<documentID-CLB-Dataset="54411"ID-DOI="http://dx.doi.org/10.3897/zookeys.1149.96233"ID-GBIF-Dataset="8dae803a-15e9-48ff-94cb-9e91de3777f1"ID-PMC="PMC10208361"ID-Pensoft-Pub="1313-2970-1149-85"ID-Pensoft-UUID="63593CB973C35658876CC7F329EB2BFF"ID-PubMed="37234448"ID-ZooBank="6F1DC883058A4137A554E374F901D2E4"ModsDocID="1313-2970-1149-85"checkinTime="1677089976593"checkinUser="pensoft"docAuthor="Su, Yo, Lin, Hsiu-Chin & Ho, Hsuan-Ching"docDate="2023"docId="29C5D38B6FFC5E749D65A52D6B123B54"docLanguage="en"docName="ZooKeys 1149: 85-101"docOrigin="ZooKeys 1149"docPubDate="2023-02-22"docSource="http://dx.doi.org/10.3897/zookeys.1149.96233"docTitle="Hoplostethus crassispinus Kotlyar 1980"docType="treatment"docVersion="5"id="63593CB973C35658876CC7F329EB2BFF"lastPageNumber="85"masterDocId="63593CB973C35658876CC7F329EB2BFF"masterDocTitle="New records of two roughy fish species of Hoplostethus and a confirmed record of H. crassispinus Kotlyar, 1980 (Trachichthyiformes, Trachichthyidae) from Taiwan"masterLastPageNumber="101"masterPageNumber="85"pageNumber="85"updateTime="1732801146578"updateUser="ExternalLinkService">
<mods:titleid="9BD0BA7E73B4F2E90D61AC3606004A67">New records of two roughy fish species of Hoplostethus and a confirmed record of H. crassispinus Kotlyar, 1980 (Trachichthyiformes, Trachichthyidae) from Taiwan</mods:title>
<mods:affiliationid="FF43830BBF325E7EF3756ED6AC5BA920">Department of Marine Biotechnology and Resources, National Sun Yat-sen University, Kaohsiung, Taiwan</mods:affiliation>
<mods:affiliationid="AC8827668E60831AB5CFD26724F2C04E">Department of Marine Biotechnology and Resources, National Sun Yat-sen University, Kaohsiung, Taiwan</mods:affiliation>
<mods:affiliationid="0343A99456CB9094AA2B6E5586936744">Doctoral Degree Program in Marine Biotechnology, National Sun Yat-sen University, Kaohsiung, Taiwan & National Museum of Marine Biology & Aquarium, Pingtung, Taiwan & Australian Museum, Sydney, Australia</mods:affiliation>
<figureCitationid="D5BB39B73A438EEB603984D10CEB0889"captionStart="Figure 7"captionStartId="F7"captionText="Figure 7. Hoplostethus crassispinus, ASIZP 0065017, 86.3 mm SL A relative position of the eye and the horizontal level of lateral-line origin (white dashed line) B coloration of the oral cavity. Images not to scale."figureDoi="10.3897/zookeys.1149.96233.figure7"httpUri="https://binary.pensoft.net/fig/814230"pageId="0"pageNumber="85">, 7</figureCitation>
<bibRefCitationid="BD1DBFF38B4F215042806F9071C783F2"author="Kotlyar, AN"journalOrPublisher="Shirshova"pageId="0"pageNumber="85"pagination="97 - 140"refId="B7"refString="Kotlyar, AN, 1986. Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P.P. Shirshova 121: 97 - 140"title="Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P. P."volume="121"year="1986">Kotlyar 1986</bibRefCitation>
<bibRefCitationid="65EB466279F172B2D128EB484B497661"author="Kotlyar, AN"journalOrPublisher="VNIRO Publishing, Moscow"pageId="0"pageNumber="85"refId="B8"refString="Kotlyar, AN, 1996. Beryciform Fishes of the World Ocean. VNIRO Publishing, Moscow"title="Beryciform Fishes of the World Ocean."year="1996">Kotlyar 1996</bibRefCitation>
<bibRefCitationid="F0F460D76C5453614E45B347355D9DDB"DOI="https://doi.org/10.3374/014.051.0103"author="Moore, JA"journalOrPublisher="Bulletin - Peabody Museum of Natural History"pageId="0"pageNumber="85"pagination="137 - 144"refId="B10"refString="Moore, JA, Dodd, KA, 2010. A new species of the roughy genus Hoplostethus (Teleostei: Trachichthyidae) from the Philippines. Bulletin - Peabody Museum of Natural History 51 (1): 137 - 144, DOI: https://doi.org/10.3374/014.051.0103"title="A new species of the roughy genus Hoplostethus (Teleostei: Trachichthyidae) from the Philippines."url="https://doi.org/10.3374/014.051.0103"volume="51"year="2010">Moore and Dodd 2010</bibRefCitation>
<bibRefCitationid="4ECBA3EB44C2CB1E10453C3E70015C71"DOI="https://doi.org/10.1134/S0032945211040096"author="Kotlyar, AN"journalOrPublisher="Journal of Ichthyology"pageId="0"pageNumber="85"pagination="484 - 486"refId="B9"refString="Kotlyar, AN, 2011. Hoplostethus robustispinus (Trachichthyidae) from the South China Sea. Journal of Ichthyology 51 (6): 484 - 486, DOI: https://doi.org/10.1134/S0032945211040096"title="Hoplostethus robustispinus (Trachichthyidae) from the South China Sea."url="https://doi.org/10.1134/S0032945211040096"volume="51"year="2011">Kotlyar 2011</bibRefCitation>
<bibRefCitationid="850311514EE0BBBF8D1B0EFA94F65B63"DOI="https://doi.org/10.24199/j.mmv.2012.69.08"author="Roberts, CD"journalOrPublisher="Memoirs of the Museum of Victoria"pageId="0"pageNumber="85"pagination="341 - 354"refId="B12"refString="Roberts, CD, Gomon, MF, 2012. A review of giant roughies of the genus Hoplostethus (Beryciformes, Trachichthyidae), with descriptions of two new Australasian species. Memoirs of the Museum of Victoria 69: 341 - 354, DOI: https://doi.org/10.24199/j.mmv.2012.69.08"title="A review of giant roughies of the genus Hoplostethus (Beryciformes, Trachichthyidae), with descriptions of two new Australasian species."url="https://doi.org/10.24199/j.mmv.2012.69.08"volume="69"year="2012">Roberts and Gomon 2012</bibRefCitation>
<bibRefCitationid="9BDD3A129251882469D5941AD937BEAF"DOI="https://doi.org/10.1111/jfb.15086"author="Su, Y"journalOrPublisher="Journal of Fish Biology"pageId="0"pageNumber="85"pagination="441 - 452"refId="B15"refString="Su, Y, Lin, H-C, Ho, H-C, 2022. Hoplostethus roseus, a new roughy fish from the western Pacific based on morphology and DNA barcoding (family Trachichthyidae). Journal of Fish Biology 101 (3): 441 - 452, DOI: https://doi.org/10.1111/jfb.15086"title="Hoplostethus roseus, a new roughy fish from the western Pacific based on morphology and DNA barcoding (family Trachichthyidae)."url="https://doi.org/10.1111/jfb.15086"volume="101"year="2022">Su et al. 2022</bibRefCitation>
<bibRefCitationid="EE99F6451C64FAECB940E76E83516ADE"DOI="https://doi.org/10.3390/jmse9111294"author="Koeda, K"journalOrPublisher="National Marine Museum of Biology and Aquarium, Checheng"pageId="0"pageNumber="85"refId="B5"refString="Koeda, K, Takashima, S, Yamakita, T, Tsuchida, S, Fujiwara, Y, 2021. Deep-sea fish fauna on the seamounts of southern Japan with taxonomic notes on the observed species. Journal of Marine Science and Engineering 9(11): e1294. https://doi.org/10.3390/jmse9111294"title="Deep-sea fish fauna on the seamounts of southern Japan with taxonomic notes on the observed species. Journal of Marine Science and Engineering 9 (11): e 1294."url="https://doi.org/10.3390/jmse9111294"year="2021">Koeda et al. 2021</bibRefCitation>
<tableCitationid="AE336B89FDB7B2662CE06250640C6449"captionStart="Table 3"captionStartId="T3"captionText="Table 3. Meristic data of Hoplostehus crassispinus. Data of type and other specimens were obtained from Kotlyar (1980) and Kotlyar (1986). Paired-fin characters are presented as left / right."httpUri="http://table.plazi.org/id/6049F671DBF26DE7817419FCC69B562F"pageId="0"pageNumber="85"tableUuid="6049F671DBF26DE7817419FCC69B562F">3</tableCitation>
<tableCitationid="836B71B1BEDC311C0619692DCE6B231F"captionStart="Table 4"captionStartId="T4"captionText="Table 4. Morphometric data of Hoplostethus crassispinus. Data of type and other specimens were obtained from Kotlyar (1980) and Kotlyar (1986). Abbreviations: A, anal-fin; C, caudal-fin; D, dorsal-fin; GR, gill raker; HF, forehead height; HL, head length; HT, holotype; NT, non-type; P, pectoral-fin; V, pelvic-fin."httpUri="http://table.plazi.org/id/285CB1B83F5162335695F6E8BA2140E4"pageId="0"pageNumber="85"tableUuid="285CB1B83F5162335695F6E8BA2140E4">4</tableCitation>
<bibRefCitationid="1C790E6E7B2303CF420D67DC5B336306"author="Kotlyar, AN"journalOrPublisher="Zoologicheskii Zhurnal [Zoologicheskii zhurnal]"pageId="0"pageNumber="85"pagination="1054 - 1059"refId="B6"refString="Kotlyar, AN, 1980. A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific. Zoologicheskii Zhurnal [Zoologicheskii zhurnal] 59 (7): 1054 - 1059"title="A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific."volume="59"year="1980">Kotlyar (1980)</bibRefCitation>
<bibRefCitationid="5A734C1AB7306DDDDD405FA9A16792F2"author="Kotlyar, AN"journalOrPublisher="Shirshova"pageId="0"pageNumber="85"pagination="97 - 140"refId="B7"refString="Kotlyar, AN, 1986. Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P.P. Shirshova 121: 97 - 140"title="Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P. P."volume="121"year="1986">Kotlyar (1986)</bibRefCitation>
<bibRefCitationid="2347AA476F1D812E365951A5E648B5AE"author="Kotlyar, AN"journalOrPublisher="Zoologicheskii Zhurnal [Zoologicheskii zhurnal]"pageId="0"pageNumber="85"pagination="1054 - 1059"refId="B6"refString="Kotlyar, AN, 1980. A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific. Zoologicheskii Zhurnal [Zoologicheskii zhurnal] 59 (7): 1054 - 1059"title="A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific."volume="59"year="1980">Kotlyar (1980)</bibRefCitation>
<bibRefCitationid="614503C7E41C51FD2C1FF520655B0BF8"author="Kotlyar, AN"journalOrPublisher="Shirshova"pageId="0"pageNumber="85"pagination="97 - 140"refId="B7"refString="Kotlyar, AN, 1986. Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P.P. Shirshova 121: 97 - 140"title="Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P. P."volume="121"year="1986">Kotlyar (1986)</bibRefCitation>
<bibRefCitationid="6265BD8826D79DF42AD31C3407B61C79"author="Kotlyar, AN"journalOrPublisher="Zoologicheskii Zhurnal [Zoologicheskii zhurnal]"pageId="0"pageNumber="85"pagination="1054 - 1059"refId="B6"refString="Kotlyar, AN, 1980. A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific. Zoologicheskii Zhurnal [Zoologicheskii zhurnal] 59 (7): 1054 - 1059"title="A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific."volume="59"year="1980">Kotlyar (1980)</bibRefCitation>
<bibRefCitationid="144CD81594E08C968AD8C480619A7822"author="Kotlyar, AN"journalOrPublisher="Shirshova"pageId="0"pageNumber="85"pagination="97 - 140"refId="B7"refString="Kotlyar, AN, 1986. Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P.P. Shirshova 121: 97 - 140"title="Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P. P."volume="121"year="1986">Kotlyar (1986)</bibRefCitation>
<bibRefCitationid="8E464589B6D42668C3D1C05663850AFD"author="Kotlyar, AN"journalOrPublisher="Zoologicheskii Zhurnal [Zoologicheskii zhurnal]"pageId="0"pageNumber="85"pagination="1054 - 1059"refId="B6"refString="Kotlyar, AN, 1980. A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific. Zoologicheskii Zhurnal [Zoologicheskii zhurnal] 59 (7): 1054 - 1059"title="A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific."volume="59"year="1980">Kotlyar (1980)</bibRefCitation>
<bibRefCitationid="4F9715AFD1725E77C7E48B3391F33B30"author="Kotlyar, AN"journalOrPublisher="Shirshova"pageId="0"pageNumber="85"pagination="97 - 140"refId="B7"refString="Kotlyar, AN, 1986. Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P.P. Shirshova 121: 97 - 140"title="Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P. P."volume="121"year="1986">Kotlyar (1986)</bibRefCitation>
<paragraphid="B0B782C2D9FCC8F355D1AFE495B3C174"pageId="0"pageNumber="85">Dorsal-fin rays VI, 13; pectoral-fin rays 17/15; pelvic-fin rays I, 6/I, 6; anal-fin rays III, 9; principal caudal-fin rays 10+9, uppermost and lowermost rays unbranched; procurrent caudal-fin rays 7 dorsally and 7 ventrally; gill rakers on outer surface of first gill arch 5+1+12=18; lateral-line scales 28; predorsal scales 23; abdominal scutes 15; vertebrae 11+16=27; pseudobranchial filaments 24; branchiostegal rays 8; supraneural and pterygiophore insertion formula: 0/0/2+1/1/1/1 (spinous dorsal fin only).</paragraph>
<paragraphid="6DFE8D58D62B1FC08F04AD6AD11147B9"pageId="0"pageNumber="85">Body oblong, distinctly longer than deep, depth at dorsal-fin origin 1.9 in SL. Head large, its length 2.7% in SL, its height slightly smaller than its length, 0.9% in HL; upper profile in front of dorsal fin rounded, slightly curved to back of head, with somewhat rounded forehead, and abrupt downturn above maxilla; forehead broad, HF1 9.4 and HF2 6.0 in HL; eyes rather large, 2.9 in HL; snout length 5.2 in HL; space between eyes convex and broad, interorbital width 3.3 in HL; crests on head bones well developed and covered with very tiny spinules.</paragraph>
Mouth large, posterior end of maxilla slightly reaching beyond vertical through posterior margin of eye. Nostrils right before anterior margin of eye, slightly higher than horizontal through center of eye; posterior nostril distinctly larger than anterior nostril; eyes rather ventrally placed, upper margin of eye distinctly lower than horizontal through lateral-line origin (Fig.
<figureCitationid="85025F4E7E981DBA5E975DCBEE77E628"captionStart="Figure 7"captionStartId="F7"captionText="Figure 7. Hoplostethus crassispinus, ASIZP 0065017, 86.3 mm SL A relative position of the eye and the horizontal level of lateral-line origin (white dashed line) B coloration of the oral cavity. Images not to scale."figureDoi="10.3897/zookeys.1149.96233.figure7"httpUri="https://binary.pensoft.net/fig/814230"pageId="0"pageNumber="85">7A</figureCitation>
<paragraphid="BED5A7FDDB96244B698920B099074E72"pageId="0"pageNumber="85">Most of lateral and medial surfaces of premaxilla and dentary covered with villiform teeth; no teeth at symphyseal notch of premaxilla and knob at symphysis of dentaries. Narrow band of villiform teeth on palatine; vomer toothless. Gill rakers on first and second arch rod-shaped, laterally compressed; those in outer row of first arch longest; those on inner row of first arch and both inner and outer rows of second to fourth arches short.</paragraph>
<paragraphid="D7AECB37602CADF750491FCBF4D19834"pageId="0"pageNumber="85">Preopercular spine rather long, its tip reaching pelvic-fin base. Longest gill raker 1.4 in eye diameter; gill filaments at angle of first gill arch very short, ca 6.2 in eye diameter, and ca 1.9 in length of longest pseudobranchial filaments.</paragraph>
<paragraphid="900A510B58F6289996A142A9D147A117"pageId="0"pageNumber="85">Body scales firmly attached, cycloid scales present on pectoral-fin region, elsewhere covered with ctenoid scales; isthmus and gular region naked; lateral-line scales enlarged, ca 2-3 times size of body scales; center of each lateral-line scale without distinct spine; enlarged scales (scutes) covering abdomen region, their bases covered with body scales, all scutes with single tip; predorsal scales enlarged and forming ridge.</paragraph>
<paragraphid="6C7CCE386679C6E6FC2E5970204F519D"pageId="0"pageNumber="85">Dorsal-fin spines progressively longer posteriorly, greatest increase in length from first to third spine; first ray unbranched, others branched; outer margin of dorsal-fin rays nearly straight. Pectoral fin truncated, slightly rounded; rather long, reaching to third anal-fin spine. Pelvic fin rather long, reaching 14th abdominal scute.</paragraph>
); all fin spines paler than body color. Membranes between dorsal-fin spines with black pigmentations. Oral cavity, including underside of tongue mostly black, with some portions slightly whitish (Fig.
<figureCitationid="CF1432C550F56BBCAB18DC5BF179363C"captionStart="Figure 7"captionStartId="F7"captionText="Figure 7. Hoplostethus crassispinus, ASIZP 0065017, 86.3 mm SL A relative position of the eye and the horizontal level of lateral-line origin (white dashed line) B coloration of the oral cavity. Images not to scale."figureDoi="10.3897/zookeys.1149.96233.figure7"httpUri="https://binary.pensoft.net/fig/814230"pageId="0"pageNumber="85">7B</figureCitation>
<bibRefCitationid="4BD93C07CCAF2CCC299976FE78A821F6"DOI="https://doi.org/10.3390/jmse9111294"author="Koeda, K"journalOrPublisher="National Marine Museum of Biology and Aquarium, Checheng"pageId="0"pageNumber="85"refId="B5"refString="Koeda, K, Takashima, S, Yamakita, T, Tsuchida, S, Fujiwara, Y, 2021. Deep-sea fish fauna on the seamounts of southern Japan with taxonomic notes on the observed species. Journal of Marine Science and Engineering 9(11): e1294. https://doi.org/10.3390/jmse9111294"title="Deep-sea fish fauna on the seamounts of southern Japan with taxonomic notes on the observed species. Journal of Marine Science and Engineering 9 (11): e 1294."url="https://doi.org/10.3390/jmse9111294"year="2021">Koeda et al. 2021</bibRefCitation>
<bibRefCitationid="F046AC564A5BF522367914A8F5552505"author="Kotlyar, AN"journalOrPublisher="Zoologicheskii Zhurnal [Zoologicheskii zhurnal]"pageId="0"pageNumber="85"pagination="1054 - 1059"refId="B6"refString="Kotlyar, AN, 1980. A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific. Zoologicheskii Zhurnal [Zoologicheskii zhurnal] 59 (7): 1054 - 1059"title="A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific."volume="59"year="1980">Kotlyar 1980</bibRefCitation>
<bibRefCitationid="CE01B340AD567C6941D57F168D2437FE"author="Kotlyar, AN"journalOrPublisher="Shirshova"pageId="0"pageNumber="85"pagination="97 - 140"refId="B7"refString="Kotlyar, AN, 1986. Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P.P. Shirshova 121: 97 - 140"title="Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P. P."volume="121"year="1986">Kotlyar 1986</bibRefCitation>
<bibRefCitationid="A43E8B10500BDE38F321CFC36089F3D9"DOI="https://doi.org/10.1134/S0032945211040096"author="Kotlyar, AN"journalOrPublisher="Journal of Ichthyology"pageId="0"pageNumber="85"pagination="484 - 486"refId="B9"refString="Kotlyar, AN, 2011. Hoplostethus robustispinus (Trachichthyidae) from the South China Sea. Journal of Ichthyology 51 (6): 484 - 486, DOI: https://doi.org/10.1134/S0032945211040096"title="Hoplostethus robustispinus (Trachichthyidae) from the South China Sea."url="https://doi.org/10.1134/S0032945211040096"volume="51"year="2011">Kotlyar (2011)</bibRefCitation>
by having a lower eye position, with the upper margin of the eye distinctly below the horizontal through lateral-line origin, a moderately long pectoral fin, with its end slightly exceeding the vertical through anal-fin origin, a blackish oral cavity, including the underside of the tongue, 15 or 17 pectoral-fin rays, and 18 total gill rakers. It can be distinguished from other Taiwanese species by its ventrally positioned eye, the upper margin of which is clearly below the horizontal through lateral-line origin (vs a more dorsally placed eye, the upper margin of the eye at the same horizontal through lateral-line origin in all species in Taiwan).
<bibRefCitationid="E88AC78A3505B21BC01D82F3FA14E243"author="Shen, S-C [Ed.]"journalOrPublisher="Department of Zoology, National Taiwan University, Taipei"pageId="0"pageNumber="85"refId="B13"refString="Shen, S-C [Ed.], Chen, C-H, Lee, S-C, Shao, K-T, Mok, H-K, Tseng, C-S, 1993. Fishes of Taiwan. Department of Zoology, National Taiwan University, Taipei"title="Fishes of Taiwan."year="1993">Shen et al. 1993</bibRefCitation>
<bibRefCitationid="1376914CCF36914449FB363BE37A6FD1"author="Shen, S-C"journalOrPublisher="National Museum of Marine Biology and Aquarium, Pingtung"pageId="0"pageNumber="85"refId="B14"refString="Shen, S-C, Wu, K-Y, 2011. Fishes of Taiwan. National Museum of Marine Biology and Aquarium, Pingtung"title="Fishes of Taiwan."year="2011">Shen and Wu 2011</bibRefCitation>
<bibRefCitationid="1B4746923FFA86B93CF9C2770880F4DF"author="Koeda, K"journalOrPublisher="National Marine Museum of Biology and Aquarium, Checheng"pageId="0"pageNumber="85"refId="B4"refString="Koeda, K, 2019. Family Trachichthyidae. In: Koeda K, Ho H-C (Eds) Fishes of Southern Taiwan. National Marine Museum of Biology and Aquarium, Checheng"title="Family Trachichthyidae. In: Koeda K, Ho H-C (Eds) Fishes of Southern Taiwan."year="2019">Koeda 2019</bibRefCitation>
<bibRefCitationid="870D28C8E36E3CBF95D2C69EE3A07D15"DOI="https://doi.org/10.1111/jfb.15086"author="Su, Y"journalOrPublisher="Journal of Fish Biology"pageId="0"pageNumber="85"pagination="441 - 452"refId="B15"refString="Su, Y, Lin, H-C, Ho, H-C, 2022. Hoplostethus roseus, a new roughy fish from the western Pacific based on morphology and DNA barcoding (family Trachichthyidae). Journal of Fish Biology 101 (3): 441 - 452, DOI: https://doi.org/10.1111/jfb.15086"title="Hoplostethus roseus, a new roughy fish from the western Pacific based on morphology and DNA barcoding (family Trachichthyidae)."url="https://doi.org/10.1111/jfb.15086"volume="101"year="2022">Su et al. 2022</bibRefCitation>
sp.); predorsal scales 21-23 (vs 15-19); pyloric caeca 40-50 (vs 36); oral cavity blackish in adults (vs oral cavity whitish in adults); upper margin of the eye clearly below the horizontal through lateral-line origin (vs at the same horizontal level); gular region naked (vs gular region covered with ctenoid scales). As mentioned previously, the taxonomic study by Dr M. Gomon is ongoing, and we will await the publication of his findings (
<bibRefCitationid="0E5CD29A7CF97DF01D744F7228A9D43E"DOI="https://doi.org/10.1111/jfb.15086"author="Su, Y"journalOrPublisher="Journal of Fish Biology"pageId="0"pageNumber="85"pagination="441 - 452"refId="B15"refString="Su, Y, Lin, H-C, Ho, H-C, 2022. Hoplostethus roseus, a new roughy fish from the western Pacific based on morphology and DNA barcoding (family Trachichthyidae). Journal of Fish Biology 101 (3): 441 - 452, DOI: https://doi.org/10.1111/jfb.15086"title="Hoplostethus roseus, a new roughy fish from the western Pacific based on morphology and DNA barcoding (family Trachichthyidae)."url="https://doi.org/10.1111/jfb.15086"volume="101"year="2022">Su et al. 2022</bibRefCitation>
<bibRefCitationid="C2965B9F8A0C7D72A9F85DB08210F9BD"author="Kotlyar, AN"journalOrPublisher="Zoologicheskii Zhurnal [Zoologicheskii zhurnal]"pageId="0"pageNumber="85"pagination="1054 - 1059"refId="B6"refString="Kotlyar, AN, 1980. A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific. Zoologicheskii Zhurnal [Zoologicheskii zhurnal] 59 (7): 1054 - 1059"title="A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific."volume="59"year="1980">Kotlyar (1980</bibRefCitation>
<bibRefCitationid="6A1D750C40488588647DF1B11833CA9C"author="Kotlyar, AN"journalOrPublisher="Shirshova"pageId="0"pageNumber="85"pagination="97 - 140"refId="B7"refString="Kotlyar, AN, 1986. Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P.P. Shirshova 121: 97 - 140"title="Systematics and distribution of species of the genus Hoplostethus Cuvier (Beryciformes, Trachichthyidae). Trudy Instituta Okeanologii Imeni P. P."volume="121"year="1986">1986</bibRefCitation>
), our specimen has very slight variations. Due to the fact that all other specimens are significantly larger than ours (135-254 vs 86.3 mm SL), these differences can be attributed to intraspecific variation. We discovered that an additional 136 mm SL specimen was used in
<bibRefCitationid="7E0601CB6C504CECB27A2505A1A0BFAE"author="Kotlyar, AN"journalOrPublisher="Zoologicheskii Zhurnal [Zoologicheskii zhurnal]"pageId="0"pageNumber="85"pagination="1054 - 1059"refId="B6"refString="Kotlyar, AN, 1980. A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific. Zoologicheskii Zhurnal [Zoologicheskii zhurnal] 59 (7): 1054 - 1059"title="A new species of the genus Hoplostethus (Trachichthyidae, Beryciformes) from the north-west Pacific."volume="59"year="1980">
