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178 lines
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<document id="2542306943D6481F0AE9465F9690DA47" ID-DOI="10.5281/zenodo.5330604" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168017471" checkinUser="felipe" docAuthor="Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong & Prokop, Jakub" docDate="2012" docId="B970A055FFE3FFC6FE0E2570FDA4FBBF" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.89-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Zetemenos sexlineatus Bode 1953" docType="treatment" docVersion="5" lastPageNumber="124" masterDocId="4549D82DFFC0FFE5FF9B2756FFB9FFFB" masterDocTitle="Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)" masterLastPageNumber="127" masterPageNumber="89" pageNumber="124" updateTime="1699171640213" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5FC640D6BEE3F884E518F3B0C5486DC3">Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)</mods:title>
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<mods:namePart id="C6A4FDE04684E3DAA5634674B1F3B8BB">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="6A709FBFEB45477CE69EA0CD8C47AE3B">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com & Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="6C621E5DB428AD6F84CB860745E25D6D">Prokin, Alexander</mods:namePart>
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<mods:affiliation id="76CD99C51ECB16D999B64FDC84D35934">Papanin Institute for Inland Water Biology, Russian Academy of Science, Borok, Russia; e-mail: prokina @ mail. ru & Voronezh State University, Research-Educational Centre “ Venevitinovo ”, Voronezh, Russia</mods:affiliation>
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<mods:namePart id="02F1647A8F662AC815314B3E70508187">Angus, Robert B.</mods:namePart>
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<mods:affiliation id="9E908BBFDB31E1388A8DAF46187BECEA">Department of Entomology, The Natural History Museum, London, UK; e-mail: r. angus @ rhul. ac. uk</mods:affiliation>
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<mods:namePart id="E66397A5FE4A2C0556D6F0DA0F2C5A2B">Pono, Alexander</mods:namePart>
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<mods:namePart id="2A5AA099AD89927D5317355E775C0AA2">Marenko</mods:namePart>
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<mods:affiliation id="D879BAA0113B93227D1A193E5B9FE7EC">Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia; e-mail: aponom @ paleo. ru</mods:affiliation>
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<mods:namePart id="BEA4CADC0B6B72B511DEE480BD740A9C">Yue, Yanli</mods:namePart>
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<mods:affiliation id="6E9CDA068AADA48F27B50840ECFE3C35">School of Life Science, Ningxia University, Ningxia, China; e-mail: yueyanli 0407 @ yahoo. cn</mods:affiliation>
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<mods:namePart id="9F3D23B0123E729DBE5CF062B8352829">Ren, Dong</mods:namePart>
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<mods:namePart id="4D74CCE9AEA0681BB63F6518E90D306B">Prokop, Jakub</mods:namePart>
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<mods:affiliation id="29125C48183D7E015FA9BF7E0882F898">Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:title id="FC22CDA8A992C82EF6261CC3D6392BE1">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="8C6E1C42E9CFE4EFE52163AC4D461208">2012</mods:date>
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<mods:number id="5710D07A3BC391B3D66A98D9AB5E33C4">2012-06-30</mods:number>
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<mods:number id="045FE90667148798697930842A73A0F8">52</mods:number>
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<mods:number id="7FB5CD08C821006C2BBD97DB86D23637">1</mods:number>
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<mods:start id="7FB86456186BB61DC9AA5A9805A0E921">89</mods:start>
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<mods:identifier id="CE7FCD46C9344763AFF2824BE03A77AA" type="DOI">10.5281/zenodo.5330604</mods:identifier>
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<mods:identifier id="0A2C9025F11872FF705397CF6D5CDFC8" type="ISSN">0374-1036</mods:identifier>
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<treatment id="B970A055FFE3FFC6FE0E2570FDA4FBBF" LSID="urn:lsid:plazi:treatment:B970A055FFE3FFC6FE0E2570FDA4FBBF" httpUri="http://treatment.plazi.org/id/B970A055FFE3FFC6FE0E2570FDA4FBBF" lastPageNumber="124" pageId="35" pageNumber="124">
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<subSubSection id="79C342C8FFE3FFC6FE0E2570FCA4FDC5" box="[405,797,550,575]" pageId="35" pageNumber="124" type="nomenclature">
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<paragraph id="31661143FFE3FFC6FE0E2570FCA4FDC5" blockId="35.[405,797,550,575]" box="[405,797,550,575]" pageId="35" pageNumber="124">
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<heading id="6A2EA62FFFE3FFC6FE0E2570FCA4FDC5" bold="true" box="[405,797,550,575]" centered="true" fontSize="8" level="2" pageId="35" pageNumber="124" reason="3">
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<emphasis id="03ADCD51FFE3FFC6FE0E2570FCA4FDC5" bold="true" box="[405,797,550,575]" pageId="35" pageNumber="124">
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†
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<taxonomicName id="F6D96AC0FFE3FFC6FE392571FCA4FDC5" authority="Bode, 1953" authorityName="Bode" authorityYear="1953" box="[418,797,550,575]" class="Insecta" family="Hydrophilidae" genus="Zetemenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="124" phylum="Arthropoda" rank="species" species="sexlineatus">
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<emphasis id="03ADCD51FFE3FFC6FE392571FD23FDC4" bold="true" box="[418,666,550,575]" italics="true" pageId="35" pageNumber="124">Zetemenos sexlineatus</emphasis>
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<bibRefCitation id="55486CB2FFE3FFC6FD3A2571FCA4FDC5" author="BODE A." box="[673,797,550,575]" pageId="35" pageNumber="124" pagination="1 - 375" refId="ref14118" refString="BODE A. 1953: Die Insektenfauna des ostniedersachsischen oberen Lias. Palaeontographica, Abteilung A 103 (1 - 4): 1 - 375." type="journal article" year="1953">Bode, 1953</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="79C342C8FFE3FFC6FD802513FD2EFDA3" box="[539,663,581,600]" pageId="35" pageNumber="124" type="description">
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<paragraph id="31661143FFE3FFC6FD802513FD2EFDA3" blockId="35.[113,663,581,661]" box="[539,663,581,600]" pageId="35" pageNumber="124">
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(
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<figureCitation id="A9E20DC6FFE3FFC6FDBA2513FDD1FDA3" box="[545,616,581,600]" captionStart="Figs" captionStartId="27.[113,152,1552,1571]" captionTargetBox="[113,1088,219,1524]" captionTargetId="figure-15@27.[113,1088,219,1524]" captionTargetPageId="27" captionText="Figs. 89–93. Representatives of Helophorus, subgenus Mesohelophorus Ponomarenko, 1977 (89–90) and taxa excluded from the helophorid lineage (91–93). 89–90 – Helophorus palaeosibiricus nom. nov., PIN 3063/841, holotype, piece and counterpiece; 91 – Helophoropsis brodiei (Giebel, 1856), holotype. 92 – Tychon antiquum (Giebel, 1856). holotype. 93 – Zetemenos sexlineatus Bode, 1953, holotype. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5453549" httpUri="https://zenodo.org/record/5453549/files/figure.png" pageId="35" pageNumber="124">Figs. 93</figureCitation>
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,
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<figureCitation id="A9E20DC6FFE3FFC6FDEA2513FD28FDA3" box="[625,657,581,600]" captionStart="Figs" captionStartId="32.[535,574,1412,1431]" captionTargetBox="[149,1128,222,1591]" captionTargetId="figure-10@32.[149,1127,212,1563]" captionTargetPageId="32" captionText="Figs.113–119.Taxa excluded from the helophorid lineage. 113 – Helo- phoropsis brodiei (Giebel, 1856), holotype; 114 – ‘Mesosperchus’ angulatus Ponomarenko, 1985, holotype; 115 – ‘Mesosperchus’ notatus Ponomarenko, 1977, holotype; 116 – Tychon antiquum (Giebel, 1856), holotype; 117–118 – ‘Mesosperchus’ schultzi Ponomarenko, 1985, holotype, piece and counterpiece; 119 – Zetemenos sexlineatus Bode, 1953, holotype. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5453551" httpUri="https://zenodo.org/record/5453551/files/figure.png" pageId="35" pageNumber="124">119</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="79C342C8FFE3FFC6FFEA253EFD9DFD6E" pageId="35" pageNumber="124" type="reference_group">
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<paragraph id="31661143FFE3FFC6FFEA253EFD9DFD6E" blockId="35.[113,663,581,661]" pageId="35" pageNumber="124">
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<treatmentCitationGroup id="11C9366DFFE3FFC6FFEA253EFD9DFD6E" pageId="35" pageNumber="124">
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<treatmentCitation id="B0783752FFE3FFC6FFEA253EFE7EFD81" author="BODE A." box="[113,455,615,634]" page="230" pageId="35" pageNumber="124" year="1953">
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<taxonomicName id="F6D96AC0FFE3FFC6FFEA253EFE7EFD81" authority="Bode, 1953: 230" authorityName="Bode" authorityPageNumber="230" authorityYear="1953" box="[113,455,615,634]" class="Insecta" family="Hydrophilidae" genus="Zetemenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="124" phylum="Arthropoda" rank="species" species="sexlineatus">
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<emphasis id="03ADCD51FFE3FFC6FFEA253EFE88FD81" box="[113,305,616,634]" italics="true" pageId="35" pageNumber="124">Zetemenos sexlineatus</emphasis>
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<bibRefCitation id="55486CB2FFE3FFC6FEAD2531FE7EFD81" author="BODE A." box="[310,455,615,634]" pageId="35" pageNumber="124" pagination="1 - 375" refId="ref14118" refString="BODE A. 1953: Die Insektenfauna des ostniedersachsischen oberen Lias. Palaeontographica, Abteilung A 103 (1 - 4): 1 - 375." type="journal article" year="1953">Bode, 1953: 230</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<treatmentCitation id="B0783752FFE3FFC6FFEA25D5FD99FD6E" author="PONOMARENKO A. G." box="[113,544,642,661]" page=")" pageId="35" pageNumber="124" year="1992">
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<taxonomicName id="F6D96AC0FFE3FFC6FFEA25D5FD99FD6E" authority=": PONOMARENKO (1992: 186)" authorityName="PONOMARENKO" authorityPageNumber="186" authorityYear="1992" box="[113,544,642,661]" class="Insecta" family="Hydrophilidae" genus="Zetemenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="124" phylum="Arthropoda" rank="species" species="sexlineatus">
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<emphasis id="03ADCD51FFE3FFC6FFEA25D5FE88FD6E" box="[113,305,643,661]" italics="true" pageId="35" pageNumber="124">Zetemenos sexlineatus</emphasis>
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:
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<bibRefCitation id="55486CB2FFE3FFC6FEA725D4FD99FD6E" author="PONOMARENKO A. G." box="[316,544,642,661]" pageId="35" pageNumber="124" pagination="179 - 188" refId="ref15829" refString="PONOMARENKO A. G. 1992: Upper Liassic beetles (Coleoptera) from Lower Saxony, Germany. Senckenbergiana Lethaea 72: 179 - 188." type="journal article" year="1992">PONOMARENKO (1992: 186)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="79C342C8FFE3FFC6FFEA25FAFDA4FBBF" pageId="35" pageNumber="124" type="materials_examined">
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<paragraph id="31661143FFE3FFC6FFEA25FAFC1AFD22" blockId="35.[113,1090,683,756]" pageId="35" pageNumber="124">
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<emphasis id="03ADCD51FFE3FFC6FFEA25FAFE88FD44" bold="true" box="[113,305,683,703]" pageId="35" pageNumber="124">
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<typeStatus id="EE62AFE1FFE3FFC6FFEA25FAFF24FD44" box="[113,157,684,703]" pageId="35" pageNumber="124">Type</typeStatus>
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locality and age.
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</emphasis>
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<collectingCountry id="49CE51D3FFE3FFC6FEAE25FAFE3CFD44" box="[309,389,684,703]" name="Germany" pageId="35" pageNumber="124">Germany</collectingCountry>
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,
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<collectingRegion id="F31DDFA1FFE3FFC6FE1725FAFDB3FD44" box="[396,522,684,703]" country="Germany" name="Niedersachsen" pageId="35" pageNumber="124">Niedersachsen</collectingRegion>
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, Grassel at Braunschweig (current
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<collectionCode id="57C88986FFE3FFC6FCB125FAFCE8FD44" box="[810,849,684,703]" pageId="35" pageNumber="124">GPS</collectionCode>
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coordinates:
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<geoCoordinate id="54ED7784FFE3FFC6FB9A25FAFB87FD44" box="[1025,1086,684,703]" degrees="10.6" direction="east" orientation="longitude" pageId="35" pageNumber="124" precision="5555" value="10.6">10.6°E</geoCoordinate>
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. Posidonia Shale Formation, Early Jurassic, Early Toarcian, 183.0–182.0 mya (
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<bibRefCitation id="55486CB2FFE3FFC6FC962590FC20FD22" author="CLAPHAM M." box="[781,921,710,729]" pageId="35" pageNumber="124" refId="ref14244" refString="CLAPHAM M. 2011 b: Grassel, Braunschweig. Paleobiology Database Collection No. 107321 [for Paleobiology Database homepage see http: // paleodb. org]. Visited on 21 st February 2012." type="url" year="2011">CLAPHAM 2011b</bibRefCitation>
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).
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</paragraph>
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<paragraph id="31661143FFE3FFC6FFEA25B6FDC9FD0F" blockId="35.[113,1090,683,756]" box="[113,624,736,756]" pageId="35" pageNumber="124">
|
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<emphasis id="03ADCD51FFE3FFC6FFEA25B6FE9AFD0F" bold="true" box="[113,291,736,756]" pageId="35" pageNumber="124">Material examined.</emphasis>
|
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|
<typeStatus id="EE62AFE1FFE3FFC6FEB325B7FEC5FD08" box="[296,380,737,756]" pageId="35" pageNumber="124" type="holotype">HOLOTYPE</typeStatus>
|
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: SMF
|
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<date id="45673783FFE3FFC6FE2C25B7FE47FD0F" box="[439,510,737,756]" pageId="35" pageNumber="124" value="1399-06">VI 1399</date>
|
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(piece only).
|
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</paragraph>
|
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<paragraph id="31661143FFE3FFC6FFEA245AFDA4FBBF" blockId="35.[113,1089,780,1093]" pageId="35" pageNumber="124">
|
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|
<emphasis id="03ADCD51FFE3FFC6FFEA245AFE96FCDF" bold="true" box="[113,303,780,804]" pageId="35" pageNumber="124">Taxonomic note.</emphasis>
|
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|
The
|
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|
|
<typeStatus id="EE62AFE1FFE3FFC6FEF2245AFE70FCDF" box="[361,457,780,804]" pageId="35" pageNumber="124" type="holotype">holotype</typeStatus>
|
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|
, which is the only available specimen of this species, is a badly preserved fossil in dorsal view showing an elongate beetle with striate elytra. Other characters are not preserved, except for the abdomen with five ventrites, narrow and elongate metepimera, and possibly the frontoclypeal suture (however, the impression on the head may also represent some ventral head structures). The latter character would be also the only possible indication of the position of the species in the
|
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<taxonomicName id="F6D96AC0FFE3FFC6FD4E24FAFC39FC3F" authorityName="Latreille" authorityYear="1802" box="[725,896,940,964]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="35" pageNumber="89" phylum="Tracheophyta" rank="superFamily" superFamily="Hydrophiloidea">Hydrophiloidea</taxonomicName>
|
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|
. However, due to the bad preservation we do not consider this character as adequate to justify the superfamily assignment of the fossil. For that reason, the species including the currently monotypic genus
|
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|
|
<taxonomicName id="F6D96AC0FFE3FFC6FF2D2358FE11FBDF" authority="Bode, 1953" authorityName="Bode" authorityYear="1953" box="[182,424,1036,1061]" class="Insecta" family="Hydrophilidae" genus="Zetemenos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="124" phylum="Arthropoda" rank="genus">
|
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|
|
<emphasis id="03ADCD51FFE3FFC6FF2D2358FE91FBDE" box="[182,296,1038,1061]" italics="true" pageId="35" pageNumber="124">Zetemenos</emphasis>
|
|||
|
|
<bibRefCitation id="55486CB2FFE3FFC6FEB5235AFE11FBDF" author="BODE A." box="[302,424,1036,1060]" pageId="35" pageNumber="124" pagination="1 - 375" refId="ref14118" refString="BODE A. 1953: Die Insektenfauna des ostniedersachsischen oberen Lias. Palaeontographica, Abteilung A 103 (1 - 4): 1 - 375." type="journal article" year="1953">Bode, 1953</bibRefCitation>
|
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|
|
</taxonomicName>
|
|||
|
|
to which it belongs are excluded from the
|
|||
|
|
<taxonomicName id="F6D96AC0FFE3FFC6FCF6235AFBADFBDF" authorityName="Latreille" authorityYear="1802" box="[877,1044,1036,1060]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="35" pageNumber="89" phylum="Tracheophyta" rank="superFamily" superFamily="Hydrophiloidea">Hydrophiloidea</taxonomicName>
|
|||
|
|
and transferred to
|
|||
|
|
<taxonomicName id="F6D96AC0FFE3FFC6FE9C237AFEC5FBBF" authority="Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop, 2012" authorityName="Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop" authorityYear="2012" box="[263,380,1068,1092]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="35" pageNumber="124" phylum="Tracheophyta" rank="order" status="incertae sedis">Coleoptera</taxonomicName>
|
|||
|
|
<emphasis id="03ADCD51FFE3FFC6FE182378FDAFFBBE" box="[387,534,1070,1093]" italics="true" pageId="35" pageNumber="124">
|
|||
|
|
<taxonomicNameLabel id="189E702AFFE3FFC6FE182378FDAFFBBE" box="[387,534,1070,1093]" pageId="35" pageNumber="124">incertae sedis</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|