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Hughes, Kusamba, Menegon, and Greenbaum
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<paragraph id="CB2985571E7AFFEA72D065C70D67FF48" blockId="29.[151,1199,151,250]" box="[151,520,187,214]" pageId="29" pageNumber="480">Albertine Rift pygmy chameleon</paragraph>
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<emphasis id="F9E259451E7AFFEA72D0645B0E61FEDC" box="[151,270,295,321]" italics="true" pageId="29" pageNumber="480">Synonymy</emphasis>
.
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<emphasis id="F9E259451E7AFFEA72D064100EDFFE19" box="[151,432,364,388]" italics="true" pageId="29" pageNumber="480">
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—
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<bibRefCitation id="AF07F8A61E7AFFEA73E864100D50FE1E" author="de Witte" box="[431,575,363,388]" firstAuthor="de Witte" pageId="29" pageNumber="480" pagination="1 - 215" refId="ref27886" refString="de Witte, G. - F. (1965) Les cameleons de l'Afrique Centrale (Republique Democratique du Congo, Republique du Rwanda et Royaume du Burundi). Annales Musee Royal de l'Afrique Centrale, Sciences Zoologiques, 142, 1 - 215, pls. I - XII." type="journal article" year="1965">de Witte 1965</bibRefCitation>
(partim), 
<bibRefCitation id="AF07F8A61E7AFFEA70E164100CE3FE1E" author="Tilbury &amp; Tolley" box="[678,908,363,388]" firstAuthor="Tilbury" pageId="29" pageNumber="480" pagination="345 - 364" refId="ref32364" refString="Tilbury, C. R. &amp; Tolley, K. A. (2015) Contributions to the herpetofauna of the Albertine Rift: two new species of chameleon (Sauria: Chamaeleonidae) from an isolated montane forest, south eastern Democratic Republic of Congo. Zootaxa, 3905 (3), 345 - 364. https: // doi. org / 10.11646 / zootaxa. 3905.3.2" type="journal article" year="2015">Tilbury &amp; Tolley 2015</bibRefCitation>
(partim), 
<bibRefCitation id="AF07F8A61E7AFFEA71B464100BDEFE19" author="Spawls" box="[1011,1201,364,388]" etAl="et al." firstAuthor="Spawls" pageId="29" pageNumber="480" refId="ref31894" refString="Spawls, S., Howell, K., Drewes, R. C. &amp; Ashe, J. (2018) Field Guide to East African Reptiles. 2 nd Edition. Bloomsbury Natural History, Oxford, 624 pp." type="book" year="2018">
Spawls 
<emphasis id="F9E259451E7AFFEA760364100B1EFE19" box="[1092,1137,364,388]" italics="true" pageId="29" pageNumber="480">et al</emphasis>
. 2018
</bibRefCitation>
(partim), 
<bibRefCitation id="AF07F8A61E7AFFEA775064100AF2FE19" author="Tilbury" box="[1303,1437,364,388]" firstAuthor="Tilbury" pageId="29" pageNumber="480" refId="ref32323" refString="Tilbury, C. R. (2018) Chameleons of Africa: An Atlas Including the Chameleons of Europe, the Middle East and Asia. 2 nd Edition. Edition Chimaira, Frankfurt am Main, 643 pp." type="book" year="2018">Tilbury 2018</bibRefCitation>
(partim)
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<taxonomicName id="0C96FED41E7AFFEA72D064D50E36FE5C" box="[151,345,425,449]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="undefined-4">
<emphasis id="F9E259451E7AFFEA72D064D50E4BFE5C" box="[151,292,425,449]" italics="true" pageId="29" pageNumber="480">Rhampholeon</emphasis>
sp. 4
</taxonomicName>
— 
<bibRefCitation id="AF07F8A61E7AFFEA733364D50D5CFE5C" author="Hughes" box="[372,563,425,449]" etAl="et al." firstAuthor="Hughes" pageId="29" pageNumber="480" pagination="125 - 141" refId="ref29435" refString="Hughes, D. F., Tolley, K. A., Behangana, M., Lukwago, W., Menegon, M., Dehling, J. M., Stipala, J., Tilbury, C. R., Khan, A. M., Kusamba, C. &amp; Greenbaum, E. (2018) Cryptic diversity in Rhampholeon boulengeri (Sauria: Chamaeleonidae), a pygmy chameleon from the Albertine Rift biodiversity hotspot. Molecular Phylogenetics and Evolution, 122, 125 - 141. https: // doi. org / 10.1016 / j. ympev. 2017.11.015" type="journal article" year="2018">
Hughes 
<emphasis id="F9E259451E7AFFEA738064D60E9BFE5C" box="[455,500,425,449]" italics="true" pageId="29" pageNumber="480">et al</emphasis>
. 2018
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<emphasis id="F9E259451E7AFFEA72D064930E7AFD94" box="[151,277,495,521]" italics="true" pageId="29" pageNumber="480">Etymology.</emphasis>
The specific epithet is derived from the Swahili word for forest, 
<emphasis id="F9E259451E7AFFEA7644648C0B2FFD94" box="[1027,1088,496,521]" italics="true" pageId="29" pageNumber="480">msitu</emphasis>
, and the German word for rift valley, 
<emphasis id="F9E259451E7AFFEA72AF676F0E58FDB0" box="[232,311,531,557]" italics="true" pageId="29" pageNumber="480">graben</emphasis>
, in reference to the Albertine Rift forests where the species occurs.
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<specimenCode id="9B302D2C1E7AFFEA737867440EBBFDCC" box="[319,468,567,594]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="480" type="Museum">UTEP 21709</specimenCode>
(field no. 
<specimenCode id="9B302D2C1E7AFFEA701567440DACFDCF" box="[594,707,567,594]" collectionCode="ELI" pageId="29" pageNumber="480">ELI 1159</specimenCode>
), adult female, 
<collectingCountry id="B381C5C71E7AFFEA713E67440C94FDCF" box="[889,1019,568,594]" name="Burundi" pageId="29" pageNumber="480">BURUNDI</collectingCountry>
, 
<collectingRegion id="09524BB51E7AFFEA764E674B0BB4FDCC" box="[1033,1243,567,593]" country="Burundi" name="Bubanza" pageId="29" pageNumber="480">Bubanza Province</collectingRegion>
, 
<collectingMunicipality id="2B4D1F2D1E7AFFEA76AD674B0FA2FDE8" pageId="29" pageNumber="480">Kibira National Park</collectingMunicipality>
, 
<location id="CE49D38C1E7AFFEA729C67270E42FDE8" LSID="urn:lsid:plazi:treatment:433F34411E7AFFE872D065EB0E43FE68:CE49D38C1E7AFFEA729C67270E42FDE8" box="[219,301,603,629]" country="Burundi" latitude="-3.06177" longLatPrecision="1" longitude="29.49343" municipality="Kibira National Park" name="Mpishi" pageId="29" pageNumber="480" stateProvince="Bubanza">Mpishi</location>
, 
<geoCoordinate id="AEA2E3901E7AFFEA737C67270EA5FDE8" box="[315,458,603,629]" degrees="03.06177" direction="south" orientation="latitude" pageId="29" pageNumber="480" precision="1" value="-3.06177">03.06177° S</geoCoordinate>
, 
<geoCoordinate id="AEA2E3901E7AFFEA739E67270D06FDEB" box="[473,617,603,630]" degrees="29.49343" direction="east" orientation="longitude" pageId="29" pageNumber="480" precision="1" value="29.49343">29.49343° E</geoCoordinate>
, 
<quantity id="0C6E28B21E7AFFEA703F67270DBFFDEB" box="[632,720,603,630]" metricMagnitude="3" metricUnit="m" metricValue="1.986" pageId="29" pageNumber="480" unit="m" value="1986.0">
<elevation id="40BB62641E7AFFEA703F67270DBFFDEB" box="[632,720,603,630]" metricMagnitude="3" metricUnit="m" metricValue="1.986" pageId="29" pageNumber="480" unit="m" value="1986.0">1986 m</elevation>
</quantity>
elevation, 
<date id="BF28A3971E7AFFEA711767270B43FDE8" box="[848,1068,603,629]" pageId="29" pageNumber="480" value="2011-12-20">
<collectingDate id="AF6C5A7F1E7AFFEA711767270B43FDE8" box="[848,1068,603,629]" pageId="29" pageNumber="480" value="2011-12-20">20 December 2011</collectingDate>
</date>
, collected by 
<collectorName id="6663E0811E7AFFEA768967200A1AFDE8" box="[1230,1397,603,630]" pageId="29" pageNumber="480">E. Greenbaum</collectorName>
, C. 
<collectorName id="6663E0811E7AFFEA72D067030E24FD07" box="[151,331,639,666]" pageId="29" pageNumber="480">Kusamba, M.M</collectorName>
. Aristote, and 
<collectorName id="6663E0811E7AFFEA73AC67FC0DF9FD04" box="[491,662,639,666]" pageId="29" pageNumber="480">W.M. Muninga</collectorName>
(
<figureCitation id="53AD99D21E7AFFEA70E267030C61FD04" box="[677,782,639,665]" captionStart="FIGURE 13" captionStartId="29.[151,250,1983,2007]" captionTargetBox="[410,1178,1445,1959]" captionTargetId="figure-470@29.[410,1178,1445,1959]" captionTargetPageId="29" captionText="FIGURE 13. Photograph of Rhampholeon msitugrabensis sp. nov. in life. (A)—Adult female holotype (UTEP 21709) from Kibira National Park, Burundi." figureDoi="http://doi.org/10.5281/zenodo.11547891" httpUri="https://zenodo.org/record/11547891/files/figure.png" pageId="29" pageNumber="480">Fig. 13A</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="CB2985571E7AFFEA728067D80C6FFD7C" blockId="29.[151,1437,495,1421]" pageId="29" pageNumber="480">
<materialsCitation id="7BFE8F0A1E7AFFEA728067D80EAEFD7C" ID-GBIF-Occurrence="4879928335" collectionCode="UTEP" pageId="29" pageNumber="480" specimenCode="UTEP 22739-22743" specimenCount="5" specimenCount-adult="4" specimenCount-juvenile="1" typeStatus="paratype">
<emphasis id="F9E259451E7AFFEA728067D80EC0FD20" box="[199,431,675,701]" italics="true" pageId="29" pageNumber="480">
<typeStatus id="142D3BF51E7AFFEA728067D80E41FD20" box="[199,302,676,701]" pageId="29" pageNumber="480" type="paratype">Paratype</typeStatus>
(
<typeStatus id="142D3BF51E7AFFEA737A67DF0ECDFD20" box="[317,418,675,701]" pageId="29" pageNumber="480" type="topotype">topotype</typeStatus>
).
</emphasis>
Same collection details as 
<typeStatus id="142D3BF51E7AFFEA709967DF0C2CFD20" box="[734,835,675,701]" pageId="29" pageNumber="480" type="holotype">holotype</typeStatus>
, 
<specimenCount id="DD904EDE1E7AFFEA710967DF0CAAFD20" box="[846,965,675,701]" count="3" pageId="29" pageNumber="480" type="adult">three adult</specimenCount>
females, 
<specimenCount id="DD904EDE1E7AFFEA767667D80BF7FD20" box="[1073,1176,675,702]" count="1" pageId="29" pageNumber="480" type="adult">one adult</specimenCount>
male, and 
<specimenCount id="DD904EDE1E7AFFEA775467D80AF3FD20" box="[1299,1436,675,702]" count="1" pageId="29" pageNumber="480" type="juvenile">one juvenile</specimenCount>
male, 
<specimenCode id="9B302D2C1E7AFFEA729B67B40EAEFD7C" box="[220,449,711,738]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="480" type="Museum">UTEP 22739–22743</specimenCode>
</materialsCitation>
<materialsCitation id="7BFE8F0A1E7AFFEA738E67BB0D92FD7C" ID-GBIF-Occurrence="4879928306" box="[457,765,711,738]" collectionCode="ELI" isTailEnumeration="true" pageId="29" pageNumber="480" specimenCode="ELI 1160-1164" specimenCount="1" typeStatus="paratype">
(field nos. 
<specimenCode id="9B302D2C1E7AFFEA700667B40D9AFD7C" box="[577,757,711,738]" collectionCode="ELI" pageId="29" pageNumber="480">ELI 1160–1164</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CB2985571E7AFFEA728067970C50FCB4" blockId="29.[151,1437,495,1421]" pageId="29" pageNumber="480">
<emphasis id="F9E259451E7AFFEA728067970EF2FC98" box="[199,413,747,773]" italics="true" pageId="29" pageNumber="480">Referred specimen</emphasis>
. 
<materialsCitation id="7BFE8F0A1E7AFFEA73F667970C53FCB4" ID-GBIF-Occurrence="4879928302" collectionCode="MTSN" country="Rwanda" elevation="2669" latitude="-2.45565" location="Mount Bigugu" longLatPrecision="1" longitude="29.24927" municipality="Nyungwe National Park" pageId="29" pageNumber="480" specimenCode="MTSN 7213" specimenCount="1" stateProvince="Ouest">
<specimenCode id="9B302D2C1E7AFFEA73F667970D28FC98" box="[433,583,747,773]" collectionCode="MTSN" pageId="29" pageNumber="480">MTSN 7213</specimenCode>
, 
<collectingCountry id="B381C5C71E7AFFEA701F67970DBBFC98" box="[600,724,747,773]" name="Rwanda" pageId="29" pageNumber="480">RWANDA</collectingCountry>
, 
<collectingRegion id="09524BB51E7AFFEA70A367900CDDFC98" box="[740,946,747,774]" country="Rwanda" name="Ouest" pageId="29" pageNumber="480">Western Province</collectingRegion>
, 
<collectingMunicipality id="2B4D1F2D1E7AFFEA718367900B8EFC98" box="[964,1249,747,774]" pageId="29" pageNumber="480">Nyungwe National Park</collectingMunicipality>
, 
<location id="CE49D38C1E7AFFEA76B567900AF3FC98" LSID="urn:lsid:plazi:treatment:433F34411E7AFFE872D065EB0E43FE68:CE49D38C1E7AFFEA76B567900AF3FC98" box="[1266,1436,747,774]" country="Rwanda" latitude="-2.45565" longLatPrecision="1" longitude="29.24927" municipality="Nyungwe National Park" name="Mount Bigugu" pageId="29" pageNumber="480" stateProvince="Ouest">Mount Bigugu</location>
(
<geoCoordinate id="AEA2E3901E7AFFEA72D866730E42FCB4" box="[159,301,783,810]" degrees="02.45565" direction="south" orientation="latitude" pageId="29" pageNumber="480" precision="1" value="-2.45565">02.45565° S</geoCoordinate>
, 
<geoCoordinate id="AEA2E3901E7AFFEA737E66730EA8FCB7" box="[313,455,783,810]" degrees="29.24927" direction="east" orientation="longitude" pageId="29" pageNumber="480" precision="1" value="29.24927">29.24927° E</geoCoordinate>
, 
<quantity id="0C6E28B21E7AFFEA739366730D46FCB7" box="[468,553,783,810]" metricMagnitude="3" metricUnit="m" metricValue="2.6689999999999996" pageId="29" pageNumber="480" unit="m" value="2669.0">
<elevation id="40BB62641E7AFFEA739366730D46FCB7" box="[468,553,783,810]" metricMagnitude="3" metricUnit="m" metricValue="2.6689999999999996" pageId="29" pageNumber="480" unit="m" value="2669.0">2669 m</elevation>
</quantity>
elevation) (
<specimenCount id="DD904EDE1E7AFFEA70F666730C5BFCB4" box="[689,820,783,809]" count="1" pageId="29" pageNumber="480" type="generic">1 specimen</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="838CD6DC1E7AFFEA728066480EACFA10" pageId="29" pageNumber="480" type="diagnosis">
<paragraph id="CB2985571E7AFFEA728066480A01FABC" blockId="29.[151,1437,495,1421]" pageId="29" pageNumber="480">
<emphasis id="F9E259451E7AFFEA728066480DCBFCD3" box="[199,676,819,846]" italics="true" pageId="29" pageNumber="480">
Diagnosis. 
<taxonomicName id="0C96FED41E7AFFEA730E664F0DCBFCD3" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[329,676,819,846]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="msitugrabensis" status="sp. nov.">
<emphasis id="F9E259451E7AFFEA730E664F0DCBFCD3" bold="true" box="[329,676,819,846]" pageId="29" pageNumber="480">Rhampholeon msitugrabensis</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="E2D1E43E1E7AFFEA70E9664F0C65FCD0" box="[686,778,819,845]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
is in the subgenus 
<taxonomicName id="0C96FED41E7AFFEA71AB664F0BEBFCD0" authorityName="Matthee, Tilbury &amp; Townsend" authorityYear="2004" box="[1004,1156,819,845]" class="Squamata" family="Chamaeleonidae" genus="Rhinodigitum" kingdom="Animalia" pageId="29" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="F9E259451E7AFFEA71AB664F0BEBFCD0" box="[1004,1156,819,845]" italics="true" pageId="29" pageNumber="480">Rhinodigitum</emphasis>
</taxonomicName>
because of its distinctly bicuspid claws, prominent rostral process, smooth plantar surfaces, and phylogenetic placement, thus easily distinguishing it from the six species in the other two subgenera (i.e., 
<taxonomicName id="0C96FED41E7AFFEA71EF66070B2BFC08" authorityName="Gunther" authorityYear="1874" box="[936,1092,891,917]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="F9E259451E7AFFEA71EF66070B2BFC08" box="[936,1092,891,917]" italics="true" pageId="29" pageNumber="480">Rhampholeon</emphasis>
</taxonomicName>
and 
<taxonomicName id="0C96FED41E7AFFEA763C66000BB5FC08" authorityName="Loveridge" authorityYear="1956" box="[1147,1242,892,917]" genus="Bicuspis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="29" pageNumber="480" phylum="Chordata" rank="genus">
<emphasis id="F9E259451E7AFFEA763C66000BB5FC08" box="[1147,1242,892,917]" italics="true" pageId="29" pageNumber="480">Bicuspis</emphasis>
</taxonomicName>
): 
<taxonomicName id="0C96FED41E7AFFEA76B466000AFAFC08" authorityName="Broadley" authorityYear="1971" box="[1267,1429,892,917]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="gorongosae">
<emphasis id="F9E259451E7AFFEA76B466000AFAFC08" box="[1267,1429,892,917]" italics="true" pageId="29" pageNumber="480">R. gorongosae</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA72D066DC0E4CFC24" authorityName="Boulenger" authorityYear="1906" box="[151,291,927,953]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="marshalli">
<emphasis id="F9E259451E7AFFEA72D066DC0E4CFC24" box="[151,291,927,953]" italics="true" pageId="29" pageNumber="480">R. marshalli</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA737466DC0ED4FC24" baseAuthorityName="Buchholz" baseAuthorityYear="1874" box="[307,443,928,953]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="spectrum">
<emphasis id="F9E259451E7AFFEA737466DC0ED4FC24" box="[307,443,928,953]" italics="true" pageId="29" pageNumber="480">R. spectrum</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA738C66DC0D23FC24" baseAuthorityName="Matschie" baseAuthorityYear="1892" box="[459,588,928,953]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="F9E259451E7AFFEA738C66DC0D23FC24" box="[459,588,928,953]" italics="true" pageId="29" pageNumber="480">R. spinosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA701B66DC0D9AFC24" baseAuthorityName="Matschie" baseAuthorityYear="1892" box="[604,757,927,953]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="temporalis">
<emphasis id="F9E259451E7AFFEA701B66DC0D9AFC24" box="[604,757,927,953]" italics="true" pageId="29" pageNumber="480">R. temporalis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="0C96FED41E7AFFEA717166DC0CF0FC24" authorityName="Mariaux &amp; Tilbury" authorityYear="2006" box="[822,927,927,953]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="viridis">
<emphasis id="F9E259451E7AFFEA717166DC0CF0FC24" box="[822,927,927,953]" italics="true" pageId="29" pageNumber="480">R. viridis</emphasis>
</taxonomicName>
. 
<emphasis id="F9E259451E7AFFEA71E966E30A02FC24" bold="true" box="[942,1389,927,954]" pageId="29" pageNumber="480">
<taxonomicName id="0C96FED41E7AFFEA71E966E30A67FC27" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[942,1288,927,954]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="msitugrabensis" status="sp. nov.">
<emphasis id="F9E259451E7AFFEA71E966E30A67FC27" bold="true" box="[942,1288,927,954]" italics="true" pageId="29" pageNumber="480">Rhampholeon msitugrabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E7AFFEA775666E30A02FC24" box="[1297,1389,927,953]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all other 
<taxonomicName id="0C96FED41E7AFFEA73B066BF0D84FC40" box="[503,747,963,989]" pageId="29" pageNumber="480">
<emphasis id="F9E259451E7AFFEA73B066BF0DFCFC40" box="[503,659,963,989]" italics="true" pageId="29" pageNumber="480">Rhampholeon</emphasis>
species
</taxonomicName>
by the following combination of traits: (1) lack of prominent mite pockets in the inguinal region distinguishes it from 
<taxonomicName id="0C96FED41E7AFFEA714266940CF4FB9C" authorityName="Mariaux &amp; Tilbury" authorityYear="2006" box="[773,923,999,1025]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="beraduccii">
<emphasis id="F9E259451E7AFFEA714266940CF4FB9C" box="[773,923,999,1025]" italics="true" pageId="29" pageNumber="480">R. beraduccii</emphasis>
</taxonomicName>
, 
<emphasis id="F9E259451E7AFFEA71E066940E86FBB8" italics="true" pageId="29" pageNumber="480">
<taxonomicName id="0C96FED41E7AFFEA71E066940B5DFB9C" authorityName="Gunther" authorityYear="1893" box="[935,1074,999,1025]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="platyceps">R. platyceps</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA767C66940B91FB9C" authorityName="Tilbury" authorityYear="1992" box="[1083,1278,999,1025]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="chapmanorum">R. chapmanorum</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA774066940AF7FB9C" authorityName="Branch, Bayliss &amp; Tolley" authorityYear="2014" box="[1287,1432,1000,1025]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="maspictus">R. maspictus</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA72D061700E61FBB8" authorityName="Branch, Bayliss &amp; Tolley" authorityYear="2014" box="[151,270,1035,1061]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="tilburyi">R. tilburyi</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA735B61700E8AFBB8" authorityName="Branch, Bayliss &amp; Tolley" authorityYear="2014" box="[284,485,1035,1061]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="bruessoworum">R. bruessoworum</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="0C96FED41E7AFFEA706361700DA2FBB8" authorityName="Branch, Bayliss &amp; Tolley" authorityYear="2014" box="[548,717,1035,1061]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="nebulauctor">
<emphasis id="F9E259451E7AFFEA706361700DA2FBB8" box="[548,717,1035,1061]" italics="true" pageId="29" pageNumber="480">R. nebulauctor</emphasis>
</taxonomicName>
; (2) presence of prominent mite pockets in the axillary region distinguishes it from 
<taxonomicName id="0C96FED41E7AFFEA73C1614C0D4EFBD4" baseAuthorityName="Loveridge" baseAuthorityYear="1953" box="[390,545,1071,1097]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="nchisiensis">
<emphasis id="F9E259451E7AFFEA73C1614C0D4EFBD4" box="[390,545,1071,1097]" italics="true" pageId="29" pageNumber="480">R. nchisiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="0C96FED41E7AFFEA701E614C0D95FBD4" authorityName="Mariaux &amp; Tilbury" authorityYear="2006" box="[601,762,1072,1097]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="F9E259451E7AFFEA701E614C0D95FBD4" box="[601,762,1072,1097]" italics="true" pageId="29" pageNumber="480">R. acuminatus</emphasis>
</taxonomicName>
; (3) distinct supra-orbital and canthal crests distinguishes it from 
<taxonomicName id="0C96FED41E7AFFEA729261280E35FBF0" authorityName="Tilbury &amp; Tolley" authorityYear="2015" box="[213,346,1107,1133]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="hattinghi">
<emphasis id="F9E259451E7AFFEA729261280E35FBF0" box="[213,346,1107,1133]" italics="true" pageId="29" pageNumber="480">R. hattinghi</emphasis>
</taxonomicName>
; (4) geographic restriction to the Albertine Rift distinguishes it from 
<taxonomicName id="0C96FED41E7AFFEA762E61280A76FBF0" authorityName="Tilbury &amp; Emmrich" authorityYear="1996" box="[1129,1305,1107,1133]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="uluguruensis">
<emphasis id="F9E259451E7AFFEA762E61280A76FBF0" box="[1129,1305,1107,1133]" italics="true" pageId="29" pageNumber="480">R. uluguruensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA776F61280AFAFBF0" authorityName="Menegon, Salvidio &amp; Tilbury" authorityYear="2002" box="[1320,1429,1108,1133]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="moyeri">
<emphasis id="F9E259451E7AFFEA776F61280AFAFBF0" box="[1320,1429,1108,1133]" italics="true" pageId="29" pageNumber="480">R. moyeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA72D061040E4FFB0C" box="[151,288,1143,1169]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="colemani">
<emphasis id="F9E259451E7AFFEA72D061040E4FFB0C" box="[151,288,1143,1169]" italics="true" pageId="29" pageNumber="480">R. colemani</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA737761040EF9FB0C" box="[304,406,1143,1169]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="sabini">
<emphasis id="F9E259451E7AFFEA737761040EF9FB0C" box="[304,406,1143,1169]" italics="true" pageId="29" pageNumber="480">R. sabini</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA73E161040D78FB0C" box="[422,535,1143,1169]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="rubeho">
<emphasis id="F9E259451E7AFFEA73E161040D78FB0C" box="[422,535,1143,1169]" italics="true" pageId="29" pageNumber="480">R. rubeho</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA706F61040DF8FB0C" box="[552,663,1143,1169]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="nicolai">
<emphasis id="F9E259451E7AFFEA706F61040DF8FB0C" box="[552,663,1143,1169]" italics="true" pageId="29" pageNumber="480">R. nicolai</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E7AFFEA70EF61040C3AFB0C" box="[680,853,1143,1169]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="waynelotteri">
<emphasis id="F9E259451E7AFFEA70EF61040C3AFB0C" box="[680,853,1143,1169]" italics="true" pageId="29" pageNumber="480">R. waynelotteri</emphasis>
</taxonomicName>
, and 
<taxonomicName id="0C96FED41E7AFFEA71DF61040B4CFB0C" box="[920,1059,1144,1169]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="princeeai">
<emphasis id="F9E259451E7AFFEA71DF61040B4CFB0C" box="[920,1059,1144,1169]" italics="true" pageId="29" pageNumber="480">R. princeeai</emphasis>
</taxonomicName>
; (5) slightly smaller head height in males, genetic divergence, and allopatric distribution distinguishes it from 
<taxonomicName id="0C96FED41E7AFFEA764961E00BC7FB28" authorityName=", Steindachner" authorityYear="1911" box="[1038,1192,1179,1205]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="F9E259451E7AFFEA764961E00BC7FB28" box="[1038,1192,1179,1205]" italics="true" pageId="29" pageNumber="480">R. boulengeri</emphasis>
</taxonomicName>
; (6) larger inter-limb lengths in both sexes distinguishes it from 
<emphasis id="F9E259451E7AFFEA703361C30CE7FB44" bold="true" box="[628,904,1215,1242]" pageId="29" pageNumber="480">
<taxonomicName id="0C96FED41E7AFFEA703361C30C47FB47" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[628,808,1215,1242]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="monteslunae" status="sp. nov.">
<emphasis id="F9E259451E7AFFEA703361C30C47FB47" bold="true" box="[628,808,1215,1242]" italics="true" pageId="29" pageNumber="480">R. monteslunae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E7AFFEA716861C30CE7FB44" box="[815,904,1215,1241]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (7) shorter inter-limb length, hind limb length, and head length in females distinguishes it from 
<emphasis id="F9E259451E7AFFEA70F3619F0CC8FB60" bold="true" box="[692,935,1251,1278]" pageId="29" pageNumber="480">
<taxonomicName id="0C96FED41E7AFFEA70F3619F0C28FB63" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[692,839,1251,1278]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="nalubaale" status="sp. nov.">
<emphasis id="F9E259451E7AFFEA70F3619F0C28FB63" bold="true" box="[692,839,1251,1278]" italics="true" pageId="29" pageNumber="480">R. nalubaale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E7AFFEA7109619F0CC8FB60" box="[846,935,1251,1277]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (8) genetic divergence and non-overlapping elevational range at sites of co-occurrence distinguishes it from 
<emphasis id="F9E259451E7AFFEA7124607B0B3BFABC" bold="true" box="[867,1108,1287,1314]" pageId="29" pageNumber="480">
<taxonomicName id="0C96FED41E7AFFEA7124607B0C9BFABF" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[867,1012,1287,1314]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="plumptrei" status="sp. nov.">
<emphasis id="F9E259451E7AFFEA7124607B0C9BFABF" bold="true" box="[867,1012,1287,1314]" italics="true" pageId="29" pageNumber="480">R. plumptrei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E7AFFEA71BC607B0B3BFABC" box="[1019,1108,1287,1313]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<emphasis id="F9E259451E7AFFEA76CC607B0A01FABC" bold="true" box="[1163,1390,1287,1314]" pageId="29" pageNumber="480">
<taxonomicName id="0C96FED41E7AFFEA76CC607B0A61FABF" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[1163,1294,1287,1314]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="bombayi" status="sp. nov.">
<emphasis id="F9E259451E7AFFEA76CC607B0A61FABF" bold="true" box="[1163,1294,1287,1314]" italics="true" pageId="29" pageNumber="480">R. bombayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E7AFFEA7752607B0A01FABC" box="[1301,1390,1287,1313]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="CB2985571E7AFFEA728060500EACFA10" blockId="29.[151,1437,495,1421]" pageId="29" pageNumber="480">
<emphasis id="F9E259451E7AFFEA728060500D09FAD8" box="[199,614,1323,1350]" italics="true" pageId="29" pageNumber="480">Genetic differentiation and variation.</emphasis>
A summary of pairwise sequence divergence for three DNA markers (
<emphasis id="F9E259451E7AFFEA772C60570AFAFAD8" box="[1387,1429,1323,1349]" italics="true" pageId="29" pageNumber="480">16S</emphasis>
, 
<emphasis id="F9E259451E7AFFEA72D060330FA3FAF4" box="[151,204,1359,1385]" italics="true" pageId="29" pageNumber="480">ND2</emphasis>
, and 
<emphasis id="F9E259451E7AFFEA734E60330E38FAF4" box="[265,343,1359,1385]" italics="true" pageId="29" pageNumber="480">RAG-1</emphasis>
) among individuals of 
<emphasis id="F9E259451E7AFFEA701C60330CE5FAF4" bold="true" box="[603,906,1359,1386]" pageId="29" pageNumber="480">
<taxonomicName id="0C96FED41E7AFFEA701C60330C45FAF7" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[603,810,1359,1386]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="msitugrabensis" status="sp. nov.">
<emphasis id="F9E259451E7AFFEA701C60330C45FAF7" bold="true" box="[603,810,1359,1386]" italics="true" pageId="29" pageNumber="480">R. msitugrabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E7AFFEA717660330CE5FAF4" box="[817,906,1359,1385]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and other 
<taxonomicName id="0C96FED41E7AFFEA764660330B9BFAF4" box="[1025,1268,1359,1385]" pageId="29" pageNumber="480">
<emphasis id="F9E259451E7AFFEA764660330BF2FAF4" box="[1025,1181,1359,1385]" italics="true" pageId="29" pageNumber="480">Rhampholeon</emphasis>
species
</taxonomicName>
is presented in Supplementary Material 1.
</paragraph>
</subSubSection>
<caption id="9FE9D5DF1E7AFFEA72D062C30EBEF866" ID-DOI="http://doi.org/10.5281/zenodo.11547891" ID-Zenodo-Dep="11547891" httpUri="https://zenodo.org/record/11547891/files/figure.png" pageId="29" pageNumber="480" startId="29.[151,250,1983,2007]" targetBox="[410,1178,1445,1959]" targetPageId="29" targetType="figure">
<paragraph id="CB2985571E7AFFEA72D062C30EBEF866" blockId="29.[151,1436,1982,2043]" pageId="29" pageNumber="480">
<emphasis id="F9E259451E7AFFEA72D062C30E4EF84A" bold="true" box="[151,289,1983,2007]" pageId="29" pageNumber="480">FIGURE 13.</emphasis>
Photograph of 
<emphasis id="F9E259451E7AFFEA738762C20C3DF84A" bold="true" box="[448,850,1982,2007]" pageId="29" pageNumber="480">
<taxonomicName id="0C96FED41E7AFFEA738762C20D95F84B" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[448,762,1982,2006]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="480" phylum="Chordata" rank="species" species="msitugrabensis" status="sp. nov.">
<emphasis id="F9E259451E7AFFEA738762C20D95F84B" bold="true" box="[448,762,1982,2006]" italics="true" pageId="29" pageNumber="480">Rhampholeon msitugrabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E7AFFEA714662C30C3DF84A" box="[769,850,1983,2007]" pageId="29" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life. (A)—Adult female holotype (UTEP 21709) from Kibira National Park, Burundi.
</paragraph>
</caption>
<subSubSection id="838CD6DC1E79FFE9728065EB0DECFE4C" pageId="30" pageNumber="481" type="description">
<paragraph id="CB2985571E79FFE9728065EB0DECFE4C" blockId="30.[151,1437,151,861]" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728065EB0DEDFF2C" box="[199,642,151,177]" italics="true" pageId="30" pageNumber="481">
Description of 
<typeStatus id="142D3BF51E79FFE9733565EB0EBDFF2C" box="[370,466,151,177]" pageId="30" pageNumber="481" type="holotype">holotype</typeStatus>
(UTEP 21709)
</emphasis>
. Adult female, SVL 
<quantity id="0C6E28B21E79FFE9713765EB0CBAFF2C" box="[880,981,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.9399999999999995" pageId="30" pageNumber="481" unit="mm" value="49.4">49.4 mm</quantity>
and TL 
<quantity id="0C6E28B21E79FFE9767065EB0BF0FF2C" box="[1079,1183,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="30" pageNumber="481" unit="mm" value="11.3">11.3 mm</quantity>
. Body shape leaf-like. Casque flattened, with short head. Neck indistinct from head. Supra-orbital crests distinct with cluster of tubercles connected by a ridge with 10 tubercles across casque and 15 tubercles from peak-to-peak of crests. Rostral process 
<quantity id="0C6E28B21E79FFE972D0647F0F9AFE80" box="[151,245,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="30" pageNumber="481" unit="mm" value="1.3">1.3 mm</quantity>
, composed of elongated tubercles. Temporal crest discrete with several enlarged tubercles extending posteriorly from mid-eye. Nares open in a posterior orientation. Canthal ridge consists of raised tubercles, one raised higher than others near snout. Ninety-six upper and 88 lower labial tubercles are present along tip of snout to rictus of mouth. Body covered in nearly homogenous, flattened tubercles. Some larger conical tubercles present on dorsal flanks around midbody. Crenulated dorsal crest, more prominent from midbody to nape. Enlarged conical tubercles present on limbs. Claws markedly bicuspid.
</paragraph>
</subSubSection>
<subSubSection id="838CD6DC1E79FFE9728064A70CA1F8C6" pageId="30" pageNumber="481" type="biology_ecology">
<paragraph id="CB2985571E79FFE9728064A70E22FD34" blockId="30.[151,1437,151,861]" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728064A70D40FE68" box="[199,559,475,501]" italics="true" pageId="30" pageNumber="481">
Coloration of 
<typeStatus id="142D3BF51E79FFE9732E64A70EA6FE68" box="[361,457,475,501]" pageId="30" pageNumber="481" type="holotype">holotype</typeStatus>
(in life).
</emphasis>
A photograph of the 
<typeStatus id="142D3BF51E79FFE9716164A70CE7FE68" box="[806,904,475,501]" pageId="30" pageNumber="481" type="holotype">holotype</typeStatus>
is presented in 
<figureCitation id="53AD99D21E79FFE9767864A70BABFE68" box="[1087,1220,475,501]" captionStart="FIGURE 13" captionStartId="29.[151,250,1983,2007]" captionTargetBox="[410,1178,1445,1959]" captionTargetId="figure-470@29.[410,1178,1445,1959]" captionTargetPageId="29" captionText="FIGURE 13. Photograph of Rhampholeon msitugrabensis sp. nov. in life. (A)—Adult female holotype (UTEP 21709) from Kibira National Park, Burundi." figureDoi="http://doi.org/10.5281/zenodo.11547891" httpUri="https://zenodo.org/record/11547891/files/figure.png" pageId="30" pageNumber="481">Figure 13A</figureCitation>
. Background color mottled light brown to gray with several darker brown patches, especially on limbs, tail, and dorsal crest. Top of head light brown and area near snout darker brown. Head exhibits several darker brown patches on ridges and below eye along mouth. Gular region white, which extends from tip of chin to neck region. Two diagonal dark brown lines extend from near dorsal crest posteriorly to lateral flanks, resembling veins of a leaf. Ventral region a much lighter color than body.
</paragraph>
<paragraph id="CB2985571E79FFE9728067CF0B4FFCC0" blockId="30.[151,1437,151,861]" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728067CF0E58FD51" box="[199,311,691,716]" italics="true" pageId="30" pageNumber="481">Variation.</emphasis>
A summary of descriptive morphometrics for 
<emphasis id="F9E259451E79FFE9711767CF0BE8FD50" bold="true" box="[848,1159,690,717]" pageId="30" pageNumber="481">
<taxonomicName id="0C96FED41E79FFE9711767CF0B4EFD50" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[848,1057,690,717]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="481" phylum="Chordata" rank="species" species="msitugrabensis" status="sp. nov.">
<emphasis id="F9E259451E79FFE9711767CF0B4EFD50" bold="true" box="[848,1057,690,717]" italics="true" pageId="30" pageNumber="481">R. msitugrabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E79FFE9766C67CF0BE8FD50" box="[1067,1159,691,717]" pageId="30" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is presented in 
<tableCitation id="8614B0EC1E79FFE9770767CF0AF8FD50" box="[1344,1431,691,717]" captionStart="TABLE 4" captionStartId="21.[152,244,151,175]" captionTargetPageId="21" captionText="TABLE 4. Summary of meristic and mensural characters in specimens of Rhampholeon msitugrabensis sp. nov. and R. monteslunae sp. nov. Linear measurements (in mm) and scale counts are given as mean ± standard deviation, followed by range in parentheses; see Materials and Methods for explanation of character abbreviations." httpUri="http://table.plazi.org/id/9FE9D5DF1E72FFE272DF65EB0BE4FF6A" pageId="30" pageNumber="481" tableUuid="9FE9D5DF1E72FFE272DF65EB0BE4FF6A">Table 4</tableCitation>
, comparative boxplots in 
<figureCitation id="53AD99D21E79FFE973E967AB0D60FD6C" box="[430,527,727,753]" captionStart="FIGURE 4" captionStartId="8.[151,250,1842,1866]" captionTargetBox="[209,1381,181,1814]" captionTargetId="figure-20@8.[205,1383,181,1818]" captionTargetPageId="8" captionText="FIGURE 4. Box plots showing variation in external traits of the Rhampholeon boulengeri complex from the Albertine Rift, Central Africa. Sample sizes for each species are denoted in the bottom right quarter. Males and females are separated to show degree of sexual-size dimorphism across members of the complex. All measurements are in mm. Plots show the median, 1st and 3rd quartiles, minimum and maximum inter-quartile ranges, and outliers. Refer to text for description of the linear measurements." figureDoi="http://doi.org/10.5281/zenodo.11547865" httpUri="https://zenodo.org/record/11547865/files/figure.png" pageId="30" pageNumber="481">Figure 4</figureCitation>
, and measurements of the type specimens in 
<tableCitation id="8614B0EC1E79FFE9764B67AB0B0EFD6C" box="[1036,1121,727,753]" captionStart="TABLE 8" captionStartId="30.[152,244,908,932]" captionTargetPageId="30" captionText="TABLE 8. Meristic and mensural characters in type specimens of Rhampholeon msitugrabensis sp. nov. Linear measurements (in mm) and scale counts are given. See Materials and Methods for explanation of character abbreviations." httpUri="http://table.plazi.org/id/9FE9D5DF1E79FFE972DF66F00E41FC71" pageId="30" pageNumber="481" tableUuid="9FE9D5DF1E79FFE972DF66F00E41FC71">Table 8</tableCitation>
. Morphological proportions are generally consistent with those of the 
<typeStatus id="142D3BF51E79FFE970C367870D85FC88" box="[644,746,763,789]" pageId="30" pageNumber="481" type="holotype">holotype</typeStatus>
. Males have smaller body sizes (M: mean 
<quantity id="0C6E28B21E79FFE976A267870A20FC88" box="[1253,1359,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="4.17" pageId="30" pageNumber="481" unit="mm" value="41.7">41.7 mm</quantity>
, range 
<quantity id="0C6E28B21E79FFE972D066630E52FCA4" box="[151,317,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" metricValueMax="4.68" metricValueMin="3.36" pageId="30" pageNumber="481" unit="mm" value="40.2" valueMax="46.8" valueMin="33.6">33.6–46.8 mm</quantity>
, n = 3; F: mean 
<quantity id="0C6E28B21E79FFE973B266630D33FCA4" box="[501,604,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="4.779999999999999" pageId="30" pageNumber="481" unit="mm" value="47.8">47.8 mm</quantity>
, range 
<quantity id="0C6E28B21E79FFE970E966620C3CFCA4" box="[686,851,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="4.755" metricValueMax="4.9399999999999995" metricValueMin="4.57" pageId="30" pageNumber="481" unit="mm" value="47.55" valueMax="49.4" valueMin="45.7">45.7–49.4 mm</quantity>
, n = 4) and longer tails than females (M: mean 
<quantity id="0C6E28B21E79FFE9772C66630FAAFCC0" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="30" pageNumber="481" unit="mm" value="12.4">12.4 mm</quantity>
, range 
<quantity id="0C6E28B21E79FFE97351663E0ED5FCC0" box="[278,442,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" metricValueMax="1.3699999999999999" metricValueMin="1.15" pageId="30" pageNumber="481" unit="mm" value="12.6" valueMax="13.7" valueMin="11.5">11.5–13.7 mm</quantity>
, n = 3; F: mean 
<quantity id="0C6E28B21E79FFE97037663F0DB9FCC0" box="[624,726,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="30" pageNumber="481" unit="mm" value="10.9">10.9 mm</quantity>
, range 
<quantity id="0C6E28B21E79FFE97160663E0CA4FCC0" box="[807,971,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" metricValueMax="1.13" metricValueMin="1.05" pageId="30" pageNumber="481" unit="mm" value="10.9" valueMax="11.3" valueMin="10.5">10.5–11.3 mm</quantity>
, n = 4).
</paragraph>
<caption id="9FE9D5DF1E79FFE972DF66F00E41FC71" ID-Table-UUID="9FE9D5DF1E79FFE972DF66F00E41FC71" httpUri="http://table.plazi.org/id/9FE9D5DF1E79FFE972DF66F00E41FC71" pageId="30" pageNumber="481" startId="30.[152,244,908,932]" targetBox="[169,1407,1020,1740]" targetIsTable="true" targetPageId="30" targetType="table">
<paragraph id="CB2985571E79FFE972DF66F00E41FC71" blockId="30.[151,1437,908,1004]" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE972DF66F00E7AFC39" bold="true" box="[152,277,908,932]" pageId="30" pageNumber="481">TABLE 8.</emphasis>
Meristic and mensural characters in type specimens of 
<emphasis id="F9E259451E79FFE971D166F00A24FC38" bold="true" box="[918,1355,908,933]" pageId="30" pageNumber="481">
<taxonomicName id="0C96FED41E79FFE971D166F00B8BFC39" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[918,1252,908,932]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="481" phylum="Chordata" rank="species" species="msitugrabensis" status="sp. nov.">
<emphasis id="F9E259451E79FFE971D166F00B8BFC39" bold="true" box="[918,1252,908,932]" italics="true" pageId="30" pageNumber="481">Rhampholeon msitugrabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E79FFE976B766F10A24FC38" box="[1264,1355,909,933]" pageId="30" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Linear measurements (in mm) and scale counts are given. See Materials and Methods for explanation of character abbreviations.
</paragraph>
</caption>
<paragraph id="CB2985571E79FFE9736066800A3BF956" pageId="30" pageNumber="481">
<table id="B99677F71E79000872EE66800A10F951" box="[169,1407,1020,1740]" gridcols="7" gridrows="21" pageId="30" pageNumber="481">
<tr id="75A687151E79000872EE66800A10FB8C" box="[169,1407,1020,1041]" gridrow="0" pageId="30" pageNumber="481" rowspan-0="1">
<th id="3677EE691E790008736066800EC7FB8C" box="[295,424,1020,1041]" gridcol="1" gridrow="0" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9736066800EC7FB8C" bold="true" box="[295,424,1020,1041]" pageId="30" pageNumber="481">UTEP 21709</emphasis>
</th>
<th id="3677EE691E790008739966800D06FB8C" box="[478,617,1020,1041]" gridcol="2" gridrow="0" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE973A466800D0BFB8C" bold="true" box="[483,612,1020,1041]" pageId="30" pageNumber="481">UTEP 22739</emphasis>
</th>
<th id="3677EE691E79000870E366800C40FB8C" box="[676,815,1020,1041]" gridcol="3" gridrow="0" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE970EE66800C46FB8C" bold="true" box="[681,809,1020,1041]" pageId="30" pageNumber="481">UTEP 22740</emphasis>
</th>
<th id="3677EE691E790008712E66800C9BFB8C" box="[873,1012,1020,1041]" gridcol="4" gridrow="0" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9712966800C80FB8C" bold="true" box="[878,1007,1020,1041]" pageId="30" pageNumber="481">UTEP 22741</emphasis>
</th>
<th id="3677EE691E790008766866800BD5FB8C" box="[1071,1210,1020,1041]" gridcol="5" gridrow="0" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9767366800BDBFB8C" bold="true" box="[1076,1204,1020,1041]" pageId="30" pageNumber="481">UTEP 22742</emphasis>
</th>
<th id="3677EE691E79000876B366800A10FB8C" box="[1268,1407,1020,1041]" gridcol="6" gridrow="0" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE976BE66800A15FB8C" bold="true" box="[1273,1402,1020,1041]" pageId="30" pageNumber="481">UTEP 22743</emphasis>
</th>
</tr>
<tr id="75A687151E79000872EE615C0A10FBA8" box="[169,1407,1056,1077]" gridrow="1" pageId="30" pageNumber="481" rowspan-0="1">
<td id="3677EE691E7900087360615C0EC7FBA8" box="[295,424,1056,1077]" gridcol="1" gridrow="1" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE97305615C0EE2FBA8" bold="true" box="[322,397,1056,1077]" pageId="30" pageNumber="481">Female</emphasis>
</td>
<td id="3677EE691E7900087399615C0D06FBA8" box="[478,617,1056,1077]" gridcol="2" gridrow="1" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9704D615C0D51FBA8" bold="true" box="[522,574,1056,1077]" pageId="30" pageNumber="481">Male</emphasis>
</td>
<td id="3677EE691E79000870E3615C0C40FBA8" box="[676,815,1056,1077]" gridcol="3" gridrow="1" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE97083615C0C60FBA8" bold="true" box="[708,783,1056,1077]" pageId="30" pageNumber="481">Female</emphasis>
</td>
<td id="3677EE691E790008712E615C0C9BFBA8" box="[873,1012,1056,1077]" gridcol="4" gridrow="1" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE971D2615C0CA6FBA8" bold="true" box="[917,969,1056,1077]" pageId="30" pageNumber="481">Male</emphasis>
</td>
<td id="3677EE691E7900087668615C0BD5FBA8" box="[1071,1210,1056,1077]" gridcol="5" gridrow="1" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE97608615C0BF5FBA8" bold="true" box="[1103,1178,1056,1077]" pageId="30" pageNumber="481">Female</emphasis>
</td>
<td id="3677EE691E79000876B3615C0A10FBA8" box="[1268,1407,1056,1077]" gridcol="6" gridrow="1" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE97753615C0A30FBA8" bold="true" box="[1300,1375,1056,1077]" pageId="30" pageNumber="481">Female</emphasis>
</td>
</tr>
<tr id="75A687151E79000872EE61380A10FBC4" box="[169,1407,1092,1113]" gridrow="2" pageId="30" pageNumber="481" rowspan-0="1">
<td id="3677EE691E790008736061380EC7FBC4" box="[295,424,1092,1113]" gridcol="1" gridrow="2" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9737E61380EF9FBC4" bold="true" box="[313,406,1092,1113]" pageId="30" pageNumber="481">Holotype</emphasis>
</td>
<td id="3677EE691E790008739961380D06FBC4" box="[478,617,1092,1113]" gridcol="2" gridrow="2" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9739961380D06FBC4" bold="true" box="[478,617,1092,1113]" pageId="30" pageNumber="481">Paratopotype</emphasis>
</td>
<td id="3677EE691E79000870E361380C40FBC4" box="[676,815,1092,1113]" gridcol="3" gridrow="2" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE970E361380C40FBC4" bold="true" box="[676,815,1092,1113]" pageId="30" pageNumber="481">Paratopotype</emphasis>
</td>
<td id="3677EE691E790008712E61380C9BFBC4" box="[873,1012,1092,1113]" gridcol="4" gridrow="2" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9712E61380C9BFBC4" bold="true" box="[873,1012,1092,1113]" pageId="30" pageNumber="481">Paratopotype</emphasis>
</td>
<td id="3677EE691E790008766861380BD5FBC4" box="[1071,1210,1092,1113]" gridcol="5" gridrow="2" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9766861380BD5FBC4" bold="true" box="[1071,1210,1092,1113]" pageId="30" pageNumber="481">Paratopotype</emphasis>
</td>
<td id="3677EE691E79000876B361380A10FBC4" box="[1268,1407,1092,1113]" gridcol="6" gridrow="2" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE976B361380A10FBC4" bold="true" box="[1268,1407,1092,1113]" pageId="30" pageNumber="481">Paratopotype</emphasis>
</td>
</tr>
<tr id="75A687151E79000872EE611A0A10FBE0" box="[169,1407,1126,1149]" gridrow="3" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE611A0F91FBE0" box="[169,254,1126,1149]" gridcol="0" gridrow="3" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE972FB611A0F84FBE1" bold="true" box="[188,235,1126,1148]" pageId="30" pageNumber="481">SVL</emphasis>
</th>
<td id="3677EE691E7900087360611A0EC7FBE0" box="[295,424,1126,1149]" gridcol="1" gridrow="3" pageId="30" pageNumber="481">49.36</td>
<td id="3677EE691E7900087399611A0D06FBE0" box="[478,617,1126,1149]" gridcol="2" gridrow="3" pageId="30" pageNumber="481">44.53</td>
<td id="3677EE691E79000870E3611A0C40FBE0" box="[676,815,1126,1149]" gridcol="3" gridrow="3" pageId="30" pageNumber="481">49.32</td>
<td id="3677EE691E790008712E611A0C9BFBE0" box="[873,1012,1126,1149]" gridcol="4" gridrow="3" pageId="30" pageNumber="481">33.58</td>
<td id="3677EE691E7900087668611A0BD5FBE0" box="[1071,1210,1126,1149]" gridcol="5" gridrow="3" pageId="30" pageNumber="481">45.67</td>
<td id="3677EE691E79000876B3611A0A10FBE0" box="[1268,1407,1126,1149]" gridcol="6" gridrow="3" pageId="30" pageNumber="481">46.82</td>
</tr>
<tr id="75A687151E79000872EE61F50A10FB02" box="[169,1407,1161,1183]" gridrow="4" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE61F50F91FB02" box="[169,254,1161,1183]" gridcol="0" gridrow="4" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728461F50F8CFB03" bold="true" box="[195,227,1161,1182]" pageId="30" pageNumber="481">TL</emphasis>
</th>
<td id="3677EE691E790008736061F50EC7FB02" box="[295,424,1161,1183]" gridcol="1" gridrow="4" pageId="30" pageNumber="481">11.27</td>
<td id="3677EE691E790008739961F50D06FB02" box="[478,617,1161,1183]" gridcol="2" gridrow="4" pageId="30" pageNumber="481">11.92</td>
<td id="3677EE691E79000870E361F50C40FB02" box="[676,815,1161,1183]" gridcol="3" gridrow="4" pageId="30" pageNumber="481">11.26</td>
<td id="3677EE691E790008712E61F50C9BFB02" box="[873,1012,1161,1183]" gridcol="4" gridrow="4" pageId="30" pageNumber="481">11.53</td>
<td id="3677EE691E790008766861F50BD5FB02" box="[1071,1210,1161,1183]" gridcol="5" gridrow="4" pageId="30" pageNumber="481">10.46</td>
<td id="3677EE691E79000876B361F50A10FB02" box="[1268,1407,1161,1183]" gridcol="6" gridrow="4" pageId="30" pageNumber="481">10.84</td>
</tr>
<tr id="75A687151E79000872EE61D00A10FB5F" box="[169,1407,1196,1218]" gridrow="5" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE61D00F91FB5F" box="[169,254,1196,1218]" gridcol="0" gridrow="5" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE972F961D00F87FB5C" bold="true" box="[190,232,1196,1217]" pageId="30" pageNumber="481">TaL</emphasis>
</th>
<td id="3677EE691E790008736061D00EC7FB5F" box="[295,424,1196,1218]" gridcol="1" gridrow="5" pageId="30" pageNumber="481">60.63</td>
<td id="3677EE691E790008739961D00D06FB5F" box="[478,617,1196,1218]" gridcol="2" gridrow="5" pageId="30" pageNumber="481">56.45</td>
<td id="3677EE691E79000870E361D00C40FB5F" box="[676,815,1196,1218]" gridcol="3" gridrow="5" pageId="30" pageNumber="481">60.58</td>
<td id="3677EE691E790008712E61D00C9BFB5F" box="[873,1012,1196,1218]" gridcol="4" gridrow="5" pageId="30" pageNumber="481">45.11</td>
<td id="3677EE691E790008766861D00BD5FB5F" box="[1071,1210,1196,1218]" gridcol="5" gridrow="5" pageId="30" pageNumber="481">56.13</td>
<td id="3677EE691E79000876B361D00A10FB5F" box="[1268,1407,1196,1218]" gridcol="6" gridrow="5" pageId="30" pageNumber="481">57.66</td>
</tr>
<tr id="75A687151E79000872EE61B30A10FB78" box="[169,1407,1231,1253]" gridrow="6" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE61B30F91FB78" box="[169,254,1231,1253]" gridcol="0" gridrow="6" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728561B30F8AFB79" bold="true" box="[194,229,1231,1252]" pageId="30" pageNumber="481">HL</emphasis>
</th>
<td id="3677EE691E790008736061B30EC7FB78" box="[295,424,1231,1253]" gridcol="1" gridrow="6" pageId="30" pageNumber="481">14.54</td>
<td id="3677EE691E790008739961B30D06FB78" box="[478,617,1231,1253]" gridcol="2" gridrow="6" pageId="30" pageNumber="481">14.42</td>
<td id="3677EE691E79000870E361B30C40FB78" box="[676,815,1231,1253]" gridcol="3" gridrow="6" pageId="30" pageNumber="481">14.38</td>
<td id="3677EE691E790008712E61B30C9BFB78" box="[873,1012,1231,1253]" gridcol="4" gridrow="6" pageId="30" pageNumber="481">12.07</td>
<td id="3677EE691E790008766861B30BD5FB78" box="[1071,1210,1231,1253]" gridcol="5" gridrow="6" pageId="30" pageNumber="481">14.07</td>
<td id="3677EE691E79000876B361B30A10FB78" box="[1268,1407,1231,1253]" gridcol="6" gridrow="6" pageId="30" pageNumber="481">14.01</td>
</tr>
<tr id="75A687151E79000872EE618E0A10FA95" box="[169,1407,1266,1288]" gridrow="7" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE618E0F91FA95" box="[169,254,1266,1288]" gridcol="0" gridrow="7" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE972FA618E0F87FA9A" bold="true" box="[189,232,1266,1287]" pageId="30" pageNumber="481">HW</emphasis>
</th>
<td id="3677EE691E7900087360618E0EC7FA95" box="[295,424,1266,1288]" gridcol="1" gridrow="7" pageId="30" pageNumber="481">8.66</td>
<td id="3677EE691E7900087399618E0D06FA95" box="[478,617,1266,1288]" gridcol="2" gridrow="7" pageId="30" pageNumber="481">8.55</td>
<td id="3677EE691E79000870E3618E0C40FA95" box="[676,815,1266,1288]" gridcol="3" gridrow="7" pageId="30" pageNumber="481">8.24</td>
<td id="3677EE691E790008712E618E0C9BFA95" box="[873,1012,1266,1288]" gridcol="4" gridrow="7" pageId="30" pageNumber="481">7.11</td>
<td id="3677EE691E7900087668618E0BD5FA95" box="[1071,1210,1266,1288]" gridcol="5" gridrow="7" pageId="30" pageNumber="481">7.89</td>
<td id="3677EE691E79000876B3618E0A10FA95" box="[1268,1407,1266,1288]" gridcol="6" gridrow="7" pageId="30" pageNumber="481">8.44</td>
</tr>
<tr id="75A687151E79000872EE60680A10FAB6" box="[169,1407,1300,1323]" gridrow="8" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE60680F91FAB6" box="[169,254,1300,1323]" gridcol="0" gridrow="8" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728760680F8AFAB4" bold="true" box="[192,229,1300,1321]" pageId="30" pageNumber="481">HH</emphasis>
</th>
<td id="3677EE691E790008736060680EC7FAB6" box="[295,424,1300,1323]" gridcol="1" gridrow="8" pageId="30" pageNumber="481">7.77</td>
<td id="3677EE691E790008739960680D06FAB6" box="[478,617,1300,1323]" gridcol="2" gridrow="8" pageId="30" pageNumber="481">8.28</td>
<td id="3677EE691E79000870E360680C40FAB6" box="[676,815,1300,1323]" gridcol="3" gridrow="8" pageId="30" pageNumber="481">7.08</td>
<td id="3677EE691E790008712E60680C9BFAB6" box="[873,1012,1300,1323]" gridcol="4" gridrow="8" pageId="30" pageNumber="481">6.76</td>
<td id="3677EE691E790008766860680BD5FAB6" box="[1071,1210,1300,1323]" gridcol="5" gridrow="8" pageId="30" pageNumber="481">7.99</td>
<td id="3677EE691E79000876B360680A10FAB6" box="[1268,1407,1300,1323]" gridcol="6" gridrow="8" pageId="30" pageNumber="481">6.77</td>
</tr>
<tr id="75A687151E79000872EE604B0A10FAD0" box="[169,1407,1335,1357]" gridrow="9" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE604B0F91FAD0" box="[169,254,1335,1357]" gridcol="0" gridrow="9" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE97287604B0F88FAD1" bold="true" box="[192,231,1335,1356]" pageId="30" pageNumber="481">ML</emphasis>
</th>
<td id="3677EE691E7900087360604B0EC7FAD0" box="[295,424,1335,1357]" gridcol="1" gridrow="9" pageId="30" pageNumber="481">9.65</td>
<td id="3677EE691E7900087399604B0D06FAD0" box="[478,617,1335,1357]" gridcol="2" gridrow="9" pageId="30" pageNumber="481">9.36</td>
<td id="3677EE691E79000870E3604B0C40FAD0" box="[676,815,1335,1357]" gridcol="3" gridrow="9" pageId="30" pageNumber="481">9.71</td>
<td id="3677EE691E790008712E604B0C9BFAD0" box="[873,1012,1335,1357]" gridcol="4" gridrow="9" pageId="30" pageNumber="481">8.24</td>
<td id="3677EE691E7900087668604B0BD5FAD0" box="[1071,1210,1335,1357]" gridcol="5" gridrow="9" pageId="30" pageNumber="481">9.74</td>
<td id="3677EE691E79000876B3604B0A10FAD0" box="[1268,1407,1335,1357]" gridcol="6" gridrow="9" pageId="30" pageNumber="481">9.28</td>
</tr>
<tr id="75A687151E79000872EE60260A10FAED" box="[169,1407,1370,1392]" gridrow="10" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE60260F91FAED" box="[169,254,1370,1392]" gridcol="0" gridrow="10" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728560260F8CFAED" bold="true" box="[194,227,1370,1392]" pageId="30" pageNumber="481">CE</emphasis>
</th>
<td id="3677EE691E790008736060260EC7FAED" box="[295,424,1370,1392]" gridcol="1" gridrow="10" pageId="30" pageNumber="481">7.18</td>
<td id="3677EE691E790008739960260D06FAED" box="[478,617,1370,1392]" gridcol="2" gridrow="10" pageId="30" pageNumber="481">7.27</td>
<td id="3677EE691E79000870E360260C40FAED" box="[676,815,1370,1392]" gridcol="3" gridrow="10" pageId="30" pageNumber="481">6.86</td>
<td id="3677EE691E790008712E60260C9BFAED" box="[873,1012,1370,1392]" gridcol="4" gridrow="10" pageId="30" pageNumber="481">6.09</td>
<td id="3677EE691E790008766860260BD5FAED" box="[1071,1210,1370,1392]" gridcol="5" gridrow="10" pageId="30" pageNumber="481">6.97</td>
<td id="3677EE691E79000876B360260A10FAED" box="[1268,1407,1370,1392]" gridcol="6" gridrow="10" pageId="30" pageNumber="481">6.69</td>
</tr>
<tr id="75A687151E79000872EE60000A10FA0E" box="[169,1407,1404,1427]" gridrow="11" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE60000F91FA0E" box="[169,254,1404,1427]" gridcol="0" gridrow="11" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728260000F8DFA0F" bold="true" box="[197,226,1404,1426]" pageId="30" pageNumber="481">SL</emphasis>
</th>
<td id="3677EE691E790008736060000EC7FA0E" box="[295,424,1404,1427]" gridcol="1" gridrow="11" pageId="30" pageNumber="481">5.11</td>
<td id="3677EE691E790008739960000D06FA0E" box="[478,617,1404,1427]" gridcol="2" gridrow="11" pageId="30" pageNumber="481">4.71</td>
<td id="3677EE691E79000870E360000C40FA0E" box="[676,815,1404,1427]" gridcol="3" gridrow="11" pageId="30" pageNumber="481">4.51</td>
<td id="3677EE691E790008712E60000C9BFA0E" box="[873,1012,1404,1427]" gridcol="4" gridrow="11" pageId="30" pageNumber="481">4.05</td>
<td id="3677EE691E790008766860000BD5FA0E" box="[1071,1210,1404,1427]" gridcol="5" gridrow="11" pageId="30" pageNumber="481">4.98</td>
<td id="3677EE691E79000876B360000A10FA0E" box="[1268,1407,1404,1427]" gridcol="6" gridrow="11" pageId="30" pageNumber="481">4.71</td>
</tr>
<tr id="75A687151E79000872EE60DC0A10FA2B" box="[169,1407,1440,1462]" gridrow="12" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE60DC0F91FA2B" box="[169,254,1440,1462]" gridcol="0" gridrow="12" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728560DC0F8CFA28" bold="true" box="[194,227,1440,1461]" pageId="30" pageNumber="481">ED</emphasis>
</th>
<td id="3677EE691E790008736060DC0EC7FA2B" box="[295,424,1440,1462]" gridcol="1" gridrow="12" pageId="30" pageNumber="481">4.13</td>
<td id="3677EE691E790008739960DC0D06FA2B" box="[478,617,1440,1462]" gridcol="2" gridrow="12" pageId="30" pageNumber="481">4.31</td>
<td id="3677EE691E79000870E360DC0C40FA2B" box="[676,815,1440,1462]" gridcol="3" gridrow="12" pageId="30" pageNumber="481">4.24</td>
<td id="3677EE691E790008712E60DC0C9BFA2B" box="[873,1012,1440,1462]" gridcol="4" gridrow="12" pageId="30" pageNumber="481">3.75</td>
<td id="3677EE691E790008766860DC0BD5FA2B" box="[1071,1210,1440,1462]" gridcol="5" gridrow="12" pageId="30" pageNumber="481">3.48</td>
<td id="3677EE691E79000876B360DC0A10FA2B" box="[1268,1407,1440,1462]" gridcol="6" gridrow="12" pageId="30" pageNumber="481">4.05</td>
</tr>
<tr id="75A687151E79000872EE60BE0A10FA45" box="[169,1407,1474,1496]" gridrow="13" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE60BE0F91FA45" box="[169,254,1474,1496]" gridcol="0" gridrow="13" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728660BE0F8BFA45" bold="true" box="[193,228,1474,1496]" pageId="30" pageNumber="481">CC</emphasis>
</th>
<td id="3677EE691E790008736060BE0EC7FA45" box="[295,424,1474,1496]" gridcol="1" gridrow="13" pageId="30" pageNumber="481">5.63</td>
<td id="3677EE691E790008739960BE0D06FA45" box="[478,617,1474,1496]" gridcol="2" gridrow="13" pageId="30" pageNumber="481">6.15</td>
<td id="3677EE691E79000870E360BE0C40FA45" box="[676,815,1474,1496]" gridcol="3" gridrow="13" pageId="30" pageNumber="481">5.51</td>
<td id="3677EE691E790008712E60BE0C9BFA45" box="[873,1012,1474,1496]" gridcol="4" gridrow="13" pageId="30" pageNumber="481">5.38</td>
<td id="3677EE691E790008766860BE0BD5FA45" box="[1071,1210,1474,1496]" gridcol="5" gridrow="13" pageId="30" pageNumber="481">5.78</td>
<td id="3677EE691E79000876B360BE0A10FA45" box="[1268,1407,1474,1496]" gridcol="6" gridrow="13" pageId="30" pageNumber="481">5.15</td>
</tr>
<tr id="75A687151E79000872EE60990A10FA66" box="[169,1407,1509,1531]" gridrow="14" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE60990F91FA66" box="[169,254,1509,1531]" gridcol="0" gridrow="14" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728060990F8FFA67" bold="true" box="[199,224,1509,1530]" pageId="30" pageNumber="481">IL</emphasis>
</th>
<td id="3677EE691E790008736060990EC7FA66" box="[295,424,1509,1531]" gridcol="1" gridrow="14" pageId="30" pageNumber="481">25.79</td>
<td id="3677EE691E790008739960990D06FA66" box="[478,617,1509,1531]" gridcol="2" gridrow="14" pageId="30" pageNumber="481">24.79</td>
<td id="3677EE691E79000870E360990C40FA66" box="[676,815,1509,1531]" gridcol="3" gridrow="14" pageId="30" pageNumber="481">25.74</td>
<td id="3677EE691E790008712E60990C9BFA66" box="[873,1012,1509,1531]" gridcol="4" gridrow="14" pageId="30" pageNumber="481">16.79</td>
<td id="3677EE691E790008766860990BD5FA66" box="[1071,1210,1509,1531]" gridcol="5" gridrow="14" pageId="30" pageNumber="481">25.5</td>
<td id="3677EE691E79000876B360990A10FA66" box="[1268,1407,1509,1531]" gridcol="6" gridrow="14" pageId="30" pageNumber="481">26.15</td>
</tr>
<tr id="75A687151E79000872EE63740A10F980" box="[169,1407,1544,1565]" gridrow="15" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE63740F91F980" box="[169,254,1544,1565]" gridcol="0" gridrow="15" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE972FB63740F84F980" bold="true" box="[188,235,1544,1565]" pageId="30" pageNumber="481">FLL</emphasis>
</th>
<td id="3677EE691E790008736063740EC7F980" box="[295,424,1544,1565]" gridcol="1" gridrow="15" pageId="30" pageNumber="481">10.13</td>
<td id="3677EE691E790008739963740D06F980" box="[478,617,1544,1565]" gridcol="2" gridrow="15" pageId="30" pageNumber="481">9.97</td>
<td id="3677EE691E79000870E363740C40F980" box="[676,815,1544,1565]" gridcol="3" gridrow="15" pageId="30" pageNumber="481">10.14</td>
<td id="3677EE691E790008712E63740C9BF980" box="[873,1012,1544,1565]" gridcol="4" gridrow="15" pageId="30" pageNumber="481">8.03</td>
<td id="3677EE691E790008766863740BD5F980" box="[1071,1210,1544,1565]" gridcol="5" gridrow="15" pageId="30" pageNumber="481">9.8</td>
<td id="3677EE691E79000876B363740A10F980" box="[1268,1407,1544,1565]" gridcol="6" gridrow="15" pageId="30" pageNumber="481">9.79</td>
</tr>
<tr id="75A687151E79000872EE63570A10F9DD" box="[169,1407,1579,1600]" gridrow="16" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE63570F91F9DD" box="[169,254,1579,1600]" gridcol="0" gridrow="16" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE972FD63570F82F9DD" bold="true" box="[186,237,1579,1600]" pageId="30" pageNumber="481">HLL</emphasis>
</th>
<td id="3677EE691E790008736063570EC7F9DD" box="[295,424,1579,1600]" gridcol="1" gridrow="16" pageId="30" pageNumber="481">9.2</td>
<td id="3677EE691E790008739963570D06F9DD" box="[478,617,1579,1600]" gridcol="2" gridrow="16" pageId="30" pageNumber="481">9.37</td>
<td id="3677EE691E79000870E363570C40F9DD" box="[676,815,1579,1600]" gridcol="3" gridrow="16" pageId="30" pageNumber="481">9.82</td>
<td id="3677EE691E790008712E63570C9BF9DD" box="[873,1012,1579,1600]" gridcol="4" gridrow="16" pageId="30" pageNumber="481">7.02</td>
<td id="3677EE691E790008766863570BD5F9DD" box="[1071,1210,1579,1600]" gridcol="5" gridrow="16" pageId="30" pageNumber="481">8.79</td>
<td id="3677EE691E79000876B363570A10F9DD" box="[1268,1407,1579,1600]" gridcol="6" gridrow="16" pageId="30" pageNumber="481">8.91</td>
</tr>
<tr id="75A687151E79000872EE63310A10F9F9" box="[169,1407,1613,1636]" gridrow="17" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE63310F91F9F9" box="[169,254,1613,1636]" gridcol="0" gridrow="17" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728463310F8BF9FF" bold="true" box="[195,228,1613,1634]" pageId="30" pageNumber="481">UL</emphasis>
</th>
<td id="3677EE691E790008736063310EC7F9F9" box="[295,424,1613,1636]" gridcol="1" gridrow="17" pageId="30" pageNumber="481">96</td>
<td id="3677EE691E790008739963310D06F9F9" box="[478,617,1613,1636]" gridcol="2" gridrow="17" pageId="30" pageNumber="481">84</td>
<td id="3677EE691E79000870E363310C40F9F9" box="[676,815,1613,1636]" gridcol="3" gridrow="17" pageId="30" pageNumber="481">88</td>
<td id="3677EE691E790008712E63310C9BF9F9" box="[873,1012,1613,1636]" gridcol="4" gridrow="17" pageId="30" pageNumber="481">96</td>
<td id="3677EE691E790008766863310BD5F9F9" box="[1071,1210,1613,1636]" gridcol="5" gridrow="17" pageId="30" pageNumber="481">94</td>
<td id="3677EE691E79000876B363310A10F9F9" box="[1268,1407,1613,1636]" gridcol="6" gridrow="17" pageId="30" pageNumber="481">94</td>
</tr>
<tr id="75A687151E79000872EE630C0A10F91B" box="[169,1407,1648,1670]" gridrow="18" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE630C0F91F91B" box="[169,254,1648,1670]" gridcol="0" gridrow="18" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE97284630C0F8CF918" bold="true" box="[195,227,1648,1669]" pageId="30" pageNumber="481">LL</emphasis>
</th>
<td id="3677EE691E7900087360630C0EC7F91B" box="[295,424,1648,1670]" gridcol="1" gridrow="18" pageId="30" pageNumber="481">88</td>
<td id="3677EE691E7900087399630C0D06F91B" box="[478,617,1648,1670]" gridcol="2" gridrow="18" pageId="30" pageNumber="481">82</td>
<td id="3677EE691E79000870E3630C0C40F91B" box="[676,815,1648,1670]" gridcol="3" gridrow="18" pageId="30" pageNumber="481">86</td>
<td id="3677EE691E790008712E630C0C9BF91B" box="[873,1012,1648,1670]" gridcol="4" gridrow="18" pageId="30" pageNumber="481">88</td>
<td id="3677EE691E7900087668630C0BD5F91B" box="[1071,1210,1648,1670]" gridcol="5" gridrow="18" pageId="30" pageNumber="481">88</td>
<td id="3677EE691E79000876B3630C0A10F91B" box="[1268,1407,1648,1670]" gridcol="6" gridrow="18" pageId="30" pageNumber="481">96</td>
</tr>
<tr id="75A687151E79000872EE63EF0A10F934" box="[169,1407,1683,1705]" gridrow="19" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE63EF0F91F934" box="[169,254,1683,1705]" gridcol="0" gridrow="19" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728463EF0F8CF935" bold="true" box="[195,227,1683,1704]" pageId="30" pageNumber="481">RP</emphasis>
</th>
<td id="3677EE691E790008736063EF0EC7F934" box="[295,424,1683,1705]" gridcol="1" gridrow="19" pageId="30" pageNumber="481">1.3</td>
<td id="3677EE691E790008739963EF0D06F934" box="[478,617,1683,1705]" gridcol="2" gridrow="19" pageId="30" pageNumber="481">1.60</td>
<td id="3677EE691E79000870E363EF0C40F934" box="[676,815,1683,1705]" gridcol="3" gridrow="19" pageId="30" pageNumber="481">0.91</td>
<td id="3677EE691E790008712E63EF0C9BF934" box="[873,1012,1683,1705]" gridcol="4" gridrow="19" pageId="30" pageNumber="481">1.01</td>
<td id="3677EE691E790008766863EF0BD5F934" box="[1071,1210,1683,1705]" gridcol="5" gridrow="19" pageId="30" pageNumber="481">1.03</td>
<td id="3677EE691E79000876B363EF0A10F934" box="[1268,1407,1683,1705]" gridcol="6" gridrow="19" pageId="30" pageNumber="481">0.77</td>
</tr>
<tr id="75A687151E79000872EE63C90A10F951" box="[169,1407,1717,1740]" gridrow="20" pageId="30" pageNumber="481">
<th id="3677EE691E79000872EE63C90F91F951" box="[169,254,1717,1740]" gridcol="0" gridrow="20" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE972EE63CA0F91F956" bold="true" box="[169,254,1717,1739]" pageId="30" pageNumber="481">TL/SVL</emphasis>
</th>
<td id="3677EE691E790008736063C90EC7F951" box="[295,424,1717,1740]" gridcol="1" gridrow="20" pageId="30" pageNumber="481">0.228</td>
<td id="3677EE691E790008739963C90D06F951" box="[478,617,1717,1740]" gridcol="2" gridrow="20" pageId="30" pageNumber="481">0.268</td>
<td id="3677EE691E79000870E363C90C40F951" box="[676,815,1717,1740]" gridcol="3" gridrow="20" pageId="30" pageNumber="481">0.228</td>
<td id="3677EE691E790008712E63C90C9BF951" box="[873,1012,1717,1740]" gridcol="4" gridrow="20" pageId="30" pageNumber="481">0.343</td>
<td id="3677EE691E790008766863C90BD5F951" box="[1071,1210,1717,1740]" gridcol="5" gridrow="20" pageId="30" pageNumber="481">0.229</td>
<td id="3677EE691E79000876B363C90A10F951" box="[1268,1407,1717,1740]" gridcol="6" gridrow="20" pageId="30" pageNumber="481">0.232</td>
</tr>
</table>
</paragraph>
<paragraph id="CB2985571E79FFE9728062610CA1F8C6" blockId="30.[151,1436,1821,2028]" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728062610E09F8AA" box="[199,358,1821,1847]" italics="true" pageId="30" pageNumber="481">Reproduction.</emphasis>
Unknown. The smallest specimen examined (
<specimenCode id="9B302D2C1E79FFE9712162620C93F8A5" box="[870,1020,1822,1848]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="30" pageNumber="481" type="Museum">UTEP 22741</specimenCode>
) was collected on 
<date id="BF28A3971E79FFE9768E62620AF3F8A5" box="[1225,1436,1821,1848]" pageId="30" pageNumber="481" value="2011-12-20">
<collectingDate id="AF6C5A7F1E79FFE9768E62620AF3F8A5" box="[1225,1436,1821,1848]" pageId="30" pageNumber="481" value="2011-12-20">20 December 2011</collectingDate>
</date>
with SVL 
<quantity id="0C6E28B21E79FFE9734C623D0E00F8C1" box="[267,367,1857,1884]" metricMagnitude="-2" metricUnit="m" metricValue="3.36" pageId="30" pageNumber="481" unit="mm" value="33.6">33.6 mm</quantity>
and TL 
<quantity id="0C6E28B21E79FFE9738B623D0D40F8C1" box="[460,559,1857,1884]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="30" pageNumber="481" unit="mm" value="11.5">11.5 mm</quantity>
from 
<location id="CE49D38C1E79FFE97034623D0C30F8C6" LSID="urn:lsid:plazi:treatment:433F34411E7AFFE872D065EB0E43FE68:CE49D38C1E79FFE97034623D0C30F8C6" box="[627,863,1857,1883]" country="Burundi" name="Kibira National Park" pageId="30" pageNumber="481">Kibira National Park</location>
, 
<collectingCountry id="B381C5C71E79FFE9712C623D0CA5F8C6" box="[875,970,1857,1883]" name="Burundi" pageId="30" pageNumber="481">Burundi</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="838CD6DC1E79FFE8728062190E43FE68" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481" type="distribution">
<paragraph id="CB2985571E79FFE8728062190E43FE68" blockId="30.[151,1436,1821,2028]" lastBlockId="31.[151,1437,150,501]" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481">
<emphasis id="F9E259451E79FFE9728062190B27F81D" box="[199,1096,1893,1920]" italics="true" pageId="30" pageNumber="481">
Distribution, natural history, and conservation. 
<taxonomicName id="0C96FED41E79FFE970AA62190B27F81D" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[749,1096,1893,1920]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="481" phylum="Chordata" rank="species" species="msitugrabensis" status="sp. nov.">
<emphasis id="F9E259451E79FFE970AA62190B27F81D" bold="true" box="[749,1096,1893,1920]" pageId="30" pageNumber="481">Rhampholeon msitugrabensis</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="E2D1E43E1E79FFE97616621A0BC2F81D" box="[1105,1197,1894,1920]" pageId="30" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
is found in montane forests at an elevation range of 
<quantity id="0C6E28B21E79FFE9704162F50DC9F839" box="[518,678,1929,1956]" metricMagnitude="3" metricUnit="m" metricValue="2.3425" metricValueMax="2.699" metricValueMin="1.986" pageId="30" pageNumber="481" unit="m" value="2342.5" valueMax="2699.0" valueMin="1986.0">1986–2699 m</quantity>
. Most specimens were collected from forest edges at Mpishi near Kibira National Park at 
<quantity id="0C6E28B21E79FFE973EF62D10E91F855" box="[424,510,1965,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.986" pageId="30" pageNumber="481" unit="m" value="1986.0">1986 m</quantity>
elevation. A single male specimen (MTSN 7213) was collected from Mt. Bigugu in Nyungwe Forest National Park at an elevation of 
<quantity id="0C6E28B21E79FFE970B762AD0C25F871" box="[752,842,2001,2028]" metricMagnitude="3" metricUnit="m" metricValue="2.699" pageId="30" pageNumber="481" unit="m" value="2699.0">2699 m</quantity>
. 
<emphasis id="F9E259451E79FFE9711062AD0BA6F871" bold="true" box="[855,1225,2001,2028]" pageId="30" pageNumber="481">
<taxonomicName id="0C96FED41E79FFE9711062AD0B0AF871" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[855,1125,2001,2028]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="481" phylum="Chordata" rank="species" species="bombayi" status="sp. nov.">
<emphasis id="F9E259451E79FFE9711062AD0B0AF871" bold="true" box="[855,1125,2001,2028]" italics="true" pageId="30" pageNumber="481">Rhampholeon bombayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E79FFE9762962AE0BA6F871" box="[1134,1225,2002,2028]" pageId="30" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was also found in Nyungwe Forest National Park but allopatrically to 
<emphasis id="F9E259451E78FFE870AC65EB0B4DFF2C" bold="true" box="[747,1058,150,177]" pageId="31" pageNumber="482">
<taxonomicName id="0C96FED41E78FFE870AC65EB0CD3FF2C" authority="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum, 2024" authorityName="Hughes &amp; Behangana &amp; Lukwago &amp; Menegon &amp; Dehling &amp; Wagner &amp; Tilbury &amp; South &amp; Kusamba &amp; Greenbaum" authorityYear="2024" box="[747,956,150,177]" family="Chamaeleonidae" genus="Rhampholeon" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="msitugrabensis" status="sp. nov.">
<emphasis id="F9E259451E78FFE870AC65EB0CD3FF2C" bold="true" box="[747,956,150,177]" italics="true" pageId="31" pageNumber="482">R. msitugrabensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E2D1E43E1E78FFE8718165EB0B4DFF2C" box="[966,1058,151,177]" pageId="31" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at Kamiranzovu Swamp with an elevation range of 
<quantity id="0C6E28B21E78FFE8732865C70D62FF48" box="[367,525,187,213]" metricMagnitude="3" metricUnit="m" metricValue="2.1665" metricValueMax="2.33" metricValueMin="2.003" pageId="31" pageNumber="482" unit="m" value="2166.5" valueMax="2330.0" valueMin="2003.0">2003–2330 m</quantity>
and, also at nearby Cyamudongo Forest with an elevation range of 
<quantity id="0C6E28B21E78FFE8775965C60FDFFF64" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" metricValueMax="2.033" metricValueMin="1.857" pageId="31" pageNumber="482" unit="m" value="1945.0" valueMax="2033.0" valueMin="1857.0">1857–2033 m</quantity>
. Specimens have been collected from two protected areas in the Rugege Highlands: Nyungwe Forest National Park, 
<collectingCountry id="B381C5C71E78FFE8729D647F0E56FE80" box="[218,313,259,285]" name="Rwanda" pageId="31" pageNumber="482">Rwanda</collectingCountry>
, and Kibira National Park, 
<collectingCountry id="B381C5C71E78FFE87035647F0DBEFE80" box="[626,721,259,285]" name="Burundi" pageId="31" pageNumber="482">Burundi</collectingCountry>
. Nyungwe Forest National Park is the largest protected area in 
<collectingCountry id="B381C5C71E78FFE872D0645B0F99FEDC" box="[151,246,295,321]" name="Rwanda" pageId="31" pageNumber="482">Rwanda</collectingCountry>
, and Kibira National Park is the largest protected area in 
<collectingCountry id="B381C5C71E78FFE871CE645B0C87FEDC" box="[905,1000,295,321]" name="Burundi" pageId="31" pageNumber="482">Burundi</collectingCountry>
. These parks form a continuous forest block, but the nutrient-rich soils of these highlands attract farmers in high densities, which puts pressure on the remaining forest fragments (
<bibRefCitation id="AF07F8A61E78FFE8739064130DD8FE14" author="Burgess" box="[471,695,367,393]" etAl="et al." firstAuthor="Burgess" pageId="31" pageNumber="482" pagination="164 - 177" refId="ref27352" refString="Burgess, N. D., Balmford, A., Cordeiro, N. J., Fjeldsa, J., Kuper, W., Rahbek, C., Sanderson, E. W., Scharlemann, J. P., Sommer, J. H. &amp; Williams, P. H. (2007) Correlations among species distributions, human density and human infrastructure across the high biodiversity tropical mountains of Africa. Biological Conservation, 134 (2), 164 - 177. https: // doi. org / 10.1016 / j. biocon. 2006.08.024" type="journal article" year="2007">
Burgess 
<emphasis id="F9E259451E78FFE8707D64130D04FE14" box="[570,619,367,393]" italics="true" pageId="31" pageNumber="482">et al</emphasis>
. 2007
</bibRefCitation>
). Other lizard species collected near the 
<typeStatus id="142D3BF51E78FFE8763864130BC0FE14" box="[1151,1199,367,393]" pageId="31" pageNumber="482">type</typeStatus>
locality were typical Albertine Rift lizard fauna, including 
<taxonomicName id="0C96FED41E78FFE8707464EF0D97FE30" baseAuthorityName="Boulenger" baseAuthorityYear="1906" box="[563,760,403,429]" class="Squamata" family="Lacertidae" genus="Adolfus" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="africanus">
<emphasis id="F9E259451E78FFE8707464EF0D97FE30" box="[563,760,403,429]" italics="true" pageId="31" pageNumber="482">Adolfus africanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E78FFE8714464EF0CBDFE30" box="[771,978,403,429]" class="Squamata" family="Chamaeleonidae" genus="Chamaeleo" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="dilepis">
<emphasis id="F9E259451E78FFE8714464EF0CBDFE30" box="[771,978,403,429]" italics="true" pageId="31" pageNumber="482">Chamaeleo dilepis</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E78FFE8719964EF0B98FE30" baseAuthorityName="Tornier" baseAuthorityYear="1902" box="[990,1271,403,429]" class="Squamata" family="Lacertidae" genus="Congolacerta" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="vauereselli">
<emphasis id="F9E259451E78FFE8719964EF0B98FE30" box="[990,1271,403,429]" italics="true" pageId="31" pageNumber="482">Congolacerta vauereselli</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E78FFE8774564EF0F96FE4C" baseAuthorityName="Moreau De Jonnes" baseAuthorityYear="1818" class="Squamata" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="mabouia">
<emphasis id="F9E259451E78FFE8774564EF0F96FE4C" italics="true" pageId="31" pageNumber="482">Hemidactylus mabouia</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E78FFE8734D64CB0E9EFE4C" authorityName="Lobón-Rovira, Bauer, Pinto, Trape, Conradie, Kusamba, Júlio, Cael, Stanley, Hughes, Behangana, Masudi, Pauwels &amp; Greenbaum" authorityYear="2023" box="[266,497,439,465]" class="Squamata" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="kibera">
<emphasis id="F9E259451E78FFE8734D64CB0E9EFE4C" box="[266,497,439,465]" italics="true" pageId="31" pageNumber="482">Lygodactylus kibera</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E78FFE8704564CB0D96FE4D" authorityName="Hughes, Kusamba, Behangana &amp; Greenbaum" authorityYear="2017" box="[514,761,439,464]" class="Squamata" family="Chamaeleonidae" genus="Kinyongia" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="rugegensis">
<emphasis id="F9E259451E78FFE8704564CB0D96FE4D" box="[514,761,439,464]" italics="true" pageId="31" pageNumber="482">Kinyongia rugegensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E78FFE8714C64CB0CAFFE4C" baseAuthorityName="Gunther" baseAuthorityYear="1895" box="[779,960,439,465]" class="Squamata" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="ellioti">
<emphasis id="F9E259451E78FFE8714C64CB0CAFFE4C" box="[779,960,439,465]" italics="true" pageId="31" pageNumber="482">Trioceros ellioti</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E78FFE8719564CB0B34FE4C" baseAuthorityName="Boulenger" baseAuthorityYear="1901" box="[978,1115,439,465]" class="Squamata" family="Chamaeleonidae" genus="Trioceros" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="johnstoni">
<emphasis id="F9E259451E78FFE8719564CB0C8DFE4D" box="[978,994,439,464]" italics="true" pageId="31" pageNumber="482">T</emphasis>
. 
<emphasis id="F9E259451E78FFE871B464CB0B34FE4C" box="[1011,1115,439,465]" italics="true" pageId="31" pageNumber="482">johnstoni</emphasis>
</taxonomicName>
, 
<taxonomicName id="0C96FED41E78FFE8762B64CB0A2DFE4D" baseAuthorityName="Peters" baseAuthorityYear="1844" box="[1132,1346,439,465]" class="Squamata" family="Scincidae" genus="Trachylepis" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="striata">
<emphasis id="F9E259451E78FFE8762B64CB0A2DFE4D" box="[1132,1346,439,465]" italics="true" pageId="31" pageNumber="482">Trachylepis striata</emphasis>
</taxonomicName>
, and 
<taxonomicName id="0C96FED41E78FFE877C164CB0E4AFE68" baseAuthorityName="Gray" baseAuthorityYear="1845" class="Squamata" family="Scincidae" genus="Trachylepis" kingdom="Animalia" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="maculilabris">
<emphasis id="F9E259451E78FFE877C164CB0AF9FE4D" box="[1414,1430,439,464]" italics="true" pageId="31" pageNumber="482">T</emphasis>
. 
<emphasis id="F9E259451E78FFE872D064A70E4AFE68" box="[151,293,475,501]" italics="true" pageId="31" pageNumber="482">maculilabris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>