, leg. Short &
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<normalizedToken id="7A1B116F932EF1D65AE002B4CE71B4D5" originalValue="García">Garcia</normalizedToken>
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, along margin, VZ08-1230-01B (1, SEMC).
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</paragraph>
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</subSubSection>
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<subSubSection id="71E0AD56DE1115E0DDA39FD90B87E323" pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph id="6FD395BED4DD0874B51CD6A5F8AC3F30" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph id="B19188B289FA76966AE9E06BD833C50E" pageId="0" pageNumber="1">
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This taxon is unique among members of this species group in having the parameres strongly sinuate along the apical 1/3 (Fig.
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<figureCitation id="2763F605D912497A30A6AD2DEA37983B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagi of the Novochares garfo species group A-C N. bidens D-F N. furcatus G-J N. garfo K-M N. tenedor A, D, G, K dorsal view B, E, H, L ventral view C, I, M lateral view F, J oblique view." figureDoi="10.3897/zookeys.1171.104142.figure12" httpUri="https://binary.pensoft.net/fig/881179" pageId="0" pageNumber="1">12K, L</figureCitation>
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). While the dorsal plate of the median lobe is slightly similar in form to
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<taxonomicName id="0EC5F73193A8465C385A28FBB22D2FD7" lsidName="N. bidens" pageId="0" pageNumber="1" rank="species" species="bidens">
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<emphasis id="81292381F6FF71EEDF945E093D2538B7" italics="true" pageId="0" pageNumber="1">N. bidens</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation id="C13F14F61BDE0D4A8D39D3DA872911B7" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagi of the Novochares garfo species group A-C N. bidens D-F N. furcatus G-J N. garfo K-M N. tenedor A, D, G, K dorsal view B, E, H, L ventral view C, I, M lateral view F, J oblique view." figureDoi="10.3897/zookeys.1171.104142.figure12" httpUri="https://binary.pensoft.net/fig/881179" pageId="0" pageNumber="1">12A, B</figureCitation>
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), the arms of the fork are less than 1/2 as long, and the plate itself is much shorter, sitting well below the apex of the parameres.
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</paragraph>
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</subSubSection>
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<subSubSection id="311B29F0A66CD2F4FFFD1D2CE982EA2E" pageId="0" pageNumber="1" type="description">
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<paragraph id="21809382F2AD62799699D0C4BEB96033" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="AADDAA6EF524EDCC837A4B3A6B3E1B2C" pageId="0" pageNumber="1">
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Body length 5.1-6.5 mm.
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<emphasis id="BFCCD1DCD90DD3695B4928AC3E1C9BBB" bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
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: Dorsal surfaces orange to yellowish brown, usually with slightly paler (yellow) clypeus and margins of pronotum and elytra.
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<emphasis id="29EB0F390870CAE3B7D63A5BD4715E3B" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
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: Maxillary palps slightly nearly 1.3
|
|||
|
<normalizedToken id="A7F55F3C4F9DFC2015D68E11CCAE8D78" originalValue="×">x</normalizedToken>
|
|||
|
longer than width of head, uniformly orange in color.
|
|||
|
<emphasis id="20D2731AF50C017476F59A959329D9BB" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
|
|||
|
: Elytra without rows of serial punctures, each with very faint rows of scarce and weakly marked systematic punctures on lateral and posterior regions. Prosternum medially broadly and weakly convex. Posterior elevation of mesoventrite somewhat transverse and broadly elevated, with low and glabrous longitudinal ridge extending anteriorly.
|
|||
|
<emphasis id="8F9BB33F3D3531280F1B58EB55CA2D20" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
|||
|
: Apical emargination of fifth ventrite relatively deep, U-shaped.
|
|||
|
<emphasis id="6DCC6466662B7BAE2C9F578FFA66AD96" bold="true" italics="true" pageId="0" pageNumber="1">Aedeagus</emphasis>
|
|||
|
: (Fig.
|
|||
|
<figureCitation id="09FD2EC9AB62E3E8557A7F27DFBE9595" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Aedeagi of the Novochares garfo species group A-C N. bidens D-F N. furcatus G-J N. garfo K-M N. tenedor A, D, G, K dorsal view B, E, H, L ventral view C, I, M lateral view F, J oblique view." figureDoi="10.3897/zookeys.1171.104142.figure12" httpUri="https://binary.pensoft.net/fig/881179" pageId="0" pageNumber="1">12K-M</figureCitation>
|
|||
|
) Overall shape pear-like, 2.3
|
|||
|
<normalizedToken id="B960E79D1F28FCBA5878E324E308346C" originalValue="×">x</normalizedToken>
|
|||
|
longer than wide, with outer lateral margins of parameres very weakly convex up to apical region; joint basal margins of parameres medially pointed in dorsal view; apical region of each paramere rounded, with outer margin laterally pointed; at closest point (along neck), dorsal inner margins of parameres separated by distance 0.54
|
|||
|
<normalizedToken id="EF950480E19A3D7F921D66C232DC3797" originalValue="×">x</normalizedToken>
|
|||
|
greatest width of a paramere; dorsal plate of median lobe with neck 0.23
|
|||
|
<normalizedToken id="0F0203C9C2A131A94BC118D460D1A583" originalValue="×">x</normalizedToken>
|
|||
|
as broad as base; base of arms of dorsal plate of median lobe dorsally concave; arms of dorsal plate somewhat parallel, diverging at apex, nearly parallel-sided for most length, nearly 0.16
|
|||
|
<normalizedToken id="9736CD4F38FB2243A0D95751F4277820" originalValue="×">x</normalizedToken>
|
|||
|
length of dorsal plate of median lobe; each arm acute and dorso-laterally pointed at apex; notch between arms at base nearly 1/2 as wide as base of an arm; basal piece nearly 0.3
|
|||
|
<normalizedToken id="B13F914D8BE3E551F33361EE15E626DC" originalValue="×">x</normalizedToken>
|
|||
|
length of a paramere. In lateral view, aedeagus straight at base, with ventral outline of parameres 3.9
|
|||
|
<normalizedToken id="62F592E15FD45A6F78081935D9B9A6FB" originalValue="×">x</normalizedToken>
|
|||
|
longer than greatest width near base; dorsal outline of aedeagus in lateral view strongly convex at base, then nearly straight and oblique along basal 1/2, then sinuate along distal 1/2; ventral outline of aedeagus in lateral view nearly straight.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="D49E9574049C15993FE95FFFA7152623" pageId="0" pageNumber="1" type="etymology">
|
|||
|
<paragraph id="79C25F5C65317BD13396D7AD7E22CA92" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|||
|
<paragraph id="C74D3D18C528335D41DEC17C6C2D1531" pageId="0" pageNumber="1">Tenedor, meaning fork in Spanish, in reference to the shape of the dorsal plate of the median lobe.</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="2BFB8EDBCC0827D152AFCF4E3001AA52" pageId="0" pageNumber="1" type="distribution">
|
|||
|
<paragraph id="1BE5E08AAB68648717F4F29E0A553780" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|||
|
<paragraph id="DA66578D1D0518725ECE384B7F2B10DF" pageId="0" pageNumber="1">
|
|||
|
Guyana, Venezuela (Fig.
|
|||
|
<figureCitation id="9B698379D524611B7842474F7F802D53" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Distribution of Novochares spp. A N. aperito (red), N. minor (yellow), N. orchis (blue) B N. bidens (red), N. furcatus (yellow), N. garfo (blue), N. tenedor (green)." figureDoi="10.3897/zookeys.1171.104142.figure13" httpUri="https://binary.pensoft.net/fig/881180" pageId="0" pageNumber="1">13B</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="D4769147F3F77135969D42F6AB429E53" pageId="0" pageNumber="1" type="habitat">
|
|||
|
<paragraph id="FFD305B15CF06B7AEFE74CBB272B4503" pageId="0" pageNumber="1">Habitat.</paragraph>
|
|||
|
<paragraph id="F62BD790775A2564329E1841621BD217" pageId="0" pageNumber="1">This species has been collected in a range of habitats, though most specimens seem to have been taken in riparian areas along the margins of streams and rivers. Other specimens have been collected in more typical marsh habitats.</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="D2E5C331BC49C6716FCFCC1E5B43A8F3" pageId="0" pageNumber="1" type="novochares minor species group">
|
|||
|
<paragraph id="86793211D460B447F22C8B6253AE3ECC" pageId="0" pageNumber="1">
|
|||
|
<taxonomicName id="27634F178657ED18DF757515A99D7EC4" authorityName="Short & Girón" authorityYear="2023" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares minor" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="minor">
|
|||
|
<emphasis id="D9C2A28438B00D2CDF945C165B4229A5" italics="true" pageId="0" pageNumber="1">Novochares minor</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
species group
|
|||
|
</paragraph>
|
|||
|
<paragraph id="4804776EDDE40968423CE0A97699CB59" pageId="0" pageNumber="1">
|
|||
|
<emphasis id="7457D1FF0B4443012198092CFDBF3CDC" bold="true" pageId="0" pageNumber="1">Species group diagnosis.</emphasis>
|
|||
|
Body length 4.2-5.4 mm.
|
|||
|
<emphasis id="E2EE2BA5A0D2F5219896E07F6761849C" bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
|
|||
|
: Dorsal surfaces very dark brown, with paler (orange to yellow) margins of pronotum and elytra.
|
|||
|
<emphasis id="19BEAF7BABEAA62D46051F9D588DF449" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|||
|
: Maxillary palps slightly longer than width of head, orange to brown in color, usually paler (yellow) at ends of each palpomere (Fig.
|
|||
|
<figureCitation id="D7FD5EAD42524100C13E6D5E26F77F8B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Dorsal habitus of Novochares spp. A N. abbreviatus B N. pilatus C N. minor D N. tenedor." figureDoi="10.3897/zookeys.1171.104142.figure7" httpUri="https://binary.pensoft.net/fig/881174" pageId="0" pageNumber="1">7C</figureCitation>
|
|||
|
).
|
|||
|
<emphasis id="8BBB74761191842C668E5AF69E5EED23" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
|
|||
|
: Ground punctation on pronotum and elytra relatively dense and shallowly impressed. Elytra without rows of serial punctures, each with very faint rows (one dorsal and two or three lateral) of scarce and weakly marked systematic punctures. Prosternum only very weakly medially convex. Posterior elevation of mesoventrite broadly and somewhat transversely elevated; mesoventrite with medial longitudinal glabrous patch extending anteriorly.
|
|||
|
<emphasis id="3A060FACDAA1719293512D45CDA2F668" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
|||
|
: Apical emargination of fifth ventrite relatively deep, U-shaped.
|
|||
|
<emphasis id="677A039D03DA9A8D70A89D400AC175BC" bold="true" italics="true" pageId="0" pageNumber="1">Aedeagus</emphasis>
|
|||
|
: (Fig.
|
|||
|
<figureCitation id="BA0F0CA6653621975A1F40338BEDD189" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Aedeagi of Novochares spp. A-C N. aperito D-F N. orchis G-K N. minor G-J Peru K Venezuela A, D, G, K dorsal view B, E, H ventral view C, F, J oblique view I lateral view." figureDoi="10.3897/zookeys.1171.104142.figure11" httpUri="https://binary.pensoft.net/fig/881178" pageId="0" pageNumber="1">11G-K</figureCitation>
|
|||
|
) Overall shape pear-like, joint basal margins of parameres somewhat truncate; apical region of each paramere sinuate and laterally pointed; parameres longer than median lobe, with apex rounded; dorsal inner margin of each paramere sinuate; dorsal plate of median lobe (in dorsal view) with broad neck and broad arms; gonopore placed at base of dorsal plate of median lobe; ventral plate of median lobe (in ventral view) membranous and short; basal piece nearly 0.3
|
|||
|
<normalizedToken id="0F1B42580724221F09D557FCE98A3F91" originalValue="×">x</normalizedToken>
|
|||
|
length of a paramere. In lateral view, aedeagus somewhat triangular, oblique at base, with ventral outline of parameres nearly 3.5
|
|||
|
<normalizedToken id="AB70C370B60AC721DF97725AC4153C7E" originalValue="×">x</normalizedToken>
|
|||
|
longer than greatest width near base.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="2437A9D928C575EF4266BC138C3FAC1A" pageId="0" pageNumber="1">
|
|||
|
<emphasis id="B553F2AB3EB5200BC73E817BE22E754B" bold="true" pageId="0" pageNumber="1">Composition.</emphasis>
|
|||
|
This group is composed of a single known species from Peru and Venezuela:
|
|||
|
<taxonomicName id="E7A52E92BE56A251E71CD3A1657F0977" lsidName="N. minor" pageId="0" pageNumber="1" rank="species" species="minor">
|
|||
|
<emphasis id="FC68588EFBA4F460924B05D0940FADEB" italics="true" pageId="0" pageNumber="1">N. minor</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
sp. nov.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="1AB6A9E35039E2D017EFF879CDD059ED" pageId="0" pageNumber="1">
|
|||
|
<emphasis id="4D6B6332EDDCB2F61C987A3F9191FBF4" bold="true" pageId="0" pageNumber="1">Remarks.</emphasis>
|
|||
|
The very small size combined with very dark to nearly black dorsal coloration serves to separate this species from most others. Most very dark species are typically much larger.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|