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<document id="FD66801D29B85AA72BF91E9EC4673A55" ID-DOI="10.11646/zootaxa.4759.3.3" ID-GBIF-Dataset="9675f3d4-930d-4e4c-97e1-bcc7e6c5120d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3741178" ID-ZooBank="FBD53C14-07AA-4E01-9739-214680A1DE4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586156589363" checkinUser="plazi" docAuthor="Skartveit, John &amp; Krizmanić, Katarina" docDate="2020" docId="C6550F6A950CFF9C23B1FDABFA8FE44A" docLanguage="en" docName="zootaxa.4759.3.3.pdf" docOrigin="Zootaxa 4759 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dilophus pinguis Skartveit &amp; Krizmanić 2020, comb.n." docType="treatment" docVersion="5" lastPageNumber="369" masterDocId="3A6C7712951CFF8E2326FFE2FF8FE07D" masterDocTitle="Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia" masterLastPageNumber="378" masterPageNumber="351" pageNumber="367" updateTime="1698735747428" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="44B006211954828FE35159570ADD1326">Revision of fossil Bibionidae (Insecta: Diptera) from the Miocene of Radoboj, Croatia</mods:title>
</mods:titleInfo>
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<mods:namePart id="C7835B318EF45B423091B46ABBE7078F">Skartveit, John</mods:namePart>
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<mods:namePart id="F0A0F5404CADB75AC814B13A98D9837B">Krizmanić, Katarina</mods:namePart>
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<mods:title id="BA6C9519E525BEDA2C681DCD7AF5AC08">Zootaxa</mods:title>
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<mods:date id="C093BADB53DBD0452954C47D645CF733">2020</mods:date>
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<mods:number id="015CC7CF8C6DA522CB9DBFB967543005">2020-04-03</mods:number>
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<mods:detail id="8BC0386536206F6D6BF872BA05C6AA4A" type="volume">
<mods:number id="C0AE9D7E48867C6E616880A575C4170D">4759</mods:number>
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<mods:number id="035562DB7E7CE6E9CB4F3062A2FB3305">3</mods:number>
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<mods:classification id="DC1FB9432AA96929A1B1023DDBFD2970">journal article</mods:classification>
<mods:identifier id="4FBF4DE403F6F9A62CBBE8F0C52BDFCD" type="DOI">10.11646/zootaxa.4759.3.3</mods:identifier>
<mods:identifier id="006C159EC8168CF7FBD4022FA507A5BF" type="GBIF-Dataset">9675f3d4-930d-4e4c-97e1-bcc7e6c5120d</mods:identifier>
<mods:identifier id="E9AA07A3474C3C1DAB25EEA4B8D3B446" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7EC809B9A425F914FB94AF11E78B2596" type="Zenodo-Dep">3741178</mods:identifier>
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<treatment id="C6550F6A950CFF9C23B1FDABFA8FE44A" ID-DOI="http://doi.org/10.5281/zenodo.3810445" ID-GBIF-Taxon="163437714" ID-Zenodo-Dep="3810445" LSID="urn:lsid:plazi:treatment:C6550F6A950CFF9C23B1FDABFA8FE44A" httpUri="http://treatment.plazi.org/id/C6550F6A950CFF9C23B1FDABFA8FE44A" lastPageId="18" lastPageNumber="369" pageId="16" pageNumber="367">
<subSubSection id="06E6EDF7950CFF9E23B1FDABFDFEE21E" box="[151,625,585,612]" pageId="16" pageNumber="367" type="nomenclature">
<paragraph id="4E43BE7C950CFF9E23B1FDABFDFEE21E" blockId="16.[151,625,585,647]" box="[151,625,585,612]" pageId="16" pageNumber="367">
<heading id="150B0910950CFF9E23B1FDABFDFEE21E" bold="true" box="[151,625,585,612]" fontSize="11" level="1" pageId="16" pageNumber="367" reason="1">
<emphasis id="7C88626E950CFF9E23B1FDABFDFEE21E" bold="true" box="[151,625,585,612]" pageId="16" pageNumber="367">
<taxonomicName id="89FCC5FF950CFF9E23B1FDABFD89E21E" ID-CoL="9HLVH" authority="(Heer, 1849)" authorityName="Skartveit &amp; Krizmanić" authorityYear="2020" baseAuthorityName="Heer" baseAuthorityYear="1849" box="[151,518,585,612]" class="Insecta" family="Bibionidae" genus="Dilophus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="pinguis" status="comb. nov.">
<emphasis id="7C88626E950CFF9E23B1FDABFEE8E21E" bold="true" box="[151,359,585,612]" italics="true" pageId="16" pageNumber="367">Dilophus pinguis</emphasis>
(
<bibRefCitation id="2A6DC38D950CFF9E2250FDABFE71E21E" author="Heer, O." box="[374,510,585,611]" pageId="16" pageNumber="367" refId="ref11225" refString="Heer, O. (1849) Die Insektenfauna der Tertiargebilde von Oeningen und von Radoboj in Croatien. Zweite Theil: Heuschrecken, Florfliegen, Aderflugler, Schmetterlinge und Fliegen. W. Engelmann, Leipzig, 264 + XVII pp." type="book" year="1849">Heer, 1849</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="67BBDF15950CFF9E2128FDABFDFEE21E" box="[526,625,585,611]" pageId="16" pageNumber="367" rank="species">comb.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="06E6EDF7950CFF9E23B1FD8FFE0DE2FA" box="[151,386,620,647]" pageId="16" pageNumber="367" type="description">
<paragraph id="4E43BE7C950CFF9E23B1FD8FFE0DE2FA" blockId="16.[151,625,585,647]" box="[151,386,620,647]" pageId="16" pageNumber="367">
(
<figureCitation id="D6C7A2F9950CFF9E23B9FD8FFEA8E2FA" box="[159,295,620,647]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStartId-0="16.[151,250,2012,2037]" captionStartId-1="17.[151,250,1073,1098]" captionStartId-2="17.[151,250,2011,2036]" captionTargetBox-0="[151,1436,1023,1987]" captionTargetBox-1="[216,1372,181,1048]" captionTargetBox-2="[217,1371,1120,1985]" captionTargetId-0="figure@16.[151,1436,1023,1987]" captionTargetId-1="figure@17.[216,1372,181,1048]" captionTargetId-2="figure@17.[217,1371,1120,1985]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="17" captionText-0="FIGURE 15. Dilophus pinguis, male, CNHM 1832." captionText-1="FIGURE 16. Dilophus pinguis, female, lectotype, SLJG 77473." captionText-2="FIGURE 17. Dilophus pinguis, female, paralectotype, NHMW 1842/0050/0022." figureDoi-0="http://doi.org/10.5281/zenodo.3741206" figureDoi-1="http://doi.org/10.5281/zenodo.3741208" figureDoi-2="http://doi.org/10.5281/zenodo.3741210" httpUri-0="https://zenodo.org/record/3741206/files/figure.png" httpUri-1="https://zenodo.org/record/3741208/files/figure.png" httpUri-2="https://zenodo.org/record/3741210/files/figure.png" pageId="16" pageNumber="367">Figs. 15–17</figureCitation>
,
<figureCitation id="D6C7A2F9950CFF9E2215FD8FFEF4E2FA" box="[307,379,621,647]" captionStart-0="FIGURES 32–35. 32" captionStart-1="FIGURES 36–38. 36" captionStartId-0="24.[151,264,1940,1965]" captionStartId-1="25.[151,264,1945,1970]" captionTargetBox-0="[159,1432,190,1903]" captionTargetBox-1="[308,1282,193,1906]" captionTargetId-0="figure@24.[151,1436,181,1914]" captionTargetId-1="figure@25.[294,1294,181,1920]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURES 32–35. 32. Bibio giganteus, female, holotype, wing, SLJG 77605. 33. Bibio giganteus, female, holotype, head and thorax, SLJG 77605. 34. Dilophus pinguis, male, hind leg, CNHM 1832. 35. Dilophus pinguis, male, hypopygium, CNHM 1832. Scale bar = 1 mm." captionText-1="FIGURES 36–38. 36. Dilophus pinguis, female, lectotype, habitus, SLJG 77473. 37. Dilophus pinguis, female, lectotype, wing, SLJG 77473. 38. «Bibio ungeri», type, habitus with tentative wing venation, SLJG, no catalog number. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3741222" figureDoi-1="http://doi.org/10.5281/zenodo.3741224" httpUri-0="https://zenodo.org/record/3741222/files/figure.png" httpUri-1="https://zenodo.org/record/3741224/files/figure.png" pageId="16" pageNumber="367">34–37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="06E6EDF7950CFF9E23B1FD53FE45E2B4" box="[151,458,689,714]" pageId="16" pageNumber="367" type="reference_group">
<paragraph id="4E43BE7C950CFF9E23B1FD53FE45E2B4" blockId="16.[151,458,689,714]" box="[151,458,689,714]" pageId="16" pageNumber="367">
<treatmentCitationGroup id="6EEC9952950CFF9E23B1FD53FE45E2B4" box="[151,458,689,714]" pageId="16" pageNumber="367">
<treatmentCitation id="CF5D986D950CFF9E23B1FD53FE45E2B4" author="Heer, O." box="[151,458,689,714]" page="220" pageId="16" pageNumber="367" year="1849">
<taxonomicName id="89FCC5FF950CFF9E23B1FD53FE45E2B4" ID-CoL="8B75N" authority="Heer, 1849: 220" authorityName="Heer" authorityPageNumber="220" authorityYear="1849" box="[151,458,689,714]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="pinguis">
Bibio pinguis
<bibRefCitation id="2A6DC38D950CFF9E2201FD50FE45E2B4" author="Heer, O." box="[295,458,689,713]" pageId="16" pageNumber="367" refId="ref11225" refString="Heer, O. (1849) Die Insektenfauna der Tertiargebilde von Oeningen und von Radoboj in Croatien. Zweite Theil: Heuschrecken, Florfliegen, Aderflugler, Schmetterlinge und Fliegen. W. Engelmann, Leipzig, 264 + XVII pp." type="book" year="1849">Heer, 1849: 220</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="06E6EDF7950CFF9C23B1FD1AFB8FE0AA" lastPageId="18" lastPageNumber="369" pageId="16" pageNumber="367" type="materials_examined">
<paragraph id="4E43BE7C950CFF9E23B1FD1AFDD9E397" blockId="16.[151,1437,759,1002]" pageId="16" pageNumber="367">
<emphasis id="7C88626E950CFF9E23B1FD1AFEC8E36F" bold="true" box="[151,327,759,786]" pageId="16" pageNumber="367">Type material:</emphasis>
<typeStatus id="914700DE950CFF9E2268FD1AFE31E36F" box="[334,446,760,786]" pageId="16" pageNumber="367" type="lectotype">Lectotype</typeStatus>
(female, hereby designated) (
<collectionCode id="28ED26B9950CFF9E202DFD1AFCC2E36F" box="[779,845,760,786]" country="Austria" httpUri="http://grbio.org/cool/nap9-3men" name="Steiermarkisches Landesmuseum Joanneum" pageId="16" pageNumber="367">SLJG</collectionCode>
77473) this is the specimen depicted by
<bibRefCitation id="2A6DC38D950CFF9E2630FD1AFA16E36F" author="Heer, O." box="[1302,1433,760,786]" pageId="16" pageNumber="367" refId="ref11225" refString="Heer, O. (1849) Die Insektenfauna der Tertiargebilde von Oeningen und von Radoboj in Croatien. Zweite Theil: Heuschrecken, Florfliegen, Aderflugler, Schmetterlinge und Fliegen. W. Engelmann, Leipzig, 264 + XVII pp." type="book" year="1849">Heer (1849</bibRefCitation>
: Plate XVI,
<figureCitation id="D6C7A2F9950CFF9E2233FCFEFEFBE34B" box="[277,372,796,822]" captionStart="FIGURE 11" captionStartId="13.[151,250,1693,1718]" captionTargetBox="[172,1415,736,1667]" captionTargetId="figure@13.[172,1415,736,1667]" captionTargetPageId="13" captionText="FIGURE 11. Bibio firmus, female, holotype, SLJG 77471." figureDoi="http://doi.org/10.5281/zenodo.3741198" httpUri="https://zenodo.org/record/3741198/files/figure.png" pageId="16" pageNumber="367">Fig. 11a</figureCitation>
). The specimen has been labeled as a
<typeStatus id="914700DE950CFF9E203CFCFEFCFEE34B" box="[794,881,796,822]" pageId="16" pageNumber="367" type="syntype">syntype</typeStatus>
of
<taxonomicName id="89FCC5FF950CFF9E20B0FCFEFB06E34B" authority="Heer" authorityName="Heer" authorityYear="1849" box="[918,1161,796,822]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="7C88626E950CFF9E20B0FCFEFBC3E34B" box="[918,1100,796,822]" italics="true" pageId="16" pageNumber="367">Bibio maculatus</emphasis>
Heer
</taxonomicName>
but this is obviously not correct. The specimen was described as
<taxonomicName id="89FCC5FF950CFF9E2171FCA2FC0DE327" box="[599,898,832,858]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="subSpecies" species="pinguis" subSpecies="radobojanus">
<emphasis id="7C88626E950CFF9E2171FCA2FC0DE327" box="[599,898,832,858]" italics="true" pageId="16" pageNumber="367">Bibio pinguis radobojanus</emphasis>
</taxonomicName>
by Heer. Specimen
<collectionCode id="28ED26B9950CFF9E2745FCA2FB2CE327" box="[1123,1187,832,858]" country="Austria" httpUri="http://grbio.org/cool/nap9-3men" name="Steiermarkisches Landesmuseum Joanneum" pageId="16" pageNumber="367">SLJG</collectionCode>
77475 is probably the counterpart of
<collectionCode id="28ED26B9950CFF9E221DFC86FEF4E303" box="[315,379,868,894]" country="Austria" httpUri="http://grbio.org/cool/nap9-3men" name="Steiermarkisches Landesmuseum Joanneum" pageId="16" pageNumber="367">SLJG</collectionCode>
77473.
<typeStatus id="914700DE950CFF9E22F2FC86FDE3E303" box="[468,620,868,894]" pageId="16" pageNumber="367" type="paralectotype">Paralectotype</typeStatus>
(female, hereby designated) (
<collectionCode id="28ED26B9950CFF9E2091FC86FB9BE303" box="[951,1044,868,894]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="367" type="Museum">NHMW</collectionCode>
1842/0050/0022), this is the specimen described as «
<taxonomicName id="89FCC5FF950CFF9E2248FC6AFE6FE3DF" box="[366,480,904,930]" pageId="16" pageNumber="367" rank="variety" variety="paulo">
var.
<emphasis id="7C88626E950CFF9E2286FC6AFE6FE3DF" box="[416,480,904,930]" italics="true" pageId="16" pageNumber="367">paulo</emphasis>
</taxonomicName>
<emphasis id="7C88626E950CFF9E22C1FC6AFDA5E3DC" box="[487,554,904,929]" italics="true" pageId="16" pageNumber="367">minor</emphasis>
» and depicted by
<bibRefCitation id="2A6DC38D950CFF9E21DCFC6AFCF2E3DF" author="Heer, O." box="[762,893,904,930]" pageId="16" pageNumber="367" refId="ref11225" refString="Heer, O. (1849) Die Insektenfauna der Tertiargebilde von Oeningen und von Radoboj in Croatien. Zweite Theil: Heuschrecken, Florfliegen, Aderflugler, Schmetterlinge und Fliegen. W. Engelmann, Leipzig, 264 + XVII pp." type="book" year="1849">Heer (1849</bibRefCitation>
: Plate XVI,
<figureCitation id="D6C7A2F9950CFF9E272FFC6AFBEBE3DF" box="[1033,1124,904,930]" captionStart="FIGURE 11" captionStartId="13.[151,250,1693,1718]" captionTargetBox="[172,1415,736,1667]" captionTargetId="figure@13.[172,1415,736,1667]" captionTargetPageId="13" captionText="FIGURE 11. Bibio firmus, female, holotype, SLJG 77471." figureDoi="http://doi.org/10.5281/zenodo.3741198" httpUri="https://zenodo.org/record/3741198/files/figure.png" pageId="16" pageNumber="367">Fig. 11c</figureCitation>
) We have not seen the type of
<taxonomicName id="89FCC5FF950CFF9E2390FC4EFE3DE3B8" box="[182,434,940,965]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="subSpecies" species="pinguis" subSpecies="oeningensis">
<emphasis id="7C88626E950CFF9E2390FC4EFE3DE3B8" box="[182,434,940,965]" italics="true" pageId="16" pageNumber="367">B. pinguis oeningensis</emphasis>
</taxonomicName>
; according to the original description (
<bibRefCitation id="2A6DC38D950CFF9E2040FC4EFB99E3BB" author="Heer, O." box="[870,1046,940,966]" pageId="16" pageNumber="367" refId="ref11225" refString="Heer, O. (1849) Die Insektenfauna der Tertiargebilde von Oeningen und von Radoboj in Croatien. Zweite Theil: Heuschrecken, Florfliegen, Aderflugler, Schmetterlinge und Fliegen. W. Engelmann, Leipzig, 264 + XVII pp." type="book" year="1849">Heer 1849: 221</bibRefCitation>
and Pl. XVI,
<figureCitation id="D6C7A2F9950CFF9E2797FC4EFA80E3BA" box="[1201,1295,940,967]" captionStart="FIGURE 11" captionStartId="13.[151,250,1693,1718]" captionTargetBox="[172,1415,736,1667]" captionTargetId="figure@13.[172,1415,736,1667]" captionTargetPageId="13" captionText="FIGURE 11. Bibio firmus, female, holotype, SLJG 77471." figureDoi="http://doi.org/10.5281/zenodo.3741198" httpUri="https://zenodo.org/record/3741198/files/figure.png" pageId="16" pageNumber="367">Fig. 11b</figureCitation>
) it would be quite similar to
<emphasis id="7C88626E950CFF9E226EFC32FDD9E397" box="[328,598,976,1002]" italics="true" pageId="16" pageNumber="367">
<taxonomicName id="89FCC5FF950CFF9E226EFC32FDDEE397" box="[328,593,976,1002]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="subSpecies" species="pinguis" subSpecies="radobojanus">B. pinguis radobojanus</taxonomicName>
.
</emphasis>
</paragraph>
<caption id="1A83EEF4950CFF9E23B1F83EFD3FE788" ID-DOI="http://doi.org/10.5281/zenodo.3741206" ID-Zenodo-Dep="3741206" box="[151,688,2012,2037]" httpUri="https://zenodo.org/record/3741206/files/figure.png" pageId="16" pageNumber="367" startId="16.[151,250,2012,2037]" targetBox="[151,1436,1023,1987]" targetPageId="16">
<paragraph id="4E43BE7C950CFF9E23B1F83EFD3FE788" blockId="16.[151,688,2012,2037]" box="[151,688,2012,2037]" pageId="16" pageNumber="367">
<emphasis id="7C88626E950CFF9E23B1F83EFE6FE788" bold="true" box="[151,480,2012,2037]" pageId="16" pageNumber="367">
FIGURE 15.
<emphasis id="7C88626E950CFF9E2200F83EFE6FE788" bold="true" box="[294,480,2012,2037]" italics="true" pageId="16" pageNumber="367">
<taxonomicName id="89FCC5FF950CFF9E2200F83EFE54E788" authorityName="Skartveit &amp; Krizmanić" authorityYear="2020" baseAuthorityName="Heer" baseAuthorityYear="1849" box="[294,475,2012,2037]" class="Insecta" family="Bibionidae" genus="Dilophus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="367" phylum="Arthropoda" rank="species" species="pinguis">Dilophus pinguis</taxonomicName>
,
</emphasis>
</emphasis>
male, CNHM 1832.
</paragraph>
</caption>
<caption id="1A83EEF4950DFF9F23B1FBD3FCA7E437" ID-DOI="http://doi.org/10.5281/zenodo.3741208" ID-Zenodo-Dep="3741208" box="[151,808,1073,1098]" httpUri="https://zenodo.org/record/3741208/files/figure.png" pageId="17" pageNumber="368" startId="17.[151,250,1073,1098]" targetBox="[216,1372,181,1048]" targetPageId="17">
<paragraph id="4E43BE7C950DFF9F23B1FBD3FCA7E437" blockId="17.[151,808,1073,1098]" box="[151,808,1073,1098]" pageId="17" pageNumber="368">
<emphasis id="7C88626E950DFF9F23B1FBD3FE56E437" bold="true" box="[151,473,1073,1098]" pageId="17" pageNumber="368">
FIGURE 16.
<taxonomicName id="89FCC5FF950DFF9F2200FBD3FE56E437" authorityName="Skartveit &amp; Krizmanić" authorityYear="2020" baseAuthorityName="Heer" baseAuthorityYear="1849" box="[294,473,1073,1098]" class="Insecta" family="Bibionidae" genus="Dilophus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="7C88626E950DFF9F2200FBD3FE56E437" bold="true" box="[294,473,1073,1098]" italics="true" pageId="17" pageNumber="368">Dilophus pinguis</emphasis>
</taxonomicName>
</emphasis>
, female, lectotype, SLJG 77473.
</paragraph>
</caption>
<caption id="1A83EEF4950DFF9F23B1F839FC5CE789" ID-DOI="http://doi.org/10.5281/zenodo.3741210" ID-Zenodo-Dep="3741210" box="[151,979,2011,2036]" httpUri="https://zenodo.org/record/3741210/files/figure.png" pageId="17" pageNumber="368" startId="17.[151,250,2011,2036]" targetBox="[217,1371,1120,1985]" targetPageId="17">
<paragraph id="4E43BE7C950DFF9F23B1F839FC5CE789" blockId="17.[151,979,2011,2036]" box="[151,979,2011,2036]" pageId="17" pageNumber="368">
<emphasis id="7C88626E950DFF9F23B1F839FE56E789" bold="true" box="[151,473,2011,2036]" pageId="17" pageNumber="368">
FIGURE 17.
<taxonomicName id="89FCC5FF950DFF9F2200F839FE56E789" authorityName="Skartveit &amp; Krizmanić" authorityYear="2020" baseAuthorityName="Heer" baseAuthorityYear="1849" box="[294,473,2011,2036]" class="Insecta" family="Bibionidae" genus="Dilophus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="368" phylum="Arthropoda" rank="species" species="pinguis">
<emphasis id="7C88626E950DFF9F2200F839FE56E789" bold="true" box="[294,473,2011,2036]" italics="true" pageId="17" pageNumber="368">Dilophus pinguis</emphasis>
</taxonomicName>
</emphasis>
, female, paralectotype, NHMW 1842/0050/0022.
</paragraph>
</caption>
<paragraph id="4E43BE7C950EFF9C23E1FF7AFD21E0CE" blockId="18.[151,1437,152,1079]" box="[199,686,152,179]" pageId="18" pageNumber="369">
<emphasis id="7C88626E950EFF9C23E1FF7AFE35E0CE" bold="true" box="[199,442,152,179]" pageId="18" pageNumber="369">Additional material.</emphasis>
Male (
<collectionCode id="28ED26B9950EFF9C212FFF7BFDD0E0CE" box="[521,607,153,179]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" lsid="urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="18" pageNumber="369" type="Museum">CNHM</collectionCode>
1832).
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FF5FFB8FE0AA" blockId="18.[151,1437,152,1079]" box="[199,1024,189,215]" pageId="18" pageNumber="369">
By far the biggest
<taxonomicName id="89FCC5FF950EFF9C22B0FF5FFE72E0AA" box="[406,509,189,215]" class="Insecta" family="Bibionidae" genus="Dilophus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="369" phylum="Arthropoda" rank="genus">
<emphasis id="7C88626E950EFF9C22B0FF5FFE72E0AA" box="[406,509,189,215]" italics="true" pageId="18" pageNumber="369">Dilophus</emphasis>
</taxonomicName>
species, fossil or recent, known from Europe.
</paragraph>
</subSubSection>
<subSubSection id="06E6EDF7950EFF9C23E1FF02FA8FE44A" pageId="18" pageNumber="369" type="description">
<paragraph id="4E43BE7C950EFF9C23E1FF02FEDEE086" blockId="18.[151,1437,152,1079]" box="[199,337,224,251]" pageId="18" pageNumber="369">
<emphasis id="7C88626E950EFF9C23E1FF02FEDEE086" bold="true" box="[199,337,224,251]" pageId="18" pageNumber="369">Description</emphasis>
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FEE7FCC7E162" blockId="18.[151,1437,152,1079]" box="[199,840,261,287]" pageId="18" pageNumber="369">
<emphasis id="7C88626E950EFF9C23E1FEE7FE8BE162" bold="true" box="[199,260,261,287]" pageId="18" pageNumber="369">Male</emphasis>
(N=1 for all measurements): Total length
<quantity id="89041399950EFF9C21FBFEE7FCCBE162" box="[733,836,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="18" pageNumber="369" unit="mm" value="12.6">12.6 mm</quantity>
.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FECBFD49E13E" blockId="18.[151,1437,152,1079]" box="[199,710,297,323]" pageId="18" pageNumber="369">Head: Not preserved in the specimen at hand.</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FEAFFBC0E11A" blockId="18.[151,1437,152,1079]" box="[199,1103,333,359]" pageId="18" pageNumber="369">
Thorax: Length
<quantity id="89041399950EFF9C225BFEAFFE59E11A" box="[381,470,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="18" pageNumber="369" unit="mm" value="3.2">3.2 mm</quantity>
. Very poorly preserved, spine combs not possible to see.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FE93FC04E1D2" blockId="18.[151,1437,152,1079]" pageId="18" pageNumber="369">
Wing: Length
<quantity id="89041399950EFF9C2249FE92FE46E1F6" box="[367,457,368,395]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="18" pageNumber="369" unit="mm" value="9.5">9.5 mm</quantity>
, width
<quantity id="89041399950EFF9C2107FE93FDF2E1F6" box="[545,637,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="369" unit="mm" value="3.3">3.3 mm</quantity>
. Dark brown fumose, anterior area a little darker than posterior. Vein measurements: basal R
<quantity id="89041399950EFF9C2286FE77FE77E1D2" box="[416,504,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="18" pageNumber="369" unit="mm" value="3.6">3.6 mm</quantity>
, distal R
<subScript id="D278BC39950EFF9C217CFE44FDEDE1C9" attach="left" box="[602,610,422,436]" fontSize="6" pageId="18" pageNumber="369">1</subScript>
<quantity id="89041399950EFF9C214FFE76FD4EE1D2" box="[617,705,404,431]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="369" unit="mm" value="2.5">2.5 mm</quantity>
, basal R
<geoCoordinate id="2BC8D8BB950EFF9C2006FE45FCDEE1D2" box="[800,849,405,437]" degrees="0.9" direction="south" orientation="latitude" pageId="18" pageNumber="369" precision="5555" value="-0.9">
<subScript id="D278BC39950EFF9C2006FE45FCA9E1C8" attach="left" box="[800,806,423,437]" fontSize="6" pageId="18" pageNumber="369">s</subScript>
<quantity id="89041399950EFF9C2008FE77FCDEE1D2" box="[814,849,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="369" unit="mm" value="0.9">0.9</quantity>
</geoCoordinate>
mm.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FE5BFD3AE266" blockId="18.[151,1437,152,1079]" pageId="18" pageNumber="369">
Legs: Fore leg relatively slender, tibia possibly with two small spines mesally, apical spines fine and short, not possible to count. Hind leg (
<figureCitation id="D6C7A2F9950EFF9C22F4FE3FFDACE18A" box="[466,547,477,503]" captionStart="FIGURES 32–35. 32" captionStartId="24.[151,264,1940,1965]" captionTargetBox="[159,1432,190,1903]" captionTargetId="figure@24.[151,1436,181,1914]" captionTargetPageId="24" captionText="FIGURES 32–35. 32. Bibio giganteus, female, holotype, wing, SLJG 77605. 33. Bibio giganteus, female, holotype, head and thorax, SLJG 77605. 34. Dilophus pinguis, male, hind leg, CNHM 1832. 35. Dilophus pinguis, male, hypopygium, CNHM 1832. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3741222" httpUri="https://zenodo.org/record/3741222/files/figure.png" pageId="18" pageNumber="369">Fig. 34</figureCitation>
) femur 4.0 mm long,
<quantity id="89041399950EFF9C2030FE3EFCF5E18A" box="[790,890,476,503]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="369" unit="mm" value="0.75">0.75 mm</quantity>
wide, moderately clavate, tibia
<quantity id="89041399950EFF9C27F8FE3FFABBE18A" box="[1246,1332,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="18" pageNumber="369" unit="mm" value="4.1">4.1 mm</quantity>
long,
<quantity id="89041399950EFF9C265FFE3FFF4CE266" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="369" unit="mm" value="0.6">0.6 mm</quantity>
wide, straight-sided, first tarsomere slender.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FDC7FB91E242" blockId="18.[151,1437,152,1079]" box="[199,1054,549,575]" pageId="18" pageNumber="369">
Abdomen: Length
<quantity id="89041399950EFF9C22BDFDC7FE7CE242" box="[411,499,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="18" pageNumber="369" unit="mm" value="8.3">8.3 mm</quantity>
, width
<quantity id="89041399950EFF9C2163FDC7FD12E242" box="[581,669,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="369" unit="mm" value="2.1">2.1 mm</quantity>
, cylindrical and relatively slender.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FDABFE28E2FA" blockId="18.[151,1437,152,1079]" pageId="18" pageNumber="369">
Terminalia (
<figureCitation id="D6C7A2F9950EFF9C2277FDABFE29E21E" box="[337,422,584,611]" captionStart="FIGURES 32–35. 32" captionStartId="24.[151,264,1940,1965]" captionTargetBox="[159,1432,190,1903]" captionTargetId="figure@24.[151,1436,181,1914]" captionTargetPageId="24" captionText="FIGURES 32–35. 32. Bibio giganteus, female, holotype, wing, SLJG 77605. 33. Bibio giganteus, female, holotype, head and thorax, SLJG 77605. 34. Dilophus pinguis, male, hind leg, CNHM 1832. 35. Dilophus pinguis, male, hypopygium, CNHM 1832. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3741222" httpUri="https://zenodo.org/record/3741222/files/figure.png" pageId="18" pageNumber="369">Fig. 35</figureCitation>
): Gonostylus digitiform as usual for genus. Shape of epandrium not possible to make out in the specimen at hand.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FD73FD91E2D6" blockId="18.[151,1437,152,1079]" box="[199,542,656,683]" pageId="18" pageNumber="369">
<emphasis id="7C88626E950EFF9C23E1FD73FE91E2D6" bold="true" box="[199,286,657,683]" pageId="18" pageNumber="369">Female</emphasis>
: Total length 15.0 mm
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FD57FEF1E36A" blockId="18.[151,1437,152,1079]" pageId="18" pageNumber="369">Head: Length 2.0 mm. Dark brown, presumably black in life. Complex eye small, rounded, protruding. Occiput approximately twice as long as complex eye. Ocellar tubercle low, not conspicuous. Antennal flagellum stout, possibly six-segmented.</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FCC3FD6BE322" blockId="18.[151,1437,152,1079]" pageId="18" pageNumber="369">
Thorax: Length
<quantity id="89041399950EFF9C225AFCC2FE5AE346" box="[380,469,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="18" pageNumber="369" unit="mm" value="5.6">5.6 mm</quantity>
. Ochreous-brown, pronotum probably a little darker. Spine rows on pronotum and mesonotum apparently consisting of short, stout spines.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FC8BFE3AE3DA" blockId="18.[151,1437,152,1079]" pageId="18" pageNumber="369">
Wing: Length
<quantity id="89041399950EFF9C2241FC8BFE42E3FE" box="[359,461,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="18" pageNumber="369" unit="mm" value="12.3">12.3 mm</quantity>
, width
<quantity id="89041399950EFF9C213BFC8BFDFAE3FE" box="[541,629,873,899]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="18" pageNumber="369" unit="mm" value="4.4">4.4 mm</quantity>
. Apparently hyaline (no pigment preserved). Veins colourless. Basal part of R
<subScript id="D278BC39950EFF9C23EEFC7DFF41E3D0" attach="left" box="[200,206,927,941]" fontSize="6" pageId="18" pageNumber="369">s</subScript>
about equal to R-M.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FC53FB10E46E" blockId="18.[151,1437,152,1079]" pageId="18" pageNumber="369">
Legs: Femorae ochreous-brown, tibiae and tarsi blackish-brown. Spine arrangement on protibia unclear: possibly three mesal spines in an oblique row. Apical spine circlet with short, widely separated spines, possibly six. Hind tibia and tarsus slender. Length of fore tibia
<quantity id="89041399950EFF9C21A1FC1BFD50E46E" box="[647,735,1017,1043]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="18" pageNumber="369" unit="mm" value="3.1">3.1 mm</quantity>
, hind femur
<quantity id="89041399950EFF9C204BFC1BFC4AE46E" box="[877,965,1017,1043]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="18" pageNumber="369" unit="mm" value="4.4">4.4 mm</quantity>
, hind tibia
<quantity id="89041399950EFF9C2764FC1BFB14E46E" box="[1090,1179,1017,1043]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="18" pageNumber="369" unit="mm" value="4.7">4.7 mm</quantity>
.
</paragraph>
<paragraph id="4E43BE7C950EFF9C23E1FBFFFA8FE44A" blockId="18.[151,1437,152,1079]" box="[199,1280,1053,1079]" pageId="18" pageNumber="369">
Abdomen: Length
<quantity id="89041399950EFF9C22BDFBFFFD8DE44A" box="[411,514,1053,1079]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="18" pageNumber="369" unit="mm" value="10.1">10.1 mm</quantity>
. Ochreous-brown. Pile not apparent. Cerci short, somewhat pointed.
</paragraph>
</subSubSection>
</treatment>
</document>
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