treatments-xml/data/17/56/87/17568786FFE0735E23A805DBFAF0C2A8.xml

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<document id="33F5C1EA29F07CAA588AE93F3DDF59A1" ID-CLB-Dataset="8966" ID-DOI="10.11646/zootaxa.4820.3.8" ID-GBIF-Dataset="f702f2bf-e817-4863-96f7-f20015f61de3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4398204" ID-ZooBank="B8AAC629-F431-49E2-B15C-80CCC0CE070C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609213172128" checkinUser="plazi" docAuthor="Alekseev, Vitalii Igorevich, Bukejs, Andris &amp; Pollock, Darren Andrew" docDate="2020" docId="17568786FFE0735E23A805DBFAF0C2A8" docLanguage="en" docName="zootaxa.4820.3.8.pdf" docOrigin="Zootaxa 4820 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Europoeurypus inglaeso Alekseev &amp; Bukejs &amp; Pollock 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="556" masterDocId="EB6FFFFEFFE3735B233F0116FFEBC10E" masterDocTitle="A new fossil Eurypinae genus and sexual dimorphism in the representatives of Omineus (Coleoptera: Mycteridae) from Tertiary amber of Europe" masterLastPageNumber="562" masterPageNumber="551" pageNumber="554" updateTime="1698848390187" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6D77FCBA6929E9DAD27AD25913CBD67C">A new fossil Eurypinae genus and sexual dimorphism in the representatives of Omineus (Coleoptera: Mycteridae) from Tertiary amber of Europe</mods:title>
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<mods:namePart id="DAAED0073D57E873D93776E386DCFB78">Alekseev, Vitalii Igorevich</mods:namePart>
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<mods:affiliation id="86C138BC33726AD5A2EEBD6F96BCB50D">Shirshov Institute of Oceanology, Russian Academy of Sciences, Nahimovskiy prospekt 36, 117997 Moscow, Russia. alekseew 0802 @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4390 - 5443 &amp; Kaliningrad Regional Amber Museum, Marshal Vasilevskii square 1, Kaliningrad 236016, Russia.</mods:affiliation>
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<mods:namePart id="AFB5A72CDD73C1A8BDDB15A918A1A84E">Bukejs, Andris</mods:namePart>
<mods:affiliation id="AE5C6873D1EB4403DA3A10183645FBA7">Institute of Life Sciences and Technologies, Daugavpils University, Vienîbas 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<mods:namePart id="CF3A8EC8D3667BE648CA714DA8330D97">Pollock, Darren Andrew</mods:namePart>
<mods:affiliation id="5ABE74EDB20D978D289673CCE8C54309">Department of Biology, Eastern New Mexico University, Portales, NM 88130 USA.</mods:affiliation>
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<paragraph id="9F403690FFE0735823A805DBFDCFC5E9" blockId="3.[151,548,1228,1291]" box="[151,548,1228,1255]" pageId="3" pageNumber="554">
<heading id="C40881FCFFE0735823A805DBFDCFC5E9" bold="true" box="[151,548,1228,1255]" fontSize="11" level="1" pageId="3" pageNumber="554" reason="1">
<emphasis id="AD8BEA82FFE0735823A805DBFDCFC5E9" bold="true" box="[151,548,1228,1255]" pageId="3" pageNumber="554">
<taxonomicName id="58FF4D13FFE0735823A805DBFE54C5E9" authority="Alekseev &amp; Bukejs &amp; Pollock, 2020" authorityName="Alekseev &amp; Bukejs &amp; Pollock" authorityYear="2020" box="[151,447,1228,1255]" class="Insecta" family="Mycteridae" genus="Europoeurypus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="554" phylum="Arthropoda" rank="species" species="inglaeso" status="sp. nov.">
<emphasis id="AD8BEA82FFE0735823A805DBFE54C5E9" bold="true" box="[151,447,1228,1255]" italics="true" pageId="3" pageNumber="554">Europoeurypus inglaeso</emphasis>
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<taxonomicNameLabel id="B6B857F9FFE0735822F905DBFDCFC5E9" box="[454,548,1229,1255]" pageId="3" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="D7E5651BFFE0735823A805E6FEF9C405" box="[151,274,1264,1291]" pageId="3" pageNumber="554" type="description">
<paragraph id="9F403690FFE0735823A805E6FEF9C405" blockId="3.[151,548,1228,1291]" box="[151,274,1264,1291]" pageId="3" pageNumber="554">
(
<figureCitation id="07C42A15FFE0735823A005E6FEE2C405" box="[159,265,1264,1291]" captionStart-0="FIGURES 12" captionStart-1="FIGURE 34" captionStart-2="FIGURE 5" captionStartId-0="2.[151,264,1899,1923]" captionStartId-1="3.[151,250,1050,1074]" captionStartId-2="4.[151,250,1955,1979]" captionTargetBox-0="[151,1435,179,1872]" captionTargetBox-1="[151,1436,181,1026]" captionTargetBox-2="[252,1339,315,1931]" captionTargetId-0="figure-23@2.[151,1435,179,1872]" captionTargetId-1="figure-18@3.[151,1436,181,1026]" captionTargetId-2="figure-78@4.[245,1342,309,1932]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 12. Europoeurypus inglaeso gen. et sp. nov., holotype, No MAIG 6081: 1—habitus, dorsal view; 2—habitus, ven- tral view. Scale bars represent 4.0 mm." captionText-1="FIGURE 34. Europoeurypus inglaeso gen. et sp. nov., holotype, No MAIG 6081: 3—abdomen, showing ventrites without discernible sex patches; 4—details of forebody, dorsal view. Abbreviations: v1v5—abdominal ventrites 15 respectively; icp—intercoxal process of ventrite 1. Scale bars represent 1.0 mm." captionText-2="FIGURE 5. Reconstruction of Europoeurypus inglaeso gen. et sp. nov. from Baltic amber: habitus, dorsal view. Body length = 10.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4398206" figureDoi-1="http://doi.org/10.5281/zenodo.4398208" figureDoi-2="http://doi.org/10.5281/zenodo.4398210" httpUri-0="https://zenodo.org/record/4398206/files/figure.png" httpUri-1="https://zenodo.org/record/4398208/files/figure.png" httpUri-2="https://zenodo.org/record/4398210/files/figure.png" pageId="3" pageNumber="554">Figs. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D7E5651BFFE0735823A8042EFF02C740" pageId="3" pageNumber="554" type="materials_examined">
<paragraph id="9F403690FFE0735823A8042EFC0DC4B0" blockId="3.[151,1437,1336,2011]" pageId="3" pageNumber="554">
<emphasis id="AD8BEA82FFE0735823A8042EFE91C45C" bold="true" box="[151,378,1336,1362]" pageId="3" pageNumber="554">Material examined</emphasis>
.
<materialsCitation id="2F973CCDFFE0735822B7042EFC0DC4B0" ID-GBIF-Occurrence="3014938302" collectionCode="MAIG" collectorName="Jonas Damzen" pageId="3" pageNumber="554" specimenCount="1" typeStatus="holotype">
<typeStatus id="40448832FFE0735822B7042EFE1EC45C" box="[392,501,1336,1362]" pageId="3" pageNumber="554" type="holotype">Holotype</typeStatus>
: No. 6081 [
<collectionCode id="F9EEAE55FFE07358214A042EFD2EC45D" box="[629,709,1336,1363]" pageId="3" pageNumber="554">MAIG</collectionCode>
] (
<collectorName id="320A5346FFE0735821E3042FFCC7C45D" box="[732,812,1337,1363]" pageId="3" pageNumber="554">No. JD</collectorName>
8112 ex coll.
<collectorName id="320A5346FFE0735820F4042FFB98C45D" box="[971,1139,1337,1363]" pageId="3" pageNumber="554">Jonas Damzen</collectorName>
), adult, apparently female.
<collectorName id="320A5346FFE0735823A8044AFF02C478" box="[151,233,1372,1398]" pageId="3" pageNumber="554">Almost</collectorName>
complete beetle inclusion (tarsomeres present on right pro- and mesotarsus only, apical parts of all left tibiae missing) included in transparent, flat, oval, orange piece of amber with dimensions 24×14×
<quantity id="58079B75FFE0735827B50497FB26C495" box="[1162,1229,1409,1435]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="554" unit="mm" value="4.0">4 mm</quantity>
.
<collectorName id="320A5346FFE0735827E80496FA9FC494" box="[1239,1396,1408,1434]" pageId="3" pageNumber="554">Syninclusions</collectorName>
are represented by detrital particles and a few stellate trichomes of plant origin.
</materialsCitation>
</paragraph>
<paragraph id="9F403690FFE0735823F804DFFB48C709" blockId="3.[151,1437,1336,2011]" pageId="3" pageNumber="554">
<emphasis id="AD8BEA82FFE0735823F804DFFEBAC4ED" bold="true" box="[199,337,1481,1507]" pageId="3" pageNumber="554">
<typeStatus id="40448832FFE0735823F804DFFEE9C4ED" box="[199,258,1481,1507]" pageId="3" pageNumber="554">Type</typeStatus>
strata
</emphasis>
. Baltic amber from Eocene amber-bearing
<emphasis id="AD8BEA82FFE07358207B04DEFC28C4EC" box="[836,963,1480,1506]" italics="true" pageId="3" pageNumber="554">Blaue Erde</emphasis>
deposits (mostly Bartonian age, as interpreted for extinct Central European resin-producing forests according to
<bibRefCitation id="FB6E4B61FFE0735820F804FAFB73C709" author="Bukejs, A. &amp; Alekseev, V. I. &amp; Pollock, D. A." box="[967,1176,1516,1543]" pageId="3" pageNumber="554" pagination="261 - 273" refId="ref4803" refString="Bukejs, A., Alekseev, V. I. &amp; Pollock, D. A. (2019) Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa, 4664 (2), 261 - 273. https: // doi. org / 10.11646 / zootaxa. 4664.2.8" type="journal article" year="2019">
Bukejs
<emphasis id="AD8BEA82FFE07358272304FBFBA6C708" box="[1052,1101,1516,1542]" italics="true" pageId="3" pageNumber="554">et al</emphasis>
. 2019
</bibRefCitation>
).
</paragraph>
<paragraph id="9F403690FFE0735823F80707FF02C740" blockId="3.[151,1437,1336,2011]" pageId="3" pageNumber="554">
<materialsCitation id="2F973CCDFFE0735823F80707FF0EC740" ID-GBIF-Occurrence="3014938303" country="Russia" county="Peninsula" location="Yantarny" municipality="Sambian" pageId="3" pageNumber="554" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="AD8BEA82FFE0735823F80707FEB4C724" bold="true" box="[199,351,1552,1579]" pageId="3" pageNumber="554">
<typeStatus id="40448832FFE0735823F80707FEE9C725" box="[199,258,1553,1579]" pageId="3" pageNumber="554">Type</typeStatus>
locality
</emphasis>
.
<location id="9A20604BFFE0735822540707FE3BC725" LSID="urn:lsid:plazi:treatment:17568786FFE0735E23A805DBFAF0C2A8:9A20604BFFE0735822540707FE3BC725" box="[363,464,1553,1579]" country="Russia" county="Peninsula" municipality="Sambian" name="Yantarny" pageId="3" pageNumber="554" stateProvince="Kaliningrad">Yantarny</location>
settlement (formerly Palmnicken),
<collectingMunicipality id="7F24ACEAFFE0735820630706FC55C724" box="[860,958,1552,1578]" pageId="3" pageNumber="554">Sambian</collectingMunicipality>
(Samland)
<collectingCounty id="76214E1CFFE0735827000706FB44C724" box="[1087,1199,1552,1578]" pageId="3" pageNumber="554">Peninsula</collectingCounty>
,
<collectingRegion id="5D3BF872FFE0735827870706FA73C724" box="[1208,1432,1552,1578]" country="Russia" name="Kaliningrad" pageId="3" pageNumber="554">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="E7E87600FFE0735823A80722FF0EC740" box="[151,229,1588,1614]" name="Russia" pageId="3" pageNumber="554">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D7E5651BFFE0735823F8074EFD3EC77D" box="[199,725,1624,1651]" pageId="3" pageNumber="554" type="diagnosis">
<paragraph id="9F403690FFE0735823F8074EFD3EC77D" blockId="3.[151,1437,1336,2011]" box="[199,725,1624,1651]" pageId="3" pageNumber="554">
<emphasis id="AD8BEA82FFE0735823F8074EFED1C77C" bold="true" box="[199,314,1624,1650]" pageId="3" pageNumber="554">Diagnosis</emphasis>
. As stated above, for the new genus.
</paragraph>
</subSubSection>
<subSubSection id="D7E5651BFFE0735E23F8076AFB84C234" lastPageId="5" lastPageNumber="556" pageId="3" pageNumber="554" type="description">
<paragraph id="9F403690FFE0735823F8076AFBEFC7B4" blockId="3.[151,1437,1336,2011]" pageId="3" pageNumber="554">
<emphasis id="AD8BEA82FFE0735823F8076AFEBAC798" bold="true" box="[199,337,1660,1686]" pageId="3" pageNumber="554">Description</emphasis>
.
<emphasis id="AD8BEA82FFE07358225F076AFE71C798" box="[352,410,1660,1686]" italics="true" pageId="3" pageNumber="554">Body</emphasis>
relatively large-sized, elongate, slightly convex dorsally; dark, unicolorous. Dorsal surface evenly punctured and covered with fine, decumbent, conspicuous pubescence.
</paragraph>
<paragraph id="9F403690FFE0735823F807D3FC22C629" blockId="3.[151,1437,1336,2011]" pageId="3" pageNumber="554">
<emphasis id="AD8BEA82FFE0735823F807D3FE82C7D0" box="[199,361,1733,1758]" italics="true" pageId="3" pageNumber="554">Measurements</emphasis>
: body length
<quantity id="58079B75FFE07358213807D3FD85C7D1" box="[519,622,1733,1759]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="554" unit="mm" value="10.2">10.2 mm</quantity>
, maximum body width
<quantity id="58079B75FFE07358204407D3FC39C7D1" box="[891,978,1733,1759]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="554" unit="mm" value="3.3">3.3 mm</quantity>
(elytra postmedially); head length about 1.0 mm, maximum width of head (across eyes)
<quantity id="58079B75FFE07358219707FFFCEBC60D" box="[680,768,1769,1795]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="554" unit="mm" value="1.8">1.8 mm</quantity>
; pronotal length
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, maximum pronotal width
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; elytral length
<quantity id="58079B75FFE07358220C061AFE60C629" box="[307,395,1804,1831]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="554" unit="mm" value="7.5">7.5 mm</quantity>
, maximum elytral width (in humeral area)
<quantity id="58079B75FFE073582053061BFC2EC629" box="[876,965,1805,1831]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="554" unit="mm" value="2.8">2.8 mm</quantity>
.
</paragraph>
<paragraph id="9F403690FFE0735F23F80626FD62C013" blockId="3.[151,1437,1336,2011]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="555" pageId="3" pageNumber="554">
<emphasis id="AD8BEA82FFE0735823F80626FEEFC644" box="[199,260,1840,1866]" italics="true" pageId="3" pageNumber="554">Head</emphasis>
moderately narrowed posterior of eyes. Compound eyes protruding, transverse, without intrafacetal setae, widely separated, with entire and very slightly emarginate anterior margin; ommatidial facets distinct and fine (as large as punctures on head). Frons convex, minimum distance between eyes 2.8× as wide as longitudinal eye diameter measured laterally. Ocular (suborbital) grooves indistinct. Frontal furrows present, straight, subparallel. Labrum with anterior margin almost straight, bearing long setae. Mandibles bidentate apically. Terminal maxillary palpomere broadly triangular. Antennal insertion slightly hidden from above. Antennae 11-segmented, short, reaching posteriorly to near one-third of pronotum, pubescent. Antennomeres 210 cylindrical, about 2× as long as wide; scape and antennomere 11 wider than other antennomeres; scape subquadrate; antennomere 11 ovoid, pointed api-cally, about 1.5× as long as antennomere 10.
</paragraph>
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<paragraph id="9F403690FFE7735F23A806B5FEE6C6D1" blockId="4.[151,1436,1955,2016]" pageId="4" pageNumber="555">
<emphasis id="AD8BEA82FFE7735F23A806B5FEFFC6B2" bold="true" box="[151,276,1955,1980]" pageId="4" pageNumber="555">FIGURE 5.</emphasis>
Reconstruction of
<taxonomicName id="58FF4D13FFE7735F22E706B2FD3BC6B5" authority="Alekseev &amp; Bukejs &amp; Pollock, 2020" authorityName="Alekseev &amp; Bukejs &amp; Pollock" authorityYear="2020" box="[472,720,1955,1979]" class="Insecta" family="Mycteridae" genus="Europoeurypus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="555" phylum="Arthropoda" rank="species" species="inglaeso" status="gen. et sp. nov.">
<emphasis id="AD8BEA82FFE7735F22E706B2FD3BC6B5" box="[472,720,1955,1979]" italics="true" pageId="4" pageNumber="555">Europoeurypus inglaeso</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B6B857F9FFE7735F21E806B2FC9EC6B2" box="[727,885,1956,1980]" pageId="4" pageNumber="555" rank="species">
<emphasis id="AD8BEA82FFE7735F21E806B2FCEFC6B2" bold="true" box="[727,772,1956,1980]" pageId="4" pageNumber="555">gen.</emphasis>
et
<emphasis id="AD8BEA82FFE7735F201B06B2FC9EC6B2" bold="true" box="[804,885,1956,1980]" pageId="4" pageNumber="555">sp. nov.</emphasis>
</taxonomicNameLabel>
from Baltic amber: habitus, dorsal view. Body length = 10.2 mm.
</paragraph>
</caption>
<paragraph id="9F403690FFE6735E23F80181FD23C317" blockId="5.[151,1437,151,934]" pageId="5" pageNumber="556">
<emphasis id="AD8BEA82FFE6735E23F80181FEF3C1BF" box="[199,280,151,177]" italics="true" pageId="5" pageNumber="556">Thorax</emphasis>
. Pronotum transverse, 1.3× as wide as long, widest at middle. Pronotal base distinctly narrower than elytral base. Anterior and lateral margins of pronotum slightly rounded; posterior margin slightly sinuate. Lateral pronotal carina present, discernible in ventral view in two-thirds of pronotal length basally. Anterior and posterior angles rounded, not produced. Pronotal disc slightly convex, uneven, with pair of small symmetrical triangular depressions at base; pronotum densely and finely punctate, distance between punctures equal to 0.20.3× diameter of one puncture. Prosternum long anterior of procoxae. Procoxal cavities apparently open posteriorly. Elytra elongate, disc moderately convex; dorsal surface evenly and irregularly punctate and pubescent. Pubescence decumbent, long (equal to 5.06.0× distance between punctures); distance between punctures equal to 0.51.0× diameter of one puncture. Scutellar shield as long as wide, rounded, densely pubescent, pointed posteriorly. Mesocoxal cavities open. Elytral epipleura narrow, widest in basal part and then gradually narrowed posteriorly, extended to abdominal ventrite 2. Metaventrite convex, densely punctate.
</paragraph>
<paragraph id="9F403690FFE6735E23F80335FC07C3A7" blockId="5.[151,1437,151,934]" pageId="5" pageNumber="556">
<emphasis id="AD8BEA82FFE6735E23F80335FF17C332" box="[199,252,547,572]" italics="true" pageId="5" pageNumber="556">Legs</emphasis>
slender. Procoxae subconical, contiguous. Meso- and metacoxae narrowly separated. Femora flattened. Tibiae straight, subequal in length to femora, with two short, equal in length spurs apically. Pro- and mesotarsi pen-tamerous. Penultimate tarsomere distinctly widened distally, flat, non-lobed. Tarsal claws swollen basally, without acute basal tooth, symmetrical. Ratio of mesotarsomere lengths 5: 3: 3: 4: 6.
</paragraph>
<paragraph id="9F403690FFE6735E23F803A5FB84C234" blockId="5.[151,1437,151,934]" pageId="5" pageNumber="556">
<emphasis id="AD8BEA82FFE6735E23F803A5FEDAC3C3" box="[199,305,691,717]" italics="true" pageId="5" pageNumber="556">Abdomen</emphasis>
with five visible abdominal ventrites. Ventrites 13 connate, ventrites 45 free; sutures between ven-trites 12 and 23 flat, sutures between ventrites 34 and 45 shelf-like. Anterior (intercoxal) process of ventrite 1 broadly triangular, pointed apically. All ventrites punctate, setose, without discernible sex patch(es). Apical margin of ventrite 5 rounded. Ratio of lengths of ventrites 15 3.5: 3.5: 3.5: 3.0: 2.5 (medially).
</paragraph>
</subSubSection>
<subSubSection id="D7E5651BFFE6735E23F80255FA60C28F" pageId="5" pageNumber="556" type="etymology">
<paragraph id="9F403690FFE6735E23F80255FA60C28F" blockId="5.[151,1437,151,934]" pageId="5" pageNumber="556">
<emphasis id="AD8BEA82FFE6735E23F80255FEA3C253" bold="true" box="[199,328,835,861]" pageId="5" pageNumber="556">Etymology</emphasis>
. The specific epithet
<taxonomicName id="58FF4D13FFE6735E217E0255FD4BC253" authorityName="Alekseev &amp; Bukejs &amp; Pollock" authorityYear="2020" box="[577,672,835,861]" class="Insecta" family="Mycteridae" genus="Europoeurypus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="556" phylum="Arthropoda" rank="species" species="inglaeso">
<emphasis id="AD8BEA82FFE6735E217E0255FD4BC253" box="[577,672,835,861]" italics="true" pageId="5" pageNumber="556">inglaeso</emphasis>
</taxonomicName>
is a Latin word combination of “in glaeso” (meaning “in amber”), used as a noun in apposition and created analogously to the widely used biological terms “
<emphasis id="AD8BEA82FFE6735E27B30271FB33C28E" box="[1164,1240,871,896]" italics="true" pageId="5" pageNumber="556">in vivo</emphasis>
” and “
<emphasis id="AD8BEA82FFE6735E26180271FA93C28E" box="[1319,1400,871,896]" italics="true" pageId="5" pageNumber="556">in vitro</emphasis>
”.
</paragraph>
</subSubSection>
<subSubSection id="D7E5651BFFE6735E23F8029DFAF0C2A8" box="[199,1307,907,934]" pageId="5" pageNumber="556" type="discussion">
<paragraph id="9F403690FFE6735E23F8029DFAF0C2A8" blockId="5.[151,1437,151,934]" box="[199,1307,907,934]" pageId="5" pageNumber="556">
<emphasis id="AD8BEA82FFE6735E23F8029DFEDFC2AB" bold="true" box="[199,308,907,933]" pageId="5" pageNumber="556">Remarks</emphasis>
. The
<typeStatus id="40448832FFE6735E224B029DFE3DC2AB" box="[372,470,907,933]" pageId="5" pageNumber="556" type="holotype">holotype</typeStatus>
apparently is female due to the lack of sex patch on abdominal ventrite(s).
</paragraph>
</subSubSection>
</treatment>
</document>