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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.853.29407" ID-GBIF-Dataset="d707464e-b671-485c-9a74-185c0b1537b8" ID-PMC="PMC6565676" ID-Pensoft-Pub="1313-2970-853-1" ID-PubMed="31217716" ID-ZBK="82596A94ACB14332AAF84EFFBE3F95E2" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-853-1" ModsDocOrigin="ZooKeys 853" ModsDocTitle="Review of the mite genus Ololaelaps (Acari, Laelapidae) and redescription of O.formidabilis Berlese" checkinTime="1559912985079" checkinUser="pensoft" docAuthor="Beaulieu, Frederic, Quintero-Gutierrez, Edwin Javier, Sandmann, Dorothee, Klarner, Bernhard, Widyastuti, Rahayu, Combita-Heredia, Orlando &amp; Scheu, Stefan" docDate="2019" docId="1FC228EA51B4D2047385BA70CEE2ED94" docLanguage="en" docName="ZooKeys 853: 1-36" docOrigin="ZooKeys 853" docSource="http://dx.doi.org/10.3897/zookeys.853.29407" docTitle="Ololaelaps Berlese 1904" docType="treatment" docVersion="4" lastPageNumber="1" masterDocId="FFFFFFFFFFA4237B4E4A2B07FFF0FF86" masterDocTitle="Review of the mite genus Ololaelaps (Acari, Laelapidae) and redescription of O. formidabilis Berlese" masterLastPageNumber="36" masterPageNumber="1" pageNumber="1" updateTime="1668167363815" updateUser="ExternalLinkService">
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<mods:title>Review of the mite genus Ololaelaps (Acari, Laelapidae) and redescription of O. formidabilis Berlese</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beaulieu, Frederic</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Quintero-Gutierrez, Edwin Javier</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sandmann, Dorothee</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Klarner, Bernhard</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Widyastuti, Rahayu</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Combita-Heredia, Orlando</mods:namePart>
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<mods:namePart>Scheu, Stefan</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>853</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.853.29407</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.853.29407</mods:identifier>
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<treatment ID-GBIF-Taxon="157436278" LSID="urn:lsid:plazi:treatment:1FC228EA51B4D2047385BA70CEE2ED94" httpUri="http://treatment.plazi.org/id/1FC228EA51B4D2047385BA70CEE2ED94" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
Genus
<taxonomicName authority="Berlese, 1904: 260" class="Arachnida" family="Laelapidae" genus="Ololaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ololaelaps" order="Mesostigmata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Ololaelaps Berlese, 1904: 260</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Laelapidae" genus="Pristolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pristolaelaps" order="Mesostigmata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Pristolaelaps</taxonomicName>
Womersley, 1956: 571. Synonymy by
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke (1962)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Ologamasidae" genus="Laelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laelaps (Hypoaspis) venetus" order="Mesostigmata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="venetus" subGenus="Hypoaspis">Laelaps (Hypoaspis) venetus</taxonomicName>
Berlese, 1903
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="1">
(adult male and female, unless stated).Well-sclerotized hypoaspidine laelapid with a hologastric (genitiventrianal) shield in female, bearing 3-5 pairs of preanal setae (plus st5), as well as the following character states: dorsal shield covering entirely idiosoma dorsally, narrowly to broadly extending onto venter; bearing 39 or slightly fewer pairs of slender setae, including px2-3 and often one Jx. A pair of well-sclerotized presternal platelets. Female with seta st4 on sternal shield or on soft cuticle (or putatively on endopodal plate). Peritrematal shield free posteriorly or variously (narrowly) fused with hologastric and/or parapodal shields, via metapodal platelet; metapodal platelet free or variously fused to above-mentioned shields; parapodal plate well-developed, subtriangular. Soft opisthogastric cuticle with 5-10 pairs of setae. Male holoventral shield broad, fused to parapodal-exopodal plates, sometimes also to peritrematal shield. Gnathotectum convex, with few to numerous fine denticles; deutosternal groove with six rows of 1-10 denticles; female cheliceral movable digit with two teeth (rarely more), fixed digit with 3-5 (exceptionally 8); palp-apotele three-tined, third tine reduced. Leg chaetotaxy normal for
<taxonomicName family="Laelapidae" lsidName="" pageId="0" pageNumber="1" rank="family">Laelapidae</taxonomicName>
; setae generally slender.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Dorsal idiosoma.Dorsal shield relatively large (435-800 in female), broadly oval to narrowly suboval (length/width ratio 1.2-1.8), completely covering idiosoma dorsally, barely to moderately extending ventrally (this can be determined most accurately before slide-mounting); shield smooth (except for fine granulation or punctuation) to strongly reticulate;
<normalizedToken originalValue="shields">shield's</normalizedToken>
ventral extension (
<normalizedToken originalValue="epipleura">'epipleura'</normalizedToken>
of
<bibRefCitation author="Bregetova, NG" journalOrPublisher="Parazitologicheskii Sbornik" pageId="0" pageNumber="1" pagination="61 - 87" title="Mites of the genus Ololaelaps Berlese, 1904 (Acarina: Laelaptide)." volume="22" year="1964">Bregetova and Koroleva (1964)</bibRefCitation>
) smooth to reticulate (sometimes in contrast to smooth dorsal region of shield); shield with a delineated marginal strip along its edge. Dorsal shield bearing 39 pairs of simple, slender, almost hair-like setae, short to moderately long, including px2-3, and often one unpaired median seta (Jx) inserted at a level between J2 and J3 (Table 2); sometimes fewer than 39 pairs of setae, with z1 (absent in
<taxonomicName lsidName="O. sellnicki" pageId="0" pageNumber="1" rank="species" species="sellnicki">O. sellnicki</taxonomicName>
), z3 (see
<bibRefCitation author="Evans, GO" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="0" pageNumber="1" pagination="109 - 370" title="Studies on the British Dermanyssidae (Acari: Mesostigmata) Part II. Classification." volume="14" year="1966">Evans and Till 1966</bibRefCitation>
), or setae in r or S series apparently absent; shield never hypertrichous; setae slender and smooth, occasionally with a few light barbs on Z5 and J5 (
<bibRefCitation author="Jordaan, LC" journalOrPublisher="Phytophylactica" pageId="0" pageNumber="1" pagination="49 - 51" title="A new species of the genus Ololaelaps Berlese, 1904 (Acari: Laelapidae) from the Afrotropical region." volume="19" year="1987">Jordaan and Loots 1987</bibRefCitation>
). Shield with 16 pairs of poroids and four or five pairs of gland openings (based on
<bibRefCitation author="Bregetova, NG" journalOrPublisher="Parazitologicheskii Sbornik" pageId="0" pageNumber="1" pagination="61 - 87" title="Mites of the genus Ololaelaps Berlese, 1904 (Acarina: Laelaptide)." volume="22" year="1964">Bregetova and Koroleva (1964)</bibRefCitation>
and specimens examined, representing a few species only): gd1, gd2 (sometimes absent), gd4 (usually conspicuous, on or near shield margin), gd6, gd9.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Ventral idiosoma.Tritosternum normal, with two pilose laciniae. Presternal region with a pair of sclerotized platelets, wedge-shaped to subrectangular, lineate (typically with 2-4 transversal lineae); typically an additional, poorly sclerotized area, lineate and granulate, anteriorly or anteromesally adjoining each platelet. Female sternal shield as long as or longer than wide, sometimes wider than long; shield length/width ratio 0.6-1.8; Shield posterior margin straight, slightly to moderately concave, or sometimes convex; shield reticulate, smooth in its posterior fourth or fifth, bearing 3-4 pairs of simple setae and 2-3 pairs of poroids, therefore sometimes including seta st4 and poroid iv3; seta st4 on sternal shield (near or on its posterolateral edge), on soft cuticle, or apparently on endopodal plate (Table 2; see Discussion). Female hologastric shield broad, laterally extending to or beyond margin of parapodal (or adcoxal,
<bibRefCitation pageId="0" pageNumber="1">Bregetova (1977a)</bibRefCitation>
) plate, overlapping sternal and endopodal shields, bearing usually five pairs of preanal setae (JV1-3, ZV1-2) in addition to seta st5, occasionally only three or four pairs of preanals (JV3 and/or ZV2 off shield) or exceptionally six (ZV3 apparently on shield in
<taxonomicName lsidName="O. rectagoni" pageId="0" pageNumber="1" rank="species" species="rectagoni">O. rectagoni</taxonomicName>
); shield setae usually moderately long, sometimes short; hologastric shield ornamented with reticulation, cells polygonal, scale-like or elongate transversally (note that shield reticulation is not drawn for some species in
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke (1962)</bibRefCitation>
, but probably present; compare
<taxonomicName lsidName="O. mooiensis" pageId="0" pageNumber="1" rank="species" species="mooiensis">O. mooiensis</taxonomicName>
in
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke (1962)</bibRefCitation>
vs
<bibRefCitation author="Marais, JF" journalOrPublisher="Revue de zoologie et de botanique africaines" pageId="0" pageNumber="1" pagination="30 - 36" title="A new mite of the genus Ololaelaps Berlese from the Congo." volume="85" year="1972">Marais and Loots (1972)</bibRefCitation>
); circumanal setae shorter than preanals, and postanal usually shorter than paranal setae; cribrum typically with 2-3 rows of spicules. Endopodal plate besides coxae
<normalizedToken originalValue="IIIIV">III-IV</normalizedToken>
well-developed (appears reduced in
<taxonomicName lsidName="O. dililoensis" pageId="0" pageNumber="1" rank="species" species="dililoensis">O. dililoensis</taxonomicName>
, but the portion of endopodal plate that is overlapped by hologastric shield may have been overlooked); plate free, more or less contiguous with sternal shield (or apparently fused to it, e.g.,
<taxonomicName lsidName="O. expansus" pageId="0" pageNumber="1" rank="species" species="expansus">O. expansus</taxonomicName>
(
<bibRefCitation author="Ma, ML" journalOrPublisher="Acta Arachnologica Sinica" pageId="0" pageNumber="1" pagination="95 - 97" title="Discovery of the genus Pristolaelaps in China, with descriptions of a new species and a new record (Acari: Mesostigmata: Laelapidae)." volume="24" year="2015">Ma 2015</bibRefCitation>
)) or slightly overlapped by it. Peritrematal shield well-developed, fused to dorsal shield anteriorly, usually free posteriorly, reaching approximately posterior margin of coxa IV, occasionally only mid-coxa IV, or moderately surpassing coxa; sometimes narrowly connected to hologastric shield and/or parapodal element; peritrematal shield posteriorly bifid in some species (
<taxonomicName lsidName="O. interruptus" pageId="0" pageNumber="1" rank="species" species="interruptus">O. interruptus</taxonomicName>
,
<taxonomicName lsidName="O. leptochelae" pageId="0" pageNumber="1" rank="species" species="leptochelae">O. leptochelae</taxonomicName>
, and an undescribed species from North America). Peritreme narrow, usually reaching anteriorly level of coxa I, sometimes slightly less. Parapodal plate well-developed, subtriangular, with outer margin convex (especially when peritrematal shield free and not extending beyond coxa IV) or straight (typically when peritrematal shield extended posteriorly or fused to hologastric shield); parapodal exceptionally not produced in
<taxonomicName lsidName="O. rectagoni" pageId="0" pageNumber="1" rank="species" species="rectagoni">O. rectagoni</taxonomicName>
(Table 2) and an undescribed species from Costa Rica; parapodal posteriorly free, more or less abutting hologastric shield, or narrowly fused to hologastric and/or peritrematal shields, via metapodal element as connecting
<normalizedToken originalValue="bridge">'bridge'</normalizedToken>
. Metapodal platelet entirely free, suboval to strip-like, or variously fused to hologastric shield, parapodal and/or peritrematal plates. Exopodal strip well-developed, fused to parapodal element posteriorly, and anteriorly to sternal shield between coxae
<normalizedToken originalValue="III">I-II</normalizedToken>
. Soft opisthogastric cuticle surrounding shield with 5-10 pairs of simple setae, often including 1-2 pairs of r-R setae isolated at level near parapodal plate; never hypertrichous. Male holoventral shield fused to parapodal-exopodal elements, sometimes also to peritrematal shield, bearing 3-5 preanals (JV1-3, ZV1-2; JV3 and ZV2 sometimes off shield, e.g.,
<taxonomicName lsidName="O. ussuriensis" pageId="0" pageNumber="1" rank="species" species="ussuriensis">O. ussuriensis</taxonomicName>
); metapodal element merged with holoventral shield.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Gnathosoma.Gnathotectum with subtriangular to rounded margin, usually finely denticulate, may appear smooth when denticles sparse or (possibly) absent. Deutosternal groove of moderate, regular width, or slightly tapering posteriorly, with six (occasionally seven, and rarely five) rows of denticles, each row bearing 1-10 denticles, most of the rows with 3-7 denticles; denticulate rows usually preceded by a smooth ridge anteriorly, and sometimes also posteriorly. Corniculi horn-like, of moderate length. Internal malae with two pairs of long projections, median pair fimbriate on its basal portion, lateral pair smooth or branched or fimbriate in its apical portion; lateral projection absent in males (and apparently in the female of
<taxonomicName lsidName="O. sitalaensis" pageId="0" pageNumber="1" rank="species" species="sitalaensis">O. sitalaensis</taxonomicName>
). Palptarsal claw three-tined, third tine reduced. Chelicerae of moderate length, chelate-dentate; female movable digit with two moderately-sized teeth, rarely more (two additional small teeth between the two typical large teeth in
<taxonomicName lsidName="O. interruptus" pageId="0" pageNumber="1" rank="species" species="interruptus">O. interruptus</taxonomicName>
; Table 2); fixed digit with 3-5 teeth, variously sized, rarely more (eight in
<taxonomicName lsidName="O. leptochelae" pageId="0" pageNumber="1" rank="species" species="leptochelae">O. leptochelae</taxonomicName>
), including a subapical, laterally offset tooth (gabelzhan); male digits each with a single tooth; pilus dentilis setiform; arthrodial process a simple corona. Male spermatodactyl 0.7-2.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as movable digit, from its departure from edge of digit; more or less straight or variously bent; junction between spermatodactyl and movable digit straight to strongly angled (
<taxonomicName lsidName="O. translineatus" pageId="0" pageNumber="1" rank="species" species="translineatus">O. translineatus</taxonomicName>
); duct inside spermatodactyl straight or sinuous. Chaetotaxy of subcapitulum and palps normal for
<taxonomicName family="Laelapidae" lsidName="" pageId="0" pageNumber="1" rank="family">Laelapidae</taxonomicName>
(sensu
<bibRefCitation author="Evans, GO" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="0" pageNumber="1" pagination="247 - 294" title="Studies on the British Dermanyssidae (Acari: Mesostigmata). Part 1. External morphology." url="https://doi.org/10.5962/bhl.part.16752" volume="13" year="1965">Evans and Till 1965</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Legs.Chaetotaxy normal for
<taxonomicName family="Laelapidae" lsidName="" pageId="0" pageNumber="1" rank="family">Laelapidae</taxonomicName>
(sensu
<bibRefCitation author="Evans, GO" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="0" pageNumber="1" pagination="247 - 294" title="Studies on the British Dermanyssidae (Acari: Mesostigmata). Part 1. External morphology." url="https://doi.org/10.5962/bhl.part.16752" volume="13" year="1965">Evans and Till 1965</bibRefCitation>
); most setae slender; ventral and/or subapical setae of tarsi
<normalizedToken originalValue="IIIV">II-IV</normalizedToken>
usually moderately thickened, sometimes lateral setae too (e.g., al2, pl2-3 of tarsus IV); setae on other leg segments occasionally thickened (e.g., pd2, ad3 on femur I, pd on femur III in
<taxonomicName lsidName="O. placentula" pageId="0" pageNumber="1" rank="species" species="placentula">O. placentula</taxonomicName>
; also dorsally on femur IV in
<taxonomicName lsidName="O. mooiensis" pageId="0" pageNumber="1" rank="species" species="mooiensis">O. mooiensis</taxonomicName>
(
<bibRefCitation author="Jordaan, LC" journalOrPublisher="Phytophylactica" pageId="0" pageNumber="1" pagination="49 - 51" title="A new species of the genus Ololaelaps Berlese, 1904 (Acari: Laelapidae) from the Afrotropical region." volume="19" year="1987">Jordaan and Loots 1987</bibRefCitation>
)). Males of some species with a spine-like seta on femur II (
<taxonomicName lsidName="O. translineatus" pageId="0" pageNumber="1" rank="species" species="translineatus">O. translineatus</taxonomicName>
); pv thickened on genu or tibia III (in undescribed species); a ventral spine on tarsus II, apparently representing pv2 (position shifted proximad) (e.g.,
<taxonomicName lsidName="O. venetus" pageId="0" pageNumber="1" rank="species" species="venetus">O. venetus</taxonomicName>
,
<taxonomicName lsidName="O. placentula" pageId="0" pageNumber="1" rank="species" species="placentula">O. placentula</taxonomicName>
,
<taxonomicName lsidName="O. ussuriensis" pageId="0" pageNumber="1" rank="species" species="ussuriensis">O. ussuriensis</taxonomicName>
); or with cuticular tubercles on various leg segments (femur and genu of
<taxonomicName lsidName="O. placentula" pageId="0" pageNumber="1" rank="species" species="placentula">O. placentula</taxonomicName>
,
<taxonomicName lsidName="O. ussuriensis" pageId="0" pageNumber="1" rank="species" species="ussuriensis">O. ussuriensis</taxonomicName>
). Ambulacra
<normalizedToken originalValue="IIV">I-IV</normalizedToken>
with well-developed paired claws and pulvillus.
</paragraph>
<paragraph pageId="0" pageNumber="1">Spermatheca.</paragraph>
<paragraph pageId="0" pageNumber="1">Spermathecal ducts well-sclerotized and discernable in some species.</paragraph>
<caption pageId="0" pageNumber="1">
<paragraph pageId="0" pageNumber="1">
Table 2. Some diagnostic features of valid
<taxonomicName class="Arachnida" family="Laelapidae" genus="Ololaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ololaelaps" order="Mesostigmata" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Ololaelaps</taxonomicName>
species based on the literature, except for a few species for which type (**) or voucher (*) specimens were examined. Species are sorted in groups based on shared features, mainly the various fusion of shields ventrally (groups may or may not reflect relatedness).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Species</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
Shared features (mostly fusion of shields
<sup pageId="0" pageNumber="1">1</sup>
)
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
Dorsal shield ornamentation
<sup pageId="0" pageNumber="1">1</sup>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
Epipleura
<sup pageId="0" pageNumber="1">2</sup>
ornament.
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Dorsal seta Jx</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
Insertion of st4
<sup pageId="0" pageNumber="1">3</sup>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
Other features
<sup pageId="0" pageNumber="1">1</sup>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Notes and references (redescriptions)</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="venetus" pageId="0" pageNumber="1" rank="species" species="venetus">venetus</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="3">HOLOGMETAPPERITPARAPspermatod</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">JV3ZV2HOLOG</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke 1962</bibRefCitation>
<bibRefCitation author="Bregetova, NG" journalOrPublisher="Parazitologicheskii Sbornik" pageId="0" pageNumber="1" pagination="61 - 87" title="Mites of the genus Ololaelaps Berlese, 1904 (Acarina: Laelaptide)." volume="22" year="1964">Bregetova and Koroleva 1964</bibRefCitation>
<bibRefCitation author="Evans, GO" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="0" pageNumber="1" pagination="109 - 370" title="Studies on the British Dermanyssidae (Acari: Mesostigmata) Part II. Classification." volume="14" year="1966">Evans and Till 1966</bibRefCitation>
<bibRefCitation pageId="0" pageNumber="1">Bregetova 1977a</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="placidus" pageId="0" pageNumber="1" rank="species" species="placidus">placidus</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">reticulant</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Hennessey, MK" journalOrPublisher="North Carolina Agricultural Research Technical Bulletin" pageId="0" pageNumber="1" pagination="1 - 123" title="Systematic revision of thirty species of free-living, soil-inhabiting gamasine mites (Acari: Mesostigmata) of North America." volume="285" year="1988">Hennessey and Farrier 1988</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="sellnicki" pageId="0" pageNumber="1" rank="species" species="sellnicki">sellnicki</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="venetus" pageId="0" pageNumber="1" rank="species" species="venetus">venetus</taxonomicName>
reticul
<bibRefCitation author="Bregetova, NG" journalOrPublisher="Parazitologicheskii Sbornik" pageId="0" pageNumber="1" pagination="61 - 87" title="Mites of the genus Ololaelaps Berlese, 1904 (Acarina: Laelaptide)." volume="22" year="1964">Bregetova and Koroleva 1964</bibRefCitation>
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<td colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">JV3ZV2HOLOGz1z3</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Evans, GO" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="0" pageNumber="1" pagination="109 - 370" title="Studies on the British Dermanyssidae (Acari: Mesostigmata) Part II. Classification." volume="14" year="1966">Evans and Till 1966</bibRefCitation>
<bibRefCitation author="Solomon, L" journalOrPublisher="Travaux du Museum d'Histoire Naturelle &quot; Grigore Antipa &quot;" pageId="0" pageNumber="1" pagination="671 - 692" title="Contribution a la connaissance de l'acaroparasitofaune des petits mammiferes de Dobroudja." volume="8" year="1968">Solomon 1968</bibRefCitation>
<bibRefCitation pageId="0" pageNumber="1">Bregetova 1977a</bibRefCitation>
<bibRefCitation author="Kavianpour, M" journalOrPublisher="Iranian Journal of Animal Biosystematics" pageId="0" pageNumber="1" pagination="159 - 170" title="A faunistic study on laelapid mites in Urmia, Iran." volume="13" year="2017">Kavianpour et al. 2017</bibRefCitation>
<bibRefCitation pageId="0" pageNumber="1">Sellnick (1940)</bibRefCitation>
<taxonomicName lsidName="O. haemisphaericus" pageId="0" pageNumber="1" rank="species" species="haemisphaericus">O. haemisphaericus</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="hemisphaera" pageId="0" pageNumber="1" rank="species" species="hemisphaera">hemisphaera</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">HOLOGMETAPPERITPARAP</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Berlese, A" journalOrPublisher="Redia" pageId="0" pageNumber="1" pagination="289 - 338" title="Centuria terza di Acari nuovi." volume="12" year="1916 b">Berlese (1916b)</bibRefCitation>
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke (1962)</bibRefCitation>
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</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="interruptus" pageId="0" pageNumber="1" rank="species" species="interruptus">interruptus</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">HOLOGMETAPPERITPARAPPERITpost</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">MD</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="leptochelae" pageId="0" pageNumber="1" rank="species" species="leptochelae">leptochelae</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">FD</td>
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<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="burdwanensis" pageId="0" pageNumber="1" rank="species" species="burdwanensis">burdwanensis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="3">HOLOGMETAPPERIT</td>
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<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="translineatus" pageId="0" pageNumber="1" rank="species" species="translineatus">translineatus</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">spermatodMDspermath</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="O. burdwanensis" pageId="0" pageNumber="1" rank="species" species="burdwanensis">O. burdwanensis</taxonomicName>
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</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="wangi" pageId="0" pageNumber="1" rank="species" species="wangi">wangi</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="O. burdwanensis" pageId="0" pageNumber="1" rank="species" species="burdwanensis">O. burdwanensis</taxonomicName>
<bibRefCitation author="Keum, E" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="485 - 505" title="New species and new records of the family Laelapidae (Acari: Mesostigmata) from Republic of Korea." url="https://doi.org/10.11646/zootaxa.4353.3.5" volume="4353" year="2017">Keum et al. 2017</bibRefCitation>
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<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="formidabilis" pageId="0" pageNumber="1" rank="species" species="formidabilis">formidabilis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">METAPPARAP</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">reticul</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">HOLOGspermatodspermath</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="O. formidabilis" pageId="0" pageNumber="1" rank="species" species="formidabilis">O. formidabilis</taxonomicName>
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke (1962)</bibRefCitation>
METAPHOLOGPARAP
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="caucasicus" pageId="0" pageNumber="1" rank="species" species="caucasicus">caucasicus</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">HOLOGMETAP</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="placentula" pageId="0" pageNumber="1" rank="species" species="placentula">placentula</taxonomicName>
<taxonomicName lsidName="ussuriensis" pageId="0" pageNumber="1" rank="species" species="ussuriensis">ussuriensis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="O. ussuriensis" pageId="0" pageNumber="1" rank="species" species="ussuriensis">O. ussuriensis</taxonomicName>
<bibRefCitation pageId="0" pageNumber="1">Bregetova 1977a</bibRefCitation>
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</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="dililoensis" pageId="0" pageNumber="1" rank="species" species="dililoensis">dililoensis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">reticulpostreticulant</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="holaspis" pageId="0" pageNumber="1" rank="species" species="holaspis">holaspis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="7">HOLOGMETAP</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Oudemans, AC" journalOrPublisher="Memoires de la Societe zoologique de France" pageId="0" pageNumber="1" pagination="5 - 32" title="Notes sur les acariens. Xe Serie (1). Parasitidae (Vel Gamasidae) Thrombididae et Oribatidae d'Italie." volume="16" year="1903">Oudemans (1903</bibRefCitation>
<bibRefCitation pageId="0" pageNumber="1">Oudemans (1902b)</bibRefCitation>
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke (1962)</bibRefCitation>
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</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="mooiensis" pageId="0" pageNumber="1" rank="species" species="mooiensis">mooiensis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">reticul</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
METAPO
<taxonomicName lsidName="gamagarensis" pageId="0" pageNumber="1" rank="species" species="gamagarensis">gamagarensis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Marais, JF" journalOrPublisher="Revue de zoologie et de botanique africaines" pageId="0" pageNumber="1" pagination="30 - 36" title="A new mite of the genus Ololaelaps Berlese from the Congo." volume="85" year="1972">Marais and Loots 1972</bibRefCitation>
<bibRefCitation author="Nemati, A" journalOrPublisher="Persian Journal of Acarology" pageId="0" pageNumber="1" pagination="115 - 191" title="A catalogue of the Iranian Mesostigmata (Acari): Additions and updates of the previous catalogue." url="10.22073/pja.v7i2.36985" volume="7" year="2018">Nemati et al. 2018</bibRefCitation>
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</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="placentula" pageId="0" pageNumber="1" rank="species" species="placentula">placentula</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PERIT</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation pageId="0" pageNumber="1">Sellnick 1940</bibRefCitation>
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke 1962</bibRefCitation>
<bibRefCitation author="Bregetova, NG" journalOrPublisher="Parazitologicheskii Sbornik" pageId="0" pageNumber="1" pagination="61 - 87" title="Mites of the genus Ololaelaps Berlese, 1904 (Acarina: Laelaptide)." volume="22" year="1964">Bregetova and Koroleva 1964</bibRefCitation>
<bibRefCitation author="Evans, GO" journalOrPublisher="Bulletin of the British Museum (Natural History), Zoology" pageId="0" pageNumber="1" pagination="109 - 370" title="Studies on the British Dermanyssidae (Acari: Mesostigmata) Part II. Classification." volume="14" year="1966">Evans and Till 1966</bibRefCitation>
<bibRefCitation author="Bai, X-L" journalOrPublisher="Acta Arachnologica Sinica" pageId="0" pageNumber="1" pagination="29 - 31" title="A new record of the genus Ololaelaps and a new record of the genus Ameroseius from China (Acari: Mesostigmata: Laelapidae, Ameroseiidae)." volume="23" year="2014">Bai and Ma 2014</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="platensis" pageId="0" pageNumber="1" rank="species" species="platensis">platensis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ZV1</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke 1962</bibRefCitation>
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</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="rectagoni" pageId="0" pageNumber="1" rank="species" species="rectagoni">rectagoni</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">j1HOLOGPARAPZV3HOLOG</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Karg, W" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="1" pagination="179 - 216" title="Raubmilben der Cohors Gamasina Leach (Acarina, Parasitiformes) vom Galapagos-Archipel." url="https://doi.org/10.1002/mmnz.19940700202" volume="70" year="1994">Karg 1994</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="sinensis" pageId="0" pageNumber="1" rank="species" species="sinensis">sinensis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Ryke, PAJ" journalOrPublisher="Revista de Biologia" pageId="0" pageNumber="1" pagination="124 - 130" title="The genus Ololaelaps Berlese (Acarina: Laelaptidae)." volume="3" year="1962">Ryke 1962</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="ussuriensis" pageId="0" pageNumber="1" rank="species" species="ussuriensis">ussuriensis</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">reticul</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation pageId="0" pageNumber="1">Bregetova 1977a</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="bregetovae" pageId="0" pageNumber="1" rank="species" species="bregetovae">bregetovae</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="7">HOLOGMETAPPERITPARAP</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">post</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="O. tasmanicus" pageId="0" pageNumber="1" rank="species" species="tasmanicus">O. tasmanicus</taxonomicName>
<taxonomicName lsidName="O. sitalensis" pageId="0" pageNumber="1" rank="species" species="sitalensis">O. sitalensis</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="expansus" pageId="0" pageNumber="1" rank="species" species="expansus">expansus</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="nasri" pageId="0" pageNumber="1" rank="species" species="nasri">nasri</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="O. obovatus" pageId="0" pageNumber="1" rank="species" species="obovatus">O. obovatus</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="obovatus" pageId="0" pageNumber="1" rank="species" species="obovatus">obovatus</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ZV1</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="paratasmanicus" pageId="0" pageNumber="1" rank="species" species="paratasmanicus">paratasmanicus</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">HOLOG</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="O. tasmanicus" pageId="0" pageNumber="1" rank="species" species="tasmanicus">O. tasmanicus</taxonomicName>
<bibRefCitation author="Ma, ML" journalOrPublisher="Acta Arachnologica Sinica" pageId="0" pageNumber="1" pagination="95 - 97" title="Discovery of the genus Pristolaelaps in China, with descriptions of a new species and a new record (Acari: Mesostigmata: Laelapidae)." volume="24" year="2015">Ma 2015</bibRefCitation>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="sitalaensis" pageId="0" pageNumber="1" rank="species" species="sitalaensis">sitalaensis</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="tasmanicus" pageId="0" pageNumber="1" rank="species" species="tasmanicus">tasmanicus</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Tenorio, JM" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="1" pagination="259 - 274" title="Hypoaspidinae (Acari: Gamasida: Laelapidae) of the Hawaiian Islands." volume="24" year="1982">Tenorio (1982)</bibRefCitation>
<bibRefCitation author="Womersley, H" journalOrPublisher="The Journal of the Linnean Society of London, Zoology" pageId="0" pageNumber="1" pagination="505 - 599" title="On some new Acarina-Mesostigmata from Australia, New Zealand and New Guinea." url="https://doi.org/10.1111/j.1096-3642.1956.tb02218.x" volume="42" year="1956">Womersley (1956)</bibRefCitation>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Tenorio, JM" journalOrPublisher="Pacific Insects" pageId="0" pageNumber="1" pagination="259 - 274" title="Hypoaspidinae (Acari: Gamasida: Laelapidae) of the Hawaiian Islands." volume="24" year="1982">Tenorio 1982</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>