394 lines
48 KiB
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394 lines
48 KiB
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<document id="AF8B85A2AB49EE6B73F97CE52DFD3824" ID-DOI="10.5281/zenodo.3709947" ID-GBIF-Dataset="f1f6714d-631a-4cc2-bbad-1fa240426fff" ID-ISSN="1942-1354" ID-Zenodo-Dep="3709947" ID-ZooBank="3A2B37FB-A2F6-4869-BFD8-2A8215128D2F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584124423284" checkinUser="felipe" docAuthor="Cano, Enio B." docDate="2018" docId="5B4E8796FFEE210FFF2AE3A6FA284C79" docLanguage="en" docName="InsectaMundi.0659.1-9.pdf.imf" docOrigin="Insecta Mundi 659" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Anomiopus cirulito Cano 2018, new species" docType="treatment" docVersion="6" lastPageNumber="3" masterDocId="A777FFEEFFED210BFFEAE37FFF884978" masterDocTitle="A new Anomiopus Westwood (Coleoptera: Scarabaeidae: Scarabaeinae) from the Mayan Biosphere Reserve, Petén, Guatemala" masterLastPageNumber="9" masterPageNumber="1" pageNumber="2" updateTime="1698728009997" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="F82CBF9782F6EDD048C16ED453A50904">
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<mods:title id="2969EA4A53C07D9C14799FAC16D37584">A new Anomiopus Westwood (Coleoptera: Scarabaeidae: Scarabaeinae) from the Mayan Biosphere Reserve, Petén, Guatemala</mods:title>
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<mods:namePart id="FC88691CDBE6D881CC42044D76DE56AD">Cano, Enio B.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="91F414FC40B5149B7491A98C5BDF49F9">text</mods:typeOfResource>
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<mods:title id="80E1E0274F988BD5ADDBA8E1A1469397">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="781E578FA5C2BEA75F73BDB18AA89F8C">
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<mods:date id="601BC3FA6BA5EF6E700FC0368F27BF54">2018</mods:date>
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<mods:detail id="B5CDC8FE8CCFBA7B72DEAC39E86728BB" type="pubDate">
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<mods:number id="E50B2E1C1E633FE9949A42E2431FC737">2018-09-28</mods:number>
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<mods:number id="08543899C5483F3B4BAC8FA6092D8420">659</mods:number>
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<mods:start id="A770F8EB2B17047C9A17C7DFA0DA359B">1</mods:start>
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<mods:end id="33D2BD1C02BFB5ECC7130DF58DD9A04A">9</mods:end>
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<mods:classification id="DEA6C029177210FFD9778B79E3F6E326">journal article</mods:classification>
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<mods:identifier id="81C75854BCA7CFE3DA01411FDEF33C9B" type="DOI">10.5281/zenodo.3709947</mods:identifier>
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<mods:identifier id="255FD198B4A8F2789991D45B0B00F25D" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="9874F9A2D60D51FF0CD4EB53FA4E508D" type="Zenodo-Dep">3709947</mods:identifier>
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<treatment id="5B4E8796FFEE210FFF2AE3A6FA284C79" ID-DOI="http://doi.org/10.5281/zenodo.3717280" ID-GBIF-Taxon="162488076" ID-Zenodo-Dep="3717280" LSID="urn:lsid:plazi:treatment:5B4E8796FFEE210FFF2AE3A6FA284C79" httpUri="http://treatment.plazi.org/id/5B4E8796FFEE210FFF2AE3A6FA284C79" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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<subSubSection id="9BFD650BFFEE2108FF2AE3A6FD404989" box="[192,712,217,241]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="D3583680FFEE2108FF2AE3A6FD404989" blockId="3.[192,712,217,274]" box="[192,712,217,241]" pageId="3" pageNumber="2">
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<heading id="881081ECFFEE2108FF2AE3A6FD404989" bold="true" box="[192,712,217,241]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
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<emphasis id="E193EA92FFEE2108FF2AE3A6FD404989" bold="true" box="[192,712,217,241]" pageId="3" pageNumber="2">
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<taxonomicName id="14E74D03FFEE2108FF2AE3A6FD9F4989" ID-CoL="676KX" authority="Cano" authorityName="Cano" authorityYear="2018" box="[192,535,217,241]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="cirulito" status="sp. nov.">
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<emphasis id="E193EA92FFEE2108FF2AE3A6FE4E4989" bold="true" box="[192,454,217,241]" italics="true" pageId="3" pageNumber="2">Anomiopus cirulito</emphasis>
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Cano
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</taxonomicName>
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,
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<taxonomicNameLabel id="FAA057E9FFEE2108FDCEE3A6FD404989" box="[548,712,217,241]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFD650BFFEE2108FF2AE386FDAE4FE9" pageId="3" pageNumber="2" type="description">
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<paragraph id="D3583680FFEE2108FF2AE386FEBC486A" blockId="3.[192,712,217,274]" box="[192,308,249,274]" pageId="3" pageNumber="2">
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(
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<figureCitation id="4BDC2A05FFEE2108FF23E386FEA4486A" box="[201,300,249,274]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="8.[192,275,1877,1899]" captionStartId-1="9.[192,276,1877,1899]" captionTargetBox-0="[217,1416,213,1843]" captionTargetBox-1="[217,1416,213,1843]" captionTargetId-0="figure@8.[217,1416,213,1843]" captionTargetId-1="figure@9.[217,1416,213,1843]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figure 3. Anomiopus cirulito n. sp. a) Dorsal habitus. b) Ventral habitus. c) Lateral view." captionText-1="Figure 4. Anomiopus cirulito, n. sp. a) Head, pronotum and left tibia. b) Detail of right tibia. c) Details of metasternum in latero-ventral view. CxII = coxa II; Mt, metasternum; Mtp = metepisternum; Ms = mesepimeron." figureDoi-0="http://doi.org/10.5281/zenodo.3709953" figureDoi-1="http://doi.org/10.5281/zenodo.3709955" httpUri-0="https://zenodo.org/record/3709953/files/figure.png" httpUri-1="https://zenodo.org/record/3709955/files/figure.png" pageId="3" pageNumber="2">Fig. 3–4</figureCitation>
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)
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</paragraph>
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<paragraph id="D3583680FFEE2108FF2AE256FE7048D9" blockId="3.[192,1440,297,417]" pageId="3" pageNumber="2">
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<emphasis id="E193EA92FFEE2108FF2AE256FD354839" bold="true" box="[192,701,297,321]" pageId="3" pageNumber="2">
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Description.
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<typeStatus id="0C5C8822FFEE2108FE87E256FE614839" box="[365,489,297,321]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
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(sex unknown):
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</emphasis>
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Length
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<quantity id="141F9B65FFEE2108FCF5E256FCED4839" box="[799,869,297,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="2" unit="mm" value="5.0">5 mm</quantity>
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, elongate, almost parallel sided (
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<figureCitation id="4BDC2A05FFEE2108FB03E256FAB4483A" box="[1257,1340,297,322]" captionStart="Figure 3" captionStartId="8.[192,275,1877,1899]" captionTargetBox="[217,1416,213,1843]" captionTargetId="figure@8.[217,1416,213,1843]" captionTargetPageId="8" captionText="Figure 3. Anomiopus cirulito n. sp. a) Dorsal habitus. b) Ventral habitus. c) Lateral view." figureDoi="http://doi.org/10.5281/zenodo.3709953" httpUri="https://zenodo.org/record/3709953/files/figure.png" pageId="3" pageNumber="2">Fig. 3a</figureCitation>
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). Dorsal and ventral surface glossy, brownish black, with mostly cupreous reflections (occasionally green); legs, pygidium and last abdominal sternite reddish brown with cupreous and some green reflections; anten- nal club yellowish brown.
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</paragraph>
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<paragraph id="D3583680FFEE2108FF2AE2C6FD824B89" blockId="3.[192,1441,441,753]" pageId="3" pageNumber="2">
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<emphasis id="E193EA92FFEE2108FF2AE2C6FE8048A9" bold="true" box="[192,264,441,465]" pageId="3" pageNumber="2">Head</emphasis>
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(
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<figureCitation id="4BDC2A05FFEE2108FEF0E2C6FEFB48AA" box="[282,371,441,466]" captionStart="Figure 4" captionStartId="9.[192,276,1877,1899]" captionTargetBox="[217,1416,213,1843]" captionTargetId="figure@9.[217,1416,213,1843]" captionTargetPageId="9" captionText="Figure 4. Anomiopus cirulito, n. sp. a) Head, pronotum and left tibia. b) Detail of right tibia. c) Details of metasternum in latero-ventral view. CxII = coxa II; Mt, metasternum; Mtp = metepisternum; Ms = mesepimeron." figureDoi="http://doi.org/10.5281/zenodo.3709955" httpUri="https://zenodo.org/record/3709955/files/figure.png" pageId="3" pageNumber="2">Fig. 4a</figureCitation>
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): Clypeal margin with four reflexed teeth distributed as follows: two elongate, median semi-cylindrical teeth separated by deep, U-shaped emargination; and shorter, triangular lateral teeth each separated from adjacent median tooth by wide notch. Genal border slightly arcuate, with weak, wide tooth forming shallow V-shaped notch with adjacent clypeal tooth. Posterior one-half of frontoclypeal suture deeper than anterior one-half. Frons with two weak, conical, tubercles. Vertex with medial area concave; occipital bead complete. Dorsal surface of head punctate, punctures separated by at least 2.5 diameters; anterior and lateral borders, and vertex behind tubercles, strongly and deeply punctate, punctures separated by at most 1.5 diameters. Maximum width of head
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<quantity id="141F9B65FFEE2108FBC9E1E5FB184BC9" box="[1059,1168,666,690]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="3" pageNumber="2" unit="mm" value="1.44">1.44 mm</quantity>
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. Dorsal surface of eyes narrow and elongated (maximum width of one eye
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<quantity id="141F9B65FFEE2108FCD5E1C5FC264BAA" box="[831,942,698,722]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="3" pageNumber="2" unit="mm" value="0.07">0.07 mm</quantity>
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); interocular distance
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<quantity id="141F9B65FFEE2108FB2FE1C5FABD4BA9" box="[1221,1333,698,722]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="3" pageNumber="2" unit="mm" value="0.93">0.93 mm</quantity>
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. Clypeal ventral process cariniform.
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</paragraph>
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<paragraph id="D3583680FFEE2108FF2AE076FD864DF9" blockId="3.[192,1441,777,1218]" pageId="3" pageNumber="2">
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<emphasis id="E193EA92FFEE2108FF2AE076FEA44A59" bold="true" box="[192,300,777,801]" pageId="3" pageNumber="2">Thorax:</emphasis>
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Pronotum evenly, broadly convex, completely finely punctate, punctures denser at sides and more dispersed and minute at disc and posteriorly. Disc with medial longitudinal sulcus distinct, shallow; sulcus extending from middle and occupying 2/3 of posterior half but not reaching posterior margin. Pronotal anterior and lateral bead present, thickening and forming strong callus at postero-lateral end (
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<figureCitation id="4BDC2A05FFEE2108FF23E0F6FE944ADA" box="[201,284,905,930]" captionStart="Figure 3" captionStartId="8.[192,275,1877,1899]" captionTargetBox="[217,1416,213,1843]" captionTargetId="figure@8.[217,1416,213,1843]" captionTargetPageId="8" captionText="Figure 3. Anomiopus cirulito n. sp. a) Dorsal habitus. b) Ventral habitus. c) Lateral view." figureDoi="http://doi.org/10.5281/zenodo.3709953" httpUri="https://zenodo.org/record/3709953/files/figure.png" pageId="3" pageNumber="2">Fig. 3c</figureCitation>
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); posterior margin not beaded. Protoracic margin (
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<figureCitation id="4BDC2A05FFEE2108FC61E0F6FC554ADA" box="[907,989,905,930]" captionStart="Figure 3" captionStartId="8.[192,275,1877,1899]" captionTargetBox="[217,1416,213,1843]" captionTargetId="figure@8.[217,1416,213,1843]" captionTargetPageId="8" captionText="Figure 3. Anomiopus cirulito n. sp. a) Dorsal habitus. b) Ventral habitus. c) Lateral view." figureDoi="http://doi.org/10.5281/zenodo.3709953" httpUri="https://zenodo.org/record/3709953/files/figure.png" pageId="3" pageNumber="2">Fig. 3c</figureCitation>
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) with marked median angle; anterolateral angles acute; posterolateral angles acute and forming a marked posteriorly-directed, tooth-like projection (
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<figureCitation id="4BDC2A05FFEE2108FEA1E0B6FE174A9A" box="[331,415,969,994]" captionStart="Figure 3" captionStartId="8.[192,275,1877,1899]" captionTargetBox="[217,1416,213,1843]" captionTargetId="figure@8.[217,1416,213,1843]" captionTargetPageId="8" captionText="Figure 3. Anomiopus cirulito n. sp. a) Dorsal habitus. b) Ventral habitus. c) Lateral view." figureDoi="http://doi.org/10.5281/zenodo.3709953" httpUri="https://zenodo.org/record/3709953/files/figure.png" pageId="3" pageNumber="2">Fig. 3c</figureCitation>
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). Pronotal fossae visible and shallow. Propleuron with complete transverse carina; proepisternum widely and deeply concave, with scattered long setae; proepimeron shining and barely convex. Sternellum (prosternum) with acute apex. Mesosternum with median transverse carina, posterior half finely micro-striated. Metasternum shining, without micro-striations (
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<figureCitation id="4BDC2A05FFEE2108FB67E756FB6A4D3A" box="[1165,1250,1065,1090]" captionStart="Figure 4" captionStartId="9.[192,276,1877,1899]" captionTargetBox="[217,1416,213,1843]" captionTargetId="figure@9.[217,1416,213,1843]" captionTargetPageId="9" captionText="Figure 4. Anomiopus cirulito, n. sp. a) Head, pronotum and left tibia. b) Detail of right tibia. c) Details of metasternum in latero-ventral view. CxII = coxa II; Mt, metasternum; Mtp = metepisternum; Ms = mesepimeron." figureDoi="http://doi.org/10.5281/zenodo.3709955" httpUri="https://zenodo.org/record/3709955/files/figure.png" pageId="3" pageNumber="2">Fig. 4c</figureCitation>
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); disc flat, with medial longitudinal sulcus clearly impressed and
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<quantity id="141F9B65FFEE2108FCF0E735FC0B4D19" box="[794,899,1098,1122]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" pageId="3" pageNumber="2" unit="mm" value="1.33">1.33 mm</quantity>
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in length. Mesepimeron and metepisternum shining, without striations.
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</paragraph>
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<paragraph id="D3583680FFEE2108FF02E7F6FA284DB9" blockId="3.[192,1441,777,1218]" pageId="3" pageNumber="2">Elytra with seven moderately impressed striae; striae 3, 5, 6, and 7 effaced apically; third shorter than fourth with striae 8 and 9 absent. Interestriae moderately convex, with minute, sparse punctures.</paragraph>
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<paragraph id="D3583680FFEE2108FF2AE7A6FB074C69" blockId="3.[192,1440,1241,1297]" pageId="3" pageNumber="2">
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<emphasis id="E193EA92FFEE2108FF2AE7A6FEC14D89" bold="true" box="[192,329,1241,1265]" pageId="3" pageNumber="2">Abdomen:</emphasis>
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Abdominal sternites smooth and shining, moderately convex; length of sternite VIII equal to combined lengths of sternites VI and VII. Pre-pygidium finely micro-striated.
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</paragraph>
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<paragraph id="D3583680FFEE2108FF2AE656FDEA4CB9" blockId="3.[192,1440,1321,1569]" pageId="3" pageNumber="2">
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<emphasis id="E193EA92FFEE2108FF2AE656FE804C39" bold="true" box="[192,264,1321,1345]" pageId="3" pageNumber="2">Legs:</emphasis>
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Pro- meso- and metacoxae finely striated. Protibiae (
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<figureCitation id="4BDC2A05FFEE2108FC7AE656FB984C39" box="[912,1040,1321,1346]" captionStart="Figure 4" captionStartId="9.[192,276,1877,1899]" captionTargetBox="[217,1416,213,1843]" captionTargetId="figure@9.[217,1416,213,1843]" captionTargetPageId="9" captionText="Figure 4. Anomiopus cirulito, n. sp. a) Head, pronotum and left tibia. b) Detail of right tibia. c) Details of metasternum in latero-ventral view. CxII = coxa II; Mt, metasternum; Mtp = metepisternum; Ms = mesepimeron." figureDoi="http://doi.org/10.5281/zenodo.3709955" httpUri="https://zenodo.org/record/3709955/files/figure.png" pageId="3" pageNumber="2">Fig. 4a, 4b</figureCitation>
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) with three apical teeth (wide at base in left tibia; slender and with an extra denticle between penultimate and antepenultimate teeth, and with two denticles between the last and penultimate teeth in the right tibia). Basal portion of lateral protibial margin with 8 (left leg) and 9 (right leg) denticles. Meso- and metatibiae rectangular, not appreciably narrowed at ends.
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</paragraph>
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<paragraph id="D3583680FFEE2108FF02E6B6FE3F4F59" blockId="3.[192,1440,1321,1569]" pageId="3" pageNumber="2">
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Metafemur with anterior longitudinal groove on ventral surface (
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<figureCitation id="4BDC2A05FFEE2108FBEFE6B6FBD14C9A" box="[1029,1113,1481,1506]" captionStart="Figure 3" captionStartId="8.[192,275,1877,1899]" captionTargetBox="[217,1416,213,1843]" captionTargetId="figure@8.[217,1416,213,1843]" captionTargetPageId="8" captionText="Figure 3. Anomiopus cirulito n. sp. a) Dorsal habitus. b) Ventral habitus. c) Lateral view." figureDoi="http://doi.org/10.5281/zenodo.3709953" httpUri="https://zenodo.org/record/3709953/files/figure.png" pageId="3" pageNumber="2">Fig. 3c</figureCitation>
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). Tarsi almost rectangular with the apical border truncated; length of first metatarsal segment slightly greater than that second and third combined.
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</paragraph>
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<paragraph id="D3583680FFEE2108FF2AE546FDAE4FE9" blockId="3.[192,1440,1593,1681]" pageId="3" pageNumber="2">
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<emphasis id="E193EA92FFEE2108FF2AE546FE194F29" bold="true" box="[192,401,1593,1617]" pageId="3" pageNumber="2">Measurements:</emphasis>
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Total length (apex of clypeal teeth to pygidum): 5.0 mm; pronotal length:
|
|||
|
|
<quantity id="141F9B65FFEE2108FAC7E546FA134F29" box="[1325,1435,1593,1617]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="3" pageNumber="2" unit="mm" value="1.62">1.62 mm</quantity>
|
|||
|
|
; pronotal width:
|
|||
|
|
<quantity id="141F9B65FFEE2108FE69E526FE674F09" box="[387,495,1625,1649]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="3" pageNumber="2" unit="mm" value="2.25">2.25 mm</quantity>
|
|||
|
|
; elytral length along suture:
|
|||
|
|
<quantity id="141F9B65FFEE2108FCA5E525FC334F09" box="[847,955,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="2.31" pageId="3" pageNumber="2" unit="mm" value="2.31">2.31 mm</quantity>
|
|||
|
|
; elytral length on sides:
|
|||
|
|
<quantity id="141F9B65FFEE2108FB09E526FAC74F09" box="[1251,1359,1625,1649]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="3" pageNumber="2" unit="mm" value="2.75">2.75 mm</quantity>
|
|||
|
|
; maximum elytral width:
|
|||
|
|
<quantity id="141F9B65FFEE2108FE5EE506FDAA4FE9" box="[436,546,1657,1681]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="3" pageNumber="2" unit="mm" value="2.25">2.25 mm</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="9BFD650BFFEE2108FF2AE5D6FCA24E5A" pageId="3" pageNumber="2" type="materials_examined">
|
|||
|
|
<paragraph id="D3583680FFEE2108FF2AE5D6FCA24E5A" blockId="3.[192,1440,1705,1826]" pageId="3" pageNumber="2">
|
|||
|
|
<emphasis id="E193EA92FFEE2108FF2AE5D6FE054FB9" bold="true" box="[192,397,1705,1729]" pageId="3" pageNumber="2">Type material:</emphasis>
|
|||
|
|
<materialsCitation id="638F3CDDFFEE2108FE72E5D6FCA24E5A" ID-GBIF-Occurrence="2575925882" collectingDate="2004-02-24" collectionCode="USAC" collectorName="A. Higueros. Deposited" country="Guatemala" county="Mayan Biosphere Reserve" elevation="299" latitude="17.72469" location="El Cedro" longLatPrecision="1" longitude="-89.50235" municipality="Rio Azul National Park" pageId="3" pageNumber="2" specimenCount="1" stateProvince="Peten Department" typeStatus="holotype">
|
|||
|
|
<typeStatus id="0C5C8822FFEE2108FE72E5D6FD824FB9" box="[408,522,1705,1729]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
|
|||
|
|
:
|
|||
|
|
<collectingCountry id="ABF07610FFEE2108FDF3E5D5FD584FBA" box="[537,720,1706,1730]" name="Guatemala" pageId="3" pageNumber="2">GUATEMALA</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="1123F862FFEE2108FD35E5D5FC424FB9" box="[735,970,1706,1730]" country="Guatemala" name="Peten" pageId="3" pageNumber="2">Petén Department</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="3A394E0CFFEE2108FC33E5D5FAAB4FB9" box="[985,1315,1705,1730]" pageId="3" pageNumber="2">Mayan Biosphere Reserve</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="333CACFAFFEE2108FAD9E5D5FEFC4F99" pageId="3" pageNumber="2">Río Azul National Park</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<location id="D638605BFFEE2108FE6BE5B6FE664F99" LSID="urn:lsid:plazi:treatment:5B4E8796FFEE210FFF2AE3A6FA284C79:D638605BFFEE2108FE6BE5B6FE664F99" box="[385,494,1737,1761]" country="Guatemala" county="Mayan Biosphere Reserve" latitude="17.72469" longLatPrecision="1" longitude="-89.50235" municipality="Rio Azul National Park" name="El Cedro" pageId="3" pageNumber="2" stateProvince="Peten Department">El Cedro</location>
|
|||
|
|
near aguada “
|
|||
|
|
<location id="D638605BFFEE2108FD4FE5B5FCDE4F99" LSID="urn:lsid:plazi:treatment:5B4E8796FFEE210FFF2AE3A6FA284C79:D638605BFFEE2108FD4FE5B5FCDE4F99" box="[677,854,1737,1762]" country="Guatemala" county="Mayan Biosphere Reserve" latitude="17.72469" longLatPrecision="1" longitude="-89.50235" municipality="Rio Azul National Park" name="Los Monifatos" pageId="3" pageNumber="2" stateProvince="Peten Department">Los Monifatos</location>
|
|||
|
|
”; coordinates
|
|||
|
|
<geoCoordinate id="B6D35047FFEE2108FBEEE5B6FBFE4F99" box="[1028,1142,1737,1761]" degrees="17.72469" direction="north" orientation="latitude" pageId="3" pageNumber="2" precision="1" value="17.72469">17.72469</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="B6D35047FFEE2108FB69E5B6FB754F99" box="[1155,1277,1737,1761]" degrees="89.50235" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="1" value="-89.50235">-89.50235</geoCoordinate>
|
|||
|
|
;
|
|||
|
|
<quantity id="141F9B65FFEE2108FAE0E5B5FAC74F9A" box="[1290,1359,1738,1762]" metricMagnitude="2" metricUnit="m" metricValue="2.9899999999999998" pageId="3" pageNumber="2" unit="m" value="299.0">
|
|||
|
|
<elevation id="58CAD1B3FFEE2108FAE0E5B5FAC74F9A" box="[1290,1359,1738,1762]" metricMagnitude="2" metricUnit="m" metricValue="2.9899999999999998" pageId="3" pageNumber="2" unit="m" value="299.0">299m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
elevation;
|
|||
|
|
<date id="A7591040FFEE2108FF15E595FE094E7A" box="[255,385,1770,1794]" pageId="3" pageNumber="2" value="2004-02-24">
|
|||
|
|
<collectingDate id="B71DE9A8FFEE2108FF15E595FE094E7A" box="[255,385,1770,1794]" pageId="3" pageNumber="2" value="2004-02-24">24 II 2004</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
.
|
|||
|
|
<collectorName id="7E125356FFEE2108FE66E595FD204E79" box="[396,680,1769,1794]" pageId="3" pageNumber="2">A. Higueros. Deposited</collectorName>
|
|||
|
|
at the Colección Entomológica de la Escuela de Biología de la Universidad de San Carlos de Guatemala (
|
|||
|
|
<collectionCode id="B5F6AE45FFEE2108FD26E475FC964E5A" box="[716,798,1802,1826]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="3" pageNumber="2">USAC</collectionCode>
|
|||
|
|
).
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="9BFD650BFFEE210FFF2AE446FB0048A9" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="biology_ecology">
|
|||
|
|
<paragraph id="D3583680FFEE210FFF2AE446FB0048A9" blockId="3.[192,1440,1849,1905]" lastBlockId="4.[192,1440,217,466]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
|
|||
|
|
<emphasis id="E193EA92FFEE2108FF2AE446FEBD4E29" bold="true" box="[192,309,1849,1873]" pageId="3" pageNumber="2">Ecology.</emphasis>
|
|||
|
|
The
|
|||
|
|
<typeStatus id="0C5C8822FFEE2108FE99E446FE534E29" box="[371,475,1849,1873]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
|
|||
|
|
was collected in a pitfall trap baited with rotten fish. Approximately the other 45,000 specimens collected in the area did not include additional specimens of the new species. The type locality is dominated by tropical rain forest growing on karst (
|
|||
|
|
<figureCitation id="4BDC2A05FFE9210FFC0FE3A6FBA34989" box="[997,1067,217,241]" captionStart="Figure 5" captionStartId="10.[192,275,1877,1899]" captionTargetId="figure@10.[192,1440,560,1496]" captionTargetPageId="10" captionText="Figure 5. Vegetation and mammals at the “aguada” (wetland) Los Monifatos, the type locality of A. cirulito n. sp., in the Mayan Biosphere Reserve. Image from a camera-trap, 23 II 2015, 12:17:25 hrs, courtesy of WCS Guatemala." figureDoi="http://doi.org/10.5281/zenodo.3709957" httpUri="https://zenodo.org/record/3709957/files/figure.png" pageId="4" pageNumber="3">Fig. 5</figureCitation>
|
|||
|
|
). According to
|
|||
|
|
<bibRefCitation id="B7764B71FFE9210FFB37E3A6FA284989" author="Gonzalez, V. R." box="[1245,1440,217,241]" pageId="4" pageNumber="3" refId="ref3350" refString="Gonzalez, V. R. 2015. Vertebrados medianos y mayores asociados a las aguadas del Biotopo Protegido Naachtun - Dos Lagunas, Peten. Tesis Licenciatura en Biologia, Facultad de Ciencias Quimicas y Farmacia, Universidad de San Carlos de Guatemala. 124 p." type="book" year="2015">González (2015)</bibRefCitation>
|
|||
|
|
the predominant vegetation of the zone includes
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFCF8E386FA954869" authority="(Kunth) Dugand" authorityName="Dugand" authorityYear="1943" baseAuthorityName="Kunth" box="[786,1309,249,273]" class="Magnoliopsida" family="Malvaceae" genus="Pseudobombax" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="species" species="ellipticum">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFCF8E386FBC24869" box="[786,1098,249,273]" italics="true" pageId="4" pageNumber="3">Pseudobombax ellipticum</emphasis>
|
|||
|
|
(Kunth) Dugand
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFAC2E385FEF04849" authority="Lam." authorityName="Lam." class="Magnoliopsida" family="Malvaceae" genus="Guazuma" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="species" species="ulmifolia">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFAC2E385FEB84849" italics="true" pageId="4" pageNumber="3">Guazuma ulmifolia</emphasis>
|
|||
|
|
Lam.
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFE68E265FE624849" box="[386,490,282,305]" class="Magnoliopsida" family="Sapotaceae" genus="Pouteria" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFE68E265FE624849" box="[386,490,282,305]" italics="true" pageId="4" pageNumber="3">Pouteria</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
spp.,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFDC4E265FCE4484A" authority="Sw." authorityName="Sw." box="[558,876,281,306]" class="Magnoliopsida" family="Moraceae" genus="Brosimum" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="species" species="alicastrum">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFDC4E265FCBF4849" box="[558,823,281,305]" italics="true" pageId="4" pageNumber="3">Brosimum alicastrum</emphasis>
|
|||
|
|
Sw.
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFC9CE266FAAF4849" authority="(Standl.) Standl." authorityName="Standl." baseAuthorityName="Standl." box="[886,1319,281,305]" class="Magnoliopsida" family="Cannabaceae" genus="Ampelocera" kingdom="Plantae" order="Rosales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="species" species="hottlei">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC9CE266FBDE4849" box="[886,1110,281,305]" italics="true" pageId="4" pageNumber="3">Ampelocera hottlei</emphasis>
|
|||
|
|
(Standl.) Standl.
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFADBE265FD034829" authority="(Britton & P. Wilson) Standl." authorityName="Standl." baseAuthorityName="Britton & P. Wilson" class="Magnoliopsida" family="Fabaceae" genus="Swartzia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="species" species="cubensis">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFADBE265FEAF4829" italics="true" pageId="4" pageNumber="3">Swartzia cubensis</emphasis>
|
|||
|
|
(Britton & P. Wilson) Standl.
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFD7FE245FC244829" authority="Standl." authorityName="Standl." box="[661,940,313,337]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="species" species="lundelli">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFD7FE245FCC34829" box="[661,843,313,337]" italics="true" pageId="4" pageNumber="3">Ocotea lundelli</emphasis>
|
|||
|
|
Standl.
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFC00E246FF7D4809" authority="Benth. Also" authorityName="Benth. Also" authorityYear="1946" class="Magnoliopsida" family="Fabaceae" genus="Lonchocarpus" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="species" species="guatemalensis">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC00E246FAC04829" box="[1002,1352,313,337]" italics="true" pageId="4" pageNumber="3">Lonchocarpus guatemalensis</emphasis>
|
|||
|
|
Benth. Also
|
|||
|
|
</taxonomicName>
|
|||
|
|
ground vegetation includes escobo (
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFD47E226FB5B480A" authority="(Heynh.) R. Evans" authorityName="R. Evans" baseAuthorityName="Heynh." box="[685,1235,345,370]" class="Liliopsida" family="Arecaceae" genus="Cryosophila" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="species" species="stauracantha">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFD47E226FC624809" box="[685,1002,345,369]" italics="true" pageId="4" pageNumber="3">Cryosophila stauracantha</emphasis>
|
|||
|
|
(Heynh.) R. Evans
|
|||
|
|
</taxonomicName>
|
|||
|
|
) and bayal (
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFA81E225FE9348E9" authorityName="C.F.P.Martius" authorityYear="1824" class="Liliopsida" family="Arecaceae" genus="Desmoncus" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="3" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFA81E225FE9348E9" italics="true" pageId="4" pageNumber="3">Desmoncus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp.). According to
|
|||
|
|
<bibRefCitation id="B7764B71FFE9210FFE1BE206FD2748E9" author="Gonzalez, V. R." box="[497,687,377,401]" pageId="4" pageNumber="3" refId="ref3350" refString="Gonzalez, V. R. 2015. Vertebrados medianos y mayores asociados a las aguadas del Biotopo Protegido Naachtun - Dos Lagunas, Peten. Tesis Licenciatura en Biologia, Facultad de Ciencias Quimicas y Farmacia, Universidad de San Carlos de Guatemala. 124 p." type="book" year="2015">González (2015)</bibRefCitation>
|
|||
|
|
mammals frequently observed by camera traps were
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFACFE206FF7148C9" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" class="Mammalia" family="Cuniculidae" genus="Cuniculus" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="3" phylum="Chordata" rank="species" species="paca">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFACFE206FF7148C9" italics="true" pageId="4" pageNumber="3">Cuniculus paca</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(L.),
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFEDDE2E5FDF048CA" authority="Gray" authorityName="Gray" authorityYear="1842" box="[311,632,410,434]" class="Mammalia" family="Dasyproctidae" genus="Dasyprocta" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="3" phylum="Chordata" rank="species" species="punctata">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFEDDE2E5FDBA48C9" box="[311,562,410,433]" italics="true" pageId="4" pageNumber="3">Dasyprocta punctata</emphasis>
|
|||
|
|
Gray
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFD69E2E6FCF948C9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[643,881,409,433]" class="Mammalia" family="Felidae" genus="Leopardus" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="3" phylum="Chordata" rank="species" species="pardalis">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFD69E2E6FCF948C9" box="[643,881,409,433]" italics="true" pageId="4" pageNumber="3">Leopardus pardalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
L.,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFC77E2E5FB8048C9" box="[925,1032,410,433]" class="Mammalia" family="Cervidae" genus="Mazama" kingdom="Animalia" order="Artiodactyla" pageId="4" pageNumber="3" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC77E2E5FB8048C9" box="[925,1032,410,433]" italics="true" pageId="4" pageNumber="3">Mazama</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
spp.,
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFBA6E2E5FB7948C9" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[1100,1265,410,433]" class="Mammalia" family="Procyonidae" genus="Nasua" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="3" phylum="Chordata" rank="species" species="narica">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFBA6E2E5FB7948C9" box="[1100,1265,410,433]" italics="true" pageId="4" pageNumber="3">Nasua narica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(L.),
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFAC5E2E6FF7048A9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Felidae" genus="Panthera" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="3" phylum="Chordata" rank="species" species="onca">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFAC5E2E6FF7048A9" italics="true" pageId="4" pageNumber="3">Panthera onca</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(L.),
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFED3E2C5FE6548A9" baseAuthorityName="Linnaeus" baseAuthorityYear="1771" box="[313,493,441,465]" class="Mammalia" family="Felidae" genus="Puma" kingdom="Animalia" order="Carnivora" pageId="4" pageNumber="3" phylum="Chordata" rank="species" species="concolor">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFED3E2C5FE6548A9" box="[313,493,441,465]" italics="true" pageId="4" pageNumber="3">Puma concolor</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(L.),
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFDC4E2C5FCBC48A9" authority="(Gill)" baseAuthorityName="Gill" baseAuthorityYear="1865" box="[558,820,441,465]" class="Mammalia" family="Tapiridae" genus="Tapirus" kingdom="Animalia" order="Perissodactyla" pageId="4" pageNumber="3" phylum="Chordata" rank="species" species="bairdii">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFDC4E2C5FD6248A9" box="[558,746,441,465]" italics="true" pageId="4" pageNumber="3">Tapirus bairdii</emphasis>
|
|||
|
|
(Gill)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFC9FE2C5FB0C48A9" authority="(Link)" baseAuthorityName="Link" baseAuthorityYear="1795" box="[885,1156,441,465]" class="Mammalia" family="Tayassuidae" genus="Tayassu" kingdom="Animalia" order="Artiodactyla" pageId="4" pageNumber="3" phylum="Chordata" rank="species" species="pecari">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC9FE2C5FBA548A9" box="[885,1069,442,465]" italics="true" pageId="4" pageNumber="3">Tayassu pecari</emphasis>
|
|||
|
|
(Link)
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="9BFD650BFFE9210FFF2AE296FAA04B59" pageId="4" pageNumber="3" type="etymology">
|
|||
|
|
<paragraph id="D3583680FFE9210FFF2AE296FAA04B59" blockId="4.[192,1440,489,546]" pageId="4" pageNumber="3">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFF2AE296FED34B79" bold="true" box="[192,347,489,513]" pageId="4" pageNumber="3">Etymology.</emphasis>
|
|||
|
|
The specific epithet is a reference to the size of the new species. Cirulito, applied here as a masculine noun in apposition, is the colloquial name for any very small thing in
|
|||
|
|
<collectingCountry id="ABF07610FFE9210FFB7DE176FAAB4B59" box="[1175,1315,521,545]" name="Guatemala" pageId="4" pageNumber="3">Guatemala</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="9BFD650BFFE9210FFF2AE146FA294A29" pageId="4" pageNumber="3" type="discussion">
|
|||
|
|
<paragraph id="D3583680FFE9210FFF2AE146FA034B89" blockId="4.[192,1441,569,850]" pageId="4" pageNumber="3">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFF2AE146FED14B29" bold="true" box="[192,345,569,593]" pageId="4" pageNumber="3">Comments.</emphasis>
|
|||
|
|
The presence of a ventral longitudinal groove on the metafemur and the two strong apical clypeal teeth place
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFE43E126FD194B09" authority="Cano, 2018" authorityName="Cano" authorityYear="2018" box="[425,657,601,625]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="cirulito" status="sp. nov.">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFE43E126FD194B09" box="[425,657,601,625]" italics="true" pageId="4" pageNumber="3">Anomiopus cirulito</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="FAA057E9FFE9210FFD72E125FD514B09" box="[664,729,602,625]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
near
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFCF4E126FC594B09" authorityName="in Canhedo" authorityYear="2006" box="[798,977,601,625]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="panamensis">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFCF4E126FC594B09" box="[798,977,601,625]" italics="true" pageId="4" pageNumber="3">A. panamensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="4BDC2A05FFE9210FFC0AE126FBAE4B0A" box="[992,1062,601,626]" captionStart="Figure 2" captionStartId="7.[192,274,1877,1899]" captionTargetBox="[218,1419,227,1843]" captionTargetId="figure@7.[213,1419,213,1845]" captionTargetPageId="7" captionText="Figure 2. Anomiopus panamensis. a) Dorsal habitus. b) Ventral habitus. c) Lateral view. d) Details of metasternum in latero-vental view. CxII = coxa II; Mt, metasternum; Mtp = metepisternum; Ms = mesepimeron." figureDoi="http://doi.org/10.5281/zenodo.3709951" httpUri="https://zenodo.org/record/3709951/files/figure.png" pageId="4" pageNumber="3">Fig. 2</figureCitation>
|
|||
|
|
), but it can be easily separated by its larger size (
|
|||
|
|
<quantity id="141F9B65FFE9210FFE77E106FE6D4BEA" box="[413,485,633,658]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="3" unit="mm" value="5.0">5 mm</quantity>
|
|||
|
|
), the shallow and minute punctures on pronotum, and the smooth sternellum (prosternum), metasternum, and mesepimeron (
|
|||
|
|
<figureCitation id="4BDC2A05FFE9210FFCE0E1E6FCE74BCA" box="[778,879,665,690]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="8.[192,275,1877,1899]" captionStartId-1="9.[192,276,1877,1899]" captionTargetBox-0="[217,1416,213,1843]" captionTargetBox-1="[217,1416,213,1843]" captionTargetId-0="figure@8.[217,1416,213,1843]" captionTargetId-1="figure@9.[217,1416,213,1843]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figure 3. Anomiopus cirulito n. sp. a) Dorsal habitus. b) Ventral habitus. c) Lateral view." captionText-1="Figure 4. Anomiopus cirulito, n. sp. a) Head, pronotum and left tibia. b) Detail of right tibia. c) Details of metasternum in latero-ventral view. CxII = coxa II; Mt, metasternum; Mtp = metepisternum; Ms = mesepimeron." figureDoi-0="http://doi.org/10.5281/zenodo.3709953" figureDoi-1="http://doi.org/10.5281/zenodo.3709955" httpUri-0="https://zenodo.org/record/3709953/files/figure.png" httpUri-1="https://zenodo.org/record/3709955/files/figure.png" pageId="4" pageNumber="3">Fig. 3–4</figureCitation>
|
|||
|
|
). In
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFC49E1E6FBDE4BC9" authorityName="in Canhedo" authorityYear="2006" box="[931,1110,665,689]" class="Insecta" family="Scarabaeidae" genus="Onthocharis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="panamensis">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC49E1E6FBDE4BC9" box="[931,1110,665,689]" italics="true" pageId="4" pageNumber="3">A. panamensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
the size varies between 3.3 and
|
|||
|
|
<quantity id="141F9B65FFE9210FFF1CE1C5FEDC4BA9" box="[246,340,698,722]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="3" unit="mm" value="4.4">4.4 mm</quantity>
|
|||
|
|
, the punctures on pronotum are deeper and stronger (
|
|||
|
|
<figureCitation id="4BDC2A05FFE9210FFC06E1C6FBD54BAA" box="[1004,1117,697,722]" captionStart="Figure 2" captionStartId="7.[192,274,1877,1899]" captionTargetBox="[218,1419,227,1843]" captionTargetId="figure@7.[213,1419,213,1845]" captionTargetPageId="7" captionText="Figure 2. Anomiopus panamensis. a) Dorsal habitus. b) Ventral habitus. c) Lateral view. d) Details of metasternum in latero-vental view. CxII = coxa II; Mt, metasternum; Mtp = metepisternum; Ms = mesepimeron." figureDoi="http://doi.org/10.5281/zenodo.3709951" httpUri="https://zenodo.org/record/3709951/files/figure.png" pageId="4" pageNumber="3">Fig. 2a, c</figureCitation>
|
|||
|
|
), and notably the surfaces of the sternellum (prosternum), mesepimeron and sides of metasternum are micro-striated (
|
|||
|
|
<figureCitation id="4BDC2A05FFE9210FFACCE1A6FAF64B89" box="[1318,1406,729,753]" captionStart="Figure 2" captionStartId="7.[192,274,1877,1899]" captionTargetBox="[218,1419,227,1843]" captionTargetId="figure@7.[213,1419,213,1845]" captionTargetPageId="7" captionText="Figure 2. Anomiopus panamensis. a) Dorsal habitus. b) Ventral habitus. c) Lateral view. d) Details of metasternum in latero-vental view. CxII = coxa II; Mt, metasternum; Mtp = metepisternum; Ms = mesepimeron." figureDoi="http://doi.org/10.5281/zenodo.3709951" httpUri="https://zenodo.org/record/3709951/files/figure.png" pageId="4" pageNumber="3">Fig. 2d</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="D3583680FFE9210FFF02E186FA294A29" blockId="4.[192,1441,569,850]" pageId="4" pageNumber="3">
|
|||
|
|
Within the cuprarius group the new species belongs to a small group of three species sharing the presence of two small conical processes on the frons. Accordingly, the following changes to the key of species of the cuprarius group of
|
|||
|
|
<bibRefCitation id="B7764B71FFE9210FFDACE046FCB44A29" author="Canhedo, V. L." box="[582,828,825,849]" pageId="4" pageNumber="3" pagination="349 - 502" refId="ref3161" refString="Canhedo, V. L. 2006. Revisao taxonomica do genero Anomiopus Westwood 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Arquivos de Zoologia 37 (4): 349 - 502." type="journal article" year="2006">Canhedo (2006: 359)</bibRefCitation>
|
|||
|
|
must be inserted to accommodate this new species:
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="9BFD650BFFE9210FFF2AE015FA284C79" pageId="4" pageNumber="3" type="key">
|
|||
|
|
<key id="A77661B1FFE9210FFF2AE015FA284C79" pageId="4" pageNumber="3">
|
|||
|
|
<keyStep id="68132E25FFE9210FFF2AE015FA284AB9" pageId="4" pageNumber="3">
|
|||
|
|
<paragraph id="D3583680FFE9210FFF2AE015FA284AF9" blockId="4.[192,1441,873,1281]" box="[192,1440,873,898]" pageId="4" pageNumber="3">
|
|||
|
|
<keyLead id="681695B5FFE9210FFF2AE015FA284AF9" box="[192,1440,873,898]" pageId="4" pageNumber="3">
|
|||
|
|
1. Frons with two conical processes at ends of frontoclypeal suture........................
|
|||
|
|
<emphasis id="E193EA92FFE9210FFA7BE016FA284AF9" bold="true" box="[1425,1440,873,897]" pageId="4" pageNumber="3">2</emphasis>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="D3583680FFE9210FFF2AE0F5FA284AB9" blockId="4.[192,1441,873,1281]" pageId="4" pageNumber="3">
|
|||
|
|
<keyLead id="681695B5FFE9210FFF2AE0F5FA284AB9" pageId="4" pageNumber="3">
|
|||
|
|
— Frons without conical processes.....................................................
|
|||
|
|
<emphasis id="E193EA92FFE9210FFEABE0D6FA284AB9" bold="true" box="[321,1440,937,961]" pageId="4" pageNumber="3">
|
|||
|
|
(to couplet 3 of the key to species of the cuprarius group in
|
|||
|
|
<bibRefCitation id="B7764B71FFE9210FFB69E0D6FA064AB9" author="Canhedo, V. L." box="[1155,1422,937,961]" pageId="4" pageNumber="3" pagination="349 - 502" refId="ref3161" refString="Canhedo, V. L. 2006. Revisao taxonomica do genero Anomiopus Westwood 1842 (Coleoptera, Scarabaeidae, Scarabaeinae). Arquivos de Zoologia 37 (4): 349 - 502." type="journal article" year="2006">Canhedo (2006: 359</bibRefCitation>
|
|||
|
|
])
|
|||
|
|
</emphasis>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="68132E25FFE9210FFF2AE0B5FA284DF9" pageId="4" pageNumber="3">
|
|||
|
|
<paragraph id="D3583680FFE9210FFF2AE0B5FA284D59" blockId="4.[192,1441,873,1281]" pageId="4" pageNumber="3">
|
|||
|
|
<keyLead id="681695B5FFE9210FFF2AE0B5FA284D59" pageId="4" pageNumber="3">
|
|||
|
|
2(1). Ventral surface of metafemur without anterior longitudinal groove; sternellum and metepisternum smooth, mesepimeron and sides of metasternum finely micro-striated.
|
|||
|
|
<collectingCountry id="ABF07610FFE9210FFED2E776FE234D59" box="[312,427,1033,1057]" name="Colombia" pageId="4" pageNumber="3">Colombia</collectingCountry>
|
|||
|
|
.......................................
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFC16E776FA284D59" ID-CoL="676L9" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[1020,1440,1033,1057]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="cuprarius">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC16E776FA284D59" bold="true" box="[1020,1440,1033,1057]" pageId="4" pageNumber="3">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC16E776FAAB4D59" bold="true" box="[1020,1315,1033,1057]" italics="true" pageId="4" pageNumber="3">Anomiopus cuprarius</emphasis>
|
|||
|
|
(Harold)
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="D3583680FFE9210FFF2AE755FA284DF9" blockId="4.[192,1441,873,1281]" pageId="4" pageNumber="3">
|
|||
|
|
<keyLead id="681695B5FFE9210FFF2AE755FA284DF9" pageId="4" pageNumber="3">
|
|||
|
|
— Ventral surface of metafemur with anterior longitudinal groove; sternellum, mesepimeron, metepisternum and sides of metasternum smooth or micro-striated. Central America,
|
|||
|
|
<collectingCountry id="ABF07610FFE9210FFED2E716FE234DF9" box="[312,427,1129,1153]" name="Colombia" pageId="4" pageNumber="3">Colombia</collectingCountry>
|
|||
|
|
...................................................................
|
|||
|
|
<emphasis id="E193EA92FFE9210FFA7BE716FA284DF9" bold="true" box="[1425,1440,1129,1153]" pageId="4" pageNumber="3">3</emphasis>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="68132E25FFE9210FFF2AE7F5FA284C79" pageId="4" pageNumber="3">
|
|||
|
|
<paragraph id="D3583680FFE9210FFF2AE7F5FA284DBA" blockId="4.[192,1441,873,1281]" pageId="4" pageNumber="3">
|
|||
|
|
<keyLead id="681695B5FFE9210FFF2AE7F5FA284DBA" pageId="4" pageNumber="3">
|
|||
|
|
3(2). Sternellum, mesepimeron, metepisternum and sides of metasternum finely micro-striated;
|
|||
|
|
<quantity id="141F9B65FFE9210FFAB4E7F5FEEF4DB9" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="4" pageNumber="3" unit="mm" value="3.65" valueMax="4.0" valueMin="3.3">3.3–4 mm</quantity>
|
|||
|
|
in length.
|
|||
|
|
<collectingCountry id="ABF07610FFE9210FFE06E7D5FDFA4DB9" box="[492,626,1193,1218]" name="Costa Rica" pageId="4" pageNumber="3">Costa Rica</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="ABF07610FFE9210FFD97E7D5FD6D4DB9" box="[637,741,1194,1217]" name="Panama" pageId="4" pageNumber="3">Panama</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="ABF07610FFE9210FFD18E7D6FCED4DB9" box="[754,869,1193,1217]" name="Colombia" pageId="4" pageNumber="3">Colombia</collectingCountry>
|
|||
|
|
......
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFC3CE7D6FA284DBA" ID-CoL="5V6MH" authority="(Paulian)" baseAuthorityName="Paulian" baseAuthorityYear="1939" box="[982,1440,1193,1218]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="panamensis">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC3CE7D6FA1F4DB9" bold="true" box="[982,1431,1193,1217]" pageId="4" pageNumber="3">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC3CE7D6FA924DB9" bold="true" box="[982,1306,1193,1217]" italics="true" pageId="4" pageNumber="3">Anomiopus panamensis</emphasis>
|
|||
|
|
(Paulian
|
|||
|
|
</emphasis>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="D3583680FFE9210FFF2AE7B5FA284C79" blockId="4.[192,1441,873,1281]" pageId="4" pageNumber="3">
|
|||
|
|
<keyLead id="681695B5FFE9210FFF2AE7B5FA284C79" pageId="4" pageNumber="3">
|
|||
|
|
— Sternellum, mesepimeron, metepisternum, and metasternum smooth, without micro-striations;
|
|||
|
|
<quantity id="141F9B65FFE9210FFED2E796FEF54C79" box="[312,381,1257,1281]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="3" unit="mm" value="5.0">5 mm</quantity>
|
|||
|
|
in length.
|
|||
|
|
<collectingCountry id="ABF07610FFE9210FFDE8E796FD034C79" box="[514,651,1257,1281]" name="Guatemala" pageId="4" pageNumber="3">Guatemala</collectingCountry>
|
|||
|
|
........................
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC16E796FA284C79" bold="true" box="[1020,1440,1257,1281]" pageId="4" pageNumber="3">
|
|||
|
|
<taxonomicName id="14E74D03FFE9210FFC16E796FAD94C79" ID-CoL="676KX" authority="Cano" authorityName="Cano" authorityYear="2018" box="[1020,1361,1257,1281]" class="Insecta" family="Scarabaeidae" genus="Anomiopus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="cirulito" status="sp. nov.">
|
|||
|
|
<emphasis id="E193EA92FFE9210FFC16E796FA8A4C79" bold="true" box="[1020,1282,1257,1281]" italics="true" pageId="4" pageNumber="3">Anomiopus cirulito</emphasis>
|
|||
|
|
Cano
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="FAA057E9FFE9210FFAB3E796FA284C79" box="[1369,1440,1257,1281]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
</key>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|