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<document id="D8B468ABD0524BD3F323F4585D93ED89" ID-CLB-Dataset="32711" ID-DOI="10.11646/zootaxa.4290.2.5" ID-GBIF-Dataset="1098c6ff-7d84-44cf-86ea-a31dd0707acd" ID-ISSN="1175-5326" ID-Zenodo-Dep="829113" ID-ZooBank="C33B5FA7-B001-415A-AD89-8D4CA5549C1D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500110851585" checkinUser="plazi" docAuthor="Mayoral, J. G. &amp; Barranco, P." docDate="2017" docId="03FD87F4591CFFBEFF65FD9AFD7AF835" docLanguage="en" docName="zootaxa.4290.2.5.pdf" docOrigin="Zootaxa 4290 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eukoenenia indalica Mayoral &amp; Barranco, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="361" masterDocId="FFC4FF8C591DFFBAFFF2FFDEFFF6FFC0" masterDocTitle="A remarkable new palpigrade (Arachnida) of the Eukoenenia mirabilis group from Spain" masterLastPageNumber="366" masterPageNumber="357" pageNumber="358" updateTime="1698465422114" updateUser="ExternalLinkService">
<mods:mods id="2F4943A767F012E0D7C52C439A2A1508" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="63C7AF004D9B8E33F48009548A663889">
<mods:title id="7B035320513FA6FAD9D7A7EBFBD35746">A remarkable new palpigrade (Arachnida) of the Eukoenenia mirabilis group from Spain</mods:title>
</mods:titleInfo>
<mods:name id="D2DCB6DD7FCFD0AA0AB5E95E7C9040D2" type="personal">
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<mods:namePart id="62C36F155CD6BF34D67F28CB0A598E84">Mayoral, J. G.</mods:namePart>
</mods:name>
<mods:name id="96F73758B749D65AA4F3F9C9D0E02A33" type="personal">
<mods:role id="422037A55B1E1AAA9617AD610FC896AC">
<mods:roleTerm id="19247F42C2EAB71BBDD064CFA38B684D">Author</mods:roleTerm>
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<mods:namePart id="B49B08A7084FDC37DA357C3C90B08F28">Barranco, P.</mods:namePart>
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<mods:typeOfResource id="8C57AC4EDAE39029EBF31CDAE6B02397">text</mods:typeOfResource>
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<mods:title id="33FD18A9BDEA9C54D1739192DC34EF3D">Zootaxa</mods:title>
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<mods:part id="FE5B6631B86A8E537E2F97235A852348">
<mods:date id="DE4129539694E34CC1AA369EE980B4F7">2017</mods:date>
<mods:detail id="1D5E035E5100A9A1EE19D8C370C81177" type="volume">
<mods:number id="BA6F96F1601BB45F727932AB483BD7F2">4290</mods:number>
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<mods:detail id="589DD0FD2BD79567FBD2D27803E54EE6" type="issue">
<mods:number id="9812B6B7FDA79B4D9D5E5B5059CBDF52">2</mods:number>
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<mods:classification id="1AC2F8743BB3D4B66EC16BEA66378181">journal article</mods:classification>
<mods:identifier id="5CD782139826E74AB9B26A7B3129C0F6" type="CLB-Dataset">32711</mods:identifier>
<mods:identifier id="F7C757E10194D7BA01395B512676CCEA" type="DOI">10.11646/zootaxa.4290.2.5</mods:identifier>
<mods:identifier id="0595BDB1C527553D896D2838CB1981F7" type="GBIF-Dataset">1098c6ff-7d84-44cf-86ea-a31dd0707acd</mods:identifier>
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<mods:identifier id="7940C4BAD72C4A3D0DC2D07C745ECA2D" type="Zenodo-Dep">829113</mods:identifier>
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</mods:mods>
<treatment id="03FD87F4591CFFBEFF65FD9AFD7AF835" ID-DOI="http://doi.org/10.5281/zenodo.6049179" ID-GBIF-Taxon="132549398" ID-Zenodo-Dep="6049179" LSID="urn:lsid:plazi:treatment:03FD87F4591CFFBEFF65FD9AFD7AF835" httpUri="http://treatment.plazi.org/id/03FD87F4591CFFBEFF65FD9AFD7AF835" lastPageId="4" lastPageNumber="361" pageId="1" pageNumber="358">
<subSubSection id="C34E6569591CFFBBFF65FD9AFED6FDBF" pageId="1" pageNumber="358" type="nomenclature">
<paragraph id="8BEB36E2591CFFBBFF65FD9AFE01FD9E" blockId="1.[151,503,580,639]" box="[151,503,580,606]" pageId="1" pageNumber="358">
<heading id="D0A3818E591CFFBBFF65FD9AFE01FD9E" bold="true" box="[151,503,580,606]" fontSize="11" level="1" pageId="1" pageNumber="358" reason="1">
<emphasis id="B920EAF0591CFFBBFF65FD9AFE01FD9E" bold="true" box="[151,503,580,606]" pageId="1" pageNumber="358">
<taxonomicName id="4C544D61591CFFBBFF65FD9AFE65FD9E" ID-CoL="6H7QF" box="[151,403,580,606]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="1" pageNumber="358" phylum="Arthropoda" rank="species" species="indalica" status="sp. nov.">
<emphasis id="B920EAF0591CFFBBFF65FD9AFE65FD9E" bold="true" box="[151,403,580,606]" italics="true" pageId="1" pageNumber="358">Eukoenenia indalica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213578B591CFFBBFE68FD9AFE01FD9E" box="[410,503,580,606]" pageId="1" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEB36E2591CFFBBFF65FDB8FED6FDBF" blockId="1.[151,503,580,639]" box="[151,288,614,639]" pageId="1" pageNumber="358">
(
<figureCitation id="136F2A67591CFFBBFF6DFDB8FEE1FDBF" box="[159,279,614,639]" captionStart-0="FIGURE 1 – 7" captionStart-1="FIGURE 8 – 9" captionStart-2="FIGURE 10 – 11" captionStart-3="FIGURE 12 – 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStartId-0="2.[151,250,1937,1959]" captionStartId-1="3.[151,250,1994,2016]" captionStartId-2="4.[151,250,1585,1607]" captionStartId-3="5.[151,250,1831,1853]" captionStartId-4="6.[151,250,1806,1828]" captionStartId-5="7.[151,250,1962,1984]" captionTargetBox-0="[272,1307,200,1908]" captionTargetBox-1="[197,1405,202,1966]" captionTargetBox-2="[180,1415,204,1561]" captionTargetBox-3="[171,1427,206,1797]" captionTargetBox-4="[159,1418,1214,1778]" captionTargetBox-5="[167,1430,851,1910]" captionTargetId-0="figure@2.[260,1320,193,1914]" captionTargetId-1="figure@3.[180,1416,193,1978]" captionTargetId-2="figure@4.[166,1430,193,1575]" captionTargetId-3="figure@5.[151,1436,195,1809]" captionTargetId-4="figure@6.[153,1433,1201,1784]" captionTargetId-5="figure@7.[151,1436,844,1920]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1 – 7. Eukoeneneia indalica sp nov. 1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2)." captionText-1="FIGURE 8 – 9. Eukoenenia indalica sp. nov. 8, ta 3 – bta 1 of left palp; 9, ti – tc of right palp. Scale bar 100 µm." captionText-2="FIGURE 10 – 11. Eukoenenia indalica sp. nov. 10, ta 3 – bta 4 of left leg I; 11, bta 3 – bta 1 + 2 of left leg I. Scale bar 100 µm." captionText-3="FIGURE 12 – 13. Eukoenenia indalica sp. nov. 12, ti – tc of left leg I; 13, bta of left leg IV. Scale bar 100 µm." captionText-4="FIGURE 14. Schematic representation of chaetotaxy on sternites IV ‒ VII of Eukoenenia indalica sp. nov., E. madeirae Strinati &amp; Condé, 1995 and E. gadorensis Mayoral &amp; Barranco, 2002. For E. madeirae, the most frequent number of setae is represented, but some variability exists." captionText-5="FIGURE 15. Eukoenenia indalica sp. nov., female genitalia, ventral view. Scale bar 50 µm." httpUri-0="https://zenodo.org/record/829115/files/figure.png" httpUri-1="https://zenodo.org/record/829117/files/figure.png" httpUri-2="https://zenodo.org/record/829119/files/figure.png" httpUri-3="https://zenodo.org/record/829121/files/figure.png" httpUri-4="https://zenodo.org/record/829123/files/figure.png" httpUri-5="https://zenodo.org/record/829125/files/figure.png" pageId="1" pageNumber="358">Figs. 1‒15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E6569591CFFBBFF65FD70FEB3FCCF" pageId="1" pageNumber="358" type="materials_examined">
<paragraph id="8BEB36E2591CFFBBFF65FD70FEB3FCCF" blockId="1.[151,1437,686,2034]" pageId="1" pageNumber="358">
<emphasis id="B920EAF0591CFFBBFF65FD70FE77FD07" bold="true" box="[151,385,686,711]" pageId="1" pageNumber="358">Material examined.</emphasis>
Holotype ♀, Cueva de la Corraliza, T.M de Fondón, Sierra de Gádor, Almería (UTM 30SWF X515000 Y4087000 Z1710),
<date id="FFEA1022591CFFBBFE1FFD0DFD57FD2B" box="[493,673,722,747]" pageId="1" pageNumber="358" value="2000-03-18">18 March 2000</date>
, D. Ortega &amp; J. Amate leg. Deposited in Universidad de Almeria (UAL-Pp-006).
</paragraph>
</subSubSection>
<subSubSection id="C34E6569591CFFBEFF35FCC4FE70F848" lastPageId="4" lastPageNumber="361" pageId="1" pageNumber="358" type="description">
<paragraph id="8BEB36E2591CFFBBFF35FCC4FC05FB9C" blockId="1.[151,1437,686,2034]" pageId="1" pageNumber="358">
<emphasis id="B920EAF0591CFFBBFF35FCC4FE27FCF2" bold="true" box="[199,465,794,819]" pageId="1" pageNumber="358">Description of female.</emphasis>
<emphasis id="B920EAF0591CFFBBFE29FCC5FDC9FCF2" box="[475,575,795,818]" italics="true" pageId="1" pageNumber="358">Prosoma</emphasis>
. Frontal organ with two elongated and granulate branches with subparallel borders, 1.6 times longer than wide (
<figureCitation id="136F2A67591CFFBBFDC1FCE0FD83FC97" box="[563,629,830,855]" captionStart="FIGURE 1 – 7" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[272,1307,200,1908]" captionTargetId="figure@2.[260,1320,193,1914]" captionTargetPageId="2" captionText="FIGURE 1 – 7. Eukoeneneia indalica sp nov. 1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2)." httpUri="https://zenodo.org/record/829115/files/figure.png" pageId="1" pageNumber="358">Fig. 1</figureCitation>
). Lateral organ with one pointed, granulate blade, more than 1.45 times longer than wide (32/~22; width of lateral organ distorted due to compression of the specimen during mounting). Propeltidium with 10 + 10 short setae. Metapeltidium with 3 + 3 setae,
<emphasis id="B920EAF0591CFFBBFC45FC59FC31FC63" box="[951,967,903,931]" italics="true" pageId="1" pageNumber="358">t1</emphasis>
(64) and
<emphasis id="B920EAF0591CFFBBFBC1FC59FBB5FC63" box="[1075,1091,903,931]" italics="true" pageId="1" pageNumber="358">t3</emphasis>
(74) shorter than
<emphasis id="B920EAF0591CFFBBFAFEFC59FAEAFC63" box="[1292,1308,903,931]" italics="true" pageId="1" pageNumber="358">t2</emphasis>
(120) (
<figureCitation id="136F2A67591CFFBBFA9CFC58FF5EFC06" captionStart="FIGURE 1 – 7" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[272,1307,200,1908]" captionTargetId="figure@2.[260,1320,193,1914]" captionTargetPageId="2" captionText="FIGURE 1 – 7. Eukoeneneia indalica sp nov. 1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2)." httpUri="https://zenodo.org/record/829115/files/figure.png" pageId="1" pageNumber="358">Fig. 2</figureCitation>
). Hand of chelicera with 6 dorsal and one ventral seta, each finger with 9 teeth. Basal segment of the chelicera with three setae in distal series (
<emphasis id="B920EAF0591CFFBBFE09FC0FFDC3FC2E" box="[507,565,977,1006]" italics="true" pageId="1" pageNumber="358">d1‒d3</emphasis>
):
<emphasis id="B920EAF0591CFFBBFDBEFC0FFD94FC2E" box="[588,610,977,1006]" italics="true" pageId="1" pageNumber="358">d3</emphasis>
&gt;
<emphasis id="B920EAF0591CFFBBFD8DFC0FFD63FC2E" box="[639,661,977,1006]" italics="true" pageId="1" pageNumber="358">d2</emphasis>
&gt;
<emphasis id="B920EAF0591CFFBBFD40FC0FFD31FC2E" box="[690,711,977,1006]" italics="true" pageId="1" pageNumber="358">d1</emphasis>
; setae
<emphasis id="B920EAF0591CFFBBFCE6FC0FFCDCFC2E" box="[788,810,977,1006]" italics="true" pageId="1" pageNumber="358">d1</emphasis>
and
<emphasis id="B920EAF0591CFFBBFCADFC0FFC83FC2E" box="[863,885,977,1006]" italics="true" pageId="1" pageNumber="358">d3</emphasis>
tapering, with small setules along their length;
<emphasis id="B920EAF0591CFFBBFA74FC0FFA6AFC2E" box="[1414,1436,977,1006]" italics="true" pageId="1" pageNumber="358">d3</emphasis>
1.7 times longer than
<emphasis id="B920EAF0591CFFBBFE7CFC26FE52FBD5" box="[398,420,1016,1045]" italics="true" pageId="1" pageNumber="358">d1</emphasis>
;
<emphasis id="B920EAF0591CFFBBFE46FC26FE3CFBD5" box="[436,458,1016,1045]" italics="true" pageId="1" pageNumber="358">d1</emphasis>
much thinner than
<emphasis id="B920EAF0591CFFBBFD55FC26FD4BFBD5" box="[679,701,1016,1045]" italics="true" pageId="1" pageNumber="358">d2</emphasis>
and
<emphasis id="B920EAF0591CFFBBFD07FC26FCFAFBD5" box="[757,780,1016,1045]" italics="true" pageId="1" pageNumber="358">d3</emphasis>
;
<emphasis id="B920EAF0591CFFBBFCE9FC26FCC7FBD5" box="[795,817,1016,1045]" italics="true" pageId="1" pageNumber="358">d2</emphasis>
(middle seta) thickened, with subparallel borders, tiny setules and a crown of spines on the truncated tip. Sternum with 11 deuto-tritosternal setae arranged in two irregular transverse rows: anterior row with 5 setae and posterior row with 6.
</paragraph>
<paragraph id="8BEB36E2591CFFBBFF35FBB9FA7BFB2C" blockId="1.[151,1437,686,2034]" pageId="1" pageNumber="358">
<emphasis id="B920EAF0591CFFBBFF35FBB9FE5DFBBF" box="[199,427,1127,1151]" italics="true" pageId="1" pageNumber="358">Chaetotaxy of coxae</emphasis>
. Pedipalp coxa with 19 setae of different lengths (
<figureCitation id="136F2A67591CFFBBFC2CFBB8FBD4FBBF" box="[990,1058,1126,1151]" captionStart="FIGURE 1 – 7" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[272,1307,200,1908]" captionTargetId="figure@2.[260,1320,193,1914]" captionTargetPageId="2" captionText="FIGURE 1 – 7. Eukoeneneia indalica sp nov. 1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2)." httpUri="https://zenodo.org/record/829115/files/figure.png" pageId="1" pageNumber="358">Fig. 3</figureCitation>
). Coxa I with 15 setae: 10 long, 3 shorter ones and 2 microsetae (
<figureCitation id="136F2A67591CFFBBFE05FB55FDCBFB63" box="[503,573,1163,1188]" captionStart="FIGURE 1 – 7" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[272,1307,200,1908]" captionTargetId="figure@2.[260,1320,193,1914]" captionTargetPageId="2" captionText="FIGURE 1 – 7. Eukoeneneia indalica sp nov. 1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2)." httpUri="https://zenodo.org/record/829115/files/figure.png" pageId="1" pageNumber="358">Fig. 4</figureCitation>
). Coxa II with 13 setae: 4 thickened and 9 normal (
<figureCitation id="136F2A67591CFFBBFB77FB55FB3CFB64" box="[1157,1226,1163,1188]" captionStart="FIGURE 1 – 7" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[272,1307,200,1908]" captionTargetId="figure@2.[260,1320,193,1914]" captionTargetPageId="2" captionText="FIGURE 1 – 7. Eukoeneneia indalica sp nov. 1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2)." httpUri="https://zenodo.org/record/829115/files/figure.png" pageId="1" pageNumber="358">Fig. 5</figureCitation>
). Coxa III with 11 setae: 4 thickened setae, one long macroseta and 6 normal setae (
<figureCitation id="136F2A67591CFFBBFC98FB70FC58FB07" box="[874,942,1198,1223]" captionStart="FIGURE 1 – 7" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[272,1307,200,1908]" captionTargetId="figure@2.[260,1320,193,1914]" captionTargetPageId="2" captionText="FIGURE 1 – 7. Eukoeneneia indalica sp nov. 1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2)." httpUri="https://zenodo.org/record/829115/files/figure.png" pageId="1" pageNumber="358">Fig. 6</figureCitation>
). Coxa IV with 14 setae: 2 thickened and 12 normal (
<figureCitation id="136F2A67591CFFBBFF07FB0DFECCFB2B" box="[245,314,1235,1260]" captionStart="FIGURE 1 – 7" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[272,1307,200,1908]" captionTargetId="figure@2.[260,1320,193,1914]" captionTargetPageId="2" captionText="FIGURE 1 – 7. Eukoeneneia indalica sp nov. 1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2)." httpUri="https://zenodo.org/record/829115/files/figure.png" pageId="1" pageNumber="358">Fig. 7</figureCitation>
). Thickened setae on coxae tapering, with a minute, obliquely projecting spine at the truncated tip.
</paragraph>
<paragraph id="8BEB36E2591CFFBBFF35FB29FCC6FA97" blockId="1.[151,1437,686,2034]" pageId="1" pageNumber="358">
<emphasis id="B920EAF0591CFFBBFF35FB29FF0AFACF" box="[199,252,1271,1295]" italics="true" pageId="1" pageNumber="358">Palp</emphasis>
(
<figureCitation id="136F2A67591CFFBBFEF8FB28FE87FACF" box="[266,369,1270,1295]" captionStart="FIGURE 8 – 9" captionStartId="3.[151,250,1994,2016]" captionTargetBox="[197,1405,202,1966]" captionTargetId="figure@3.[180,1416,193,1978]" captionTargetPageId="3" captionText="FIGURE 8 – 9. Eukoenenia indalica sp. nov. 8, ta 3 – bta 1 of left palp; 9, ti – tc of right palp. Scale bar 100 µm." httpUri="https://zenodo.org/record/829117/files/figure.png" pageId="1" pageNumber="358">Figs 8‒9</figureCitation>
): ta3 with 10 normal setae, 2 bifurcate setae (
<emphasis id="B920EAF0591CFFBBFC99FB29FC88FACF" box="[875,894,1271,1295]" italics="true" pageId="1" pageNumber="358">fs</emphasis>
), 1 curved seta (
<emphasis id="B920EAF0591CFFBBFBC5FB26FBB8FACF" box="[1079,1102,1272,1295]" italics="true" pageId="1" pageNumber="358">cs</emphasis>
), 4 macrosetae and seta
<emphasis id="B920EAF0591CFFBBFA92FB26FA9DFACF" box="[1376,1387,1272,1295]" italics="true" pageId="1" pageNumber="358">r</emphasis>
; ta2 with 3 normal setae and 3 macrosetae; ta1 with 2 normal setae; bta2 with 4 normal setae and 2 macrosetae; bta1 with 3 setae; ti with 9 setae; fe with 8 setae; tc with 7 setae.
</paragraph>
<paragraph id="8BEB36E2591CFFBBFF35FABAFC43F9F0" blockId="1.[151,1437,686,2034]" pageId="1" pageNumber="358">
<emphasis id="B920EAF0591CFFBBFF35FABAFEF7FABB" box="[199,257,1380,1403]" italics="true" pageId="1" pageNumber="358">Leg I</emphasis>
(
<figureCitation id="136F2A67591CFFBBFEFDFABDFE64FABB" box="[271,402,1379,1404]" captionStart-0="FIGURE 10 – 11" captionStart-1="FIGURE 12 – 13" captionStartId-0="4.[151,250,1585,1607]" captionStartId-1="5.[151,250,1831,1853]" captionTargetBox-0="[180,1415,204,1561]" captionTargetBox-1="[171,1427,206,1797]" captionTargetId-0="figure@4.[166,1430,193,1575]" captionTargetId-1="figure@5.[151,1436,195,1809]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 10 – 11. Eukoenenia indalica sp. nov. 10, ta 3 – bta 4 of left leg I; 11, bta 3 – bta 1 + 2 of left leg I. Scale bar 100 µm." captionText-1="FIGURE 12 – 13. Eukoenenia indalica sp. nov. 12, ti – tc of left leg I; 13, bta of left leg IV. Scale bar 100 µm." httpUri-0="https://zenodo.org/record/829119/files/figure.png" httpUri-1="https://zenodo.org/record/829121/files/figure.png" pageId="1" pageNumber="358">Figs 10‒12</figureCitation>
): ta3 with 15 normal setae, 3 macrosetae, 2 bifurcate setae (
<emphasis id="B920EAF0591CFFBBFBD8FABDFBCBFABB" box="[1066,1085,1379,1403]" italics="true" pageId="1" pageNumber="358">fs</emphasis>
) with equal branches and
<emphasis id="B920EAF0591CFFBBFA92FABAFA9DFABB" box="[1376,1387,1380,1403]" italics="true" pageId="1" pageNumber="358">r</emphasis>
; ta2 with 5 normal setae, 1 trichobothrium (
<emphasis id="B920EAF0591CFFBBFDAAFA59FD98FA5F" box="[600,622,1415,1439]" italics="true" pageId="1" pageNumber="358">tb</emphasis>
) and 1 bifurcate seta (
<emphasis id="B920EAF0591CFFBBFC83FA59FC72FA5F" box="[881,900,1415,1439]" italics="true" pageId="1" pageNumber="358">fs</emphasis>
) with equal branches; ta1 with 5 normal setae, one considerably smaller than the others; bta4 with 5 normal setae, 1
<emphasis id="B920EAF0591CFFBBFC68FA75FC46FA03" box="[922,944,1451,1475]" italics="true" pageId="1" pageNumber="358">tb</emphasis>
and 1
<emphasis id="B920EAF0591CFFBBFC08FA75FBFBFA03" box="[1018,1037,1451,1475]" italics="true" pageId="1" pageNumber="358">fs</emphasis>
with unequal branches; bta3 with
<emphasis id="B920EAF0591CFFBBFA79FA72FA63FA03" box="[1419,1429,1452,1475]" italics="true" pageId="1" pageNumber="358">r</emphasis>
,
<emphasis id="B920EAF0591CFFBBFF65FA0EFF4EFA27" box="[151,184,1488,1511]" italics="true" pageId="1" pageNumber="358">grt</emphasis>
and 1 microseta; bta2 with 4 normal setae, 2
<emphasis id="B920EAF0591CFFBBFD41FA11FD3FFA27" box="[691,713,1487,1511]" italics="true" pageId="1" pageNumber="358">tb</emphasis>
, and 1
<emphasis id="B920EAF0591CFFBBFCE8FA11FCDBFA27" box="[794,813,1487,1511]" italics="true" pageId="1" pageNumber="358">fs</emphasis>
with equal branches; bta1 with 2 normal setae, 2
<emphasis id="B920EAF0591CFFBBFAA5FA11FA9BFA27" box="[1367,1389,1487,1511]" italics="true" pageId="1" pageNumber="358">tb</emphasis>
and 1
<emphasis id="B920EAF0591CFFBBFF5EFA2DFF49F9CB" box="[172,191,1523,1547]" italics="true" pageId="1" pageNumber="358">fs</emphasis>
with unequal branches; ti with 9 normal setae; pa with 9 normal setae and 1
<emphasis id="B920EAF0591CFFBBFBD2FA2DFBC0F9CB" box="[1056,1078,1523,1547]" italics="true" pageId="1" pageNumber="358">tb</emphasis>
; fe with 9 normal setae; tc with 11 normal setae, two of which are considerably smaller than the others.
</paragraph>
<paragraph id="8BEB36E2591CFFBBFF35F9E2FF17F900" blockId="1.[151,1437,686,2034]" pageId="1" pageNumber="358">
<emphasis id="B920EAF0591CFFBBFF35F9E2FE41F993" box="[199,439,1595,1619]" italics="true" pageId="1" pageNumber="358">Basitarsus of leg IV</emphasis>
(IVbta) (
<figureCitation id="136F2A67591CFFBBFDDFF9E5FD72F993" box="[557,644,1595,1620]" captionStart="FIGURE 12 – 13" captionStartId="5.[151,250,1831,1853]" captionTargetBox="[171,1427,206,1797]" captionTargetId="figure@5.[151,1436,195,1809]" captionTargetPageId="5" captionText="FIGURE 12 – 13. Eukoenenia indalica sp. nov. 12, ti – tc of left leg I; 13, bta of left leg IV. Scale bar 100 µm." httpUri="https://zenodo.org/record/829121/files/figure.png" pageId="1" pageNumber="358">Fig. 13</figureCitation>
) with seven setae: 2
<emphasis id="B920EAF0591CFFBBFC62F9E5FC43F993" box="[912,949,1595,1619]" italics="true" pageId="1" pageNumber="358">esd</emphasis>
inserted most distally;
<emphasis id="B920EAF0591CFFBBFB26F9E5FB0EF993" box="[1236,1272,1595,1619]" italics="true" pageId="1" pageNumber="358">gla</emphasis>
, 2
<emphasis id="B920EAF0591CFFBBFADAF9E2FABBF993" box="[1320,1357,1596,1619]" italics="true" pageId="1" pageNumber="358">esp</emphasis>
and
<emphasis id="B920EAF0591CFFBBFA63F9E2FA6AF993" box="[1425,1436,1596,1619]" italics="true" pageId="1" pageNumber="358">r</emphasis>
approximately in the middle of segment;
<emphasis id="B920EAF0591CFFBBFD9EF981FD67F9B7" box="[620,657,1631,1655]" italics="true" pageId="1" pageNumber="358">esd</emphasis>
and
<emphasis id="B920EAF0591CFFBBFD38F9BEFD19F9B7" box="[714,751,1632,1655]" italics="true" pageId="1" pageNumber="358">esp</emphasis>
with a small spine at its base (characteristic of macrosetae);
<emphasis id="B920EAF0591CFFBBFF65F95AFF4EF95B" box="[151,184,1668,1691]" italics="true" pageId="1" pageNumber="358">grt</emphasis>
proximad of the other setae, with a small and sharp spine at its base. Lengths and ratios given in measurements below.
</paragraph>
<paragraph id="8BEB36E2591CFFBBFF35F915FEFDF865" blockId="1.[151,1437,686,2034]" pageId="1" pageNumber="358">
<emphasis id="B920EAF0591CFFBBFF35F915FEADF923" box="[199,347,1739,1763]" italics="true" pageId="1" pageNumber="358">Opisthosoma</emphasis>
. Tergites II–VII each with 3 + 3 dorsal setae:
<emphasis id="B920EAF0591CFFBBFC84F912FC70F928" box="[886,902,1740,1768]" italics="true" pageId="1" pageNumber="358">t1</emphasis>
,
<emphasis id="B920EAF0591CFFBBFC64F912FC53F928" box="[918,933,1740,1768]" italics="true" pageId="1" pageNumber="358">t3</emphasis>
and one slender lateral seta
<emphasis id="B920EAF0591CFFBBFB03F912FB0AF923" box="[1265,1276,1740,1763]" italics="true" pageId="1" pageNumber="358">s</emphasis>
on each side. Sternite III with 2 + 2 setae. Sternites IV and V with 3 + 3 setae, inner ones (
<emphasis id="B920EAF0591CFFBBFC06F92CFBFFF8CE" box="[1012,1033,1778,1806]" italics="true" pageId="1" pageNumber="358">a1</emphasis>
,
<emphasis id="B920EAF0591CFFBBFBE5F92CFBDBF8CE" box="[1047,1069,1778,1806]" italics="true" pageId="1" pageNumber="358">a2</emphasis>
) 75 long and
<geoCoordinate id="EE605025591CFFBBFB3BF92CFB08F8CA" box="[1225,1278,1777,1806]" direction="south" orientation="latitude" pageId="1" pageNumber="358" precision="925" value="-1.75">
<emphasis id="B920EAF0591CFFBBFB3BF92CFB2AF8CE" box="[1225,1244,1778,1806]" italics="true" pageId="1" pageNumber="358">s1</emphasis>
45
</geoCoordinate>
long. Sternite VI with a wart-like structure in medial plane of segment bearing 4 thickened setae, medial 2 setae (
<emphasis id="B920EAF0591CFFBBFB03F8C7FAF1F8F5" box="[1265,1287,1817,1845]" italics="true" pageId="1" pageNumber="358">a1</emphasis>
) 50 long and lateral ones (
<emphasis id="B920EAF0591CFFBBFED4F89EFECAF89C" box="[294,316,1856,1884]" italics="true" pageId="1" pageNumber="358">a2</emphasis>
) 63 long. The setae
<emphasis id="B920EAF0591CFFBBFDEDF89EFDC4F89C" box="[543,562,1856,1884]" italics="true" pageId="1" pageNumber="358">s1</emphasis>
on each side of wart-like structure are 58 long (
<figureCitation id="136F2A67591CFFBBFBB4F8E1FB6FF898" box="[1094,1177,1855,1880]" captionStart="FIGURE 14" captionStartId="6.[151,250,1806,1828]" captionTargetBox="[159,1418,1214,1778]" captionTargetId="figure@6.[153,1433,1201,1784]" captionTargetPageId="6" captionText="FIGURE 14. Schematic representation of chaetotaxy on sternites IV ‒ VII of Eukoenenia indalica sp. nov., E. madeirae Strinati &amp; Condé, 1995 and E. gadorensis Mayoral &amp; Barranco, 2002. For E. madeirae, the most frequent number of setae is represented, but some variability exists." httpUri="https://zenodo.org/record/829123/files/figure.png" pageId="1" pageNumber="358">Fig. 14</figureCitation>
). Sternite VII with 3+3 setae,
<emphasis id="B920EAF0591CFFBBFF10F8B9FF0EF843" box="[226,248,1895,1923]" italics="true" pageId="1" pageNumber="358">a1</emphasis>
80,
<emphasis id="B920EAF0591CFFBBFEC1F8B9FEBFF843" box="[307,329,1895,1923]" italics="true" pageId="1" pageNumber="358">a2</emphasis>
60 and
<geoCoordinate id="EE605025591CFFBBFE40F8B9FE1BF8BF" box="[434,493,1895,1923]" direction="south" orientation="latitude" pageId="1" pageNumber="358" precision="925" value="-1.6666667">
<emphasis id="B920EAF0591CFFBBFE40F8B9FE33F843" box="[434,453,1895,1923]" italics="true" pageId="1" pageNumber="358">s1</emphasis>
40
</geoCoordinate>
long. Segments VIII–XI with 10, 6, 6 and 8 setae, respectively. Flagellum not preserved.
</paragraph>
<paragraph id="8BEB36E2591CFFBEFF35F86FFDCDF915" blockId="1.[151,1437,686,2034]" lastBlockId="4.[151,1437,1646,2037]" lastPageId="4" lastPageNumber="361" pageId="1" pageNumber="358">
<emphasis id="B920EAF0591CFFBBFF35F86FFE9CF809" box="[199,362,1969,1993]" italics="true" pageId="1" pageNumber="358">Genital region</emphasis>
. Two lobes with a total of 22 setae. First lobe with 9 + 9 setae in 5 transverse rows: 1 + 1, 1 + 1, 1 + 1, 1 + 1, and 4 + 4 on posterior margin of lobe; of these,
<emphasis id="B920EAF0591CFFBBFCC7F808FCBDF832" box="[821,843,2006,2034]" italics="true" pageId="1" pageNumber="358">a3</emphasis>
and
<emphasis id="B920EAF0591CFFBBFC72F808FC60F832" box="[896,918,2006,2034]" italics="true" pageId="1" pageNumber="358">a4</emphasis>
(34‒38) are approximately twice the length of
<emphasis id="B920EAF05919FFBEFF65F9B1FF5BF94B" box="[151,173,1647,1675]" italics="true" pageId="4" pageNumber="361">a1</emphasis>
and
<emphasis id="B920EAF05919FFBEFF18F9B1FEF6F94B" box="[234,256,1647,1675]" italics="true" pageId="4" pageNumber="361">a2</emphasis>
(18). Posterior margin of first genital lobe without serrations and with a rounded median indentation between
<emphasis id="B920EAF05919FFBEFF0EF948FEE4F972" box="[252,274,1686,1714]" italics="true" pageId="4" pageNumber="361">a1</emphasis>
setae. The two halves of the second lobe each with 2 setae (
<emphasis id="B920EAF05919FFBEFC36F948FC26F96D" box="[964,976,1686,1709]" italics="true" pageId="4" pageNumber="361">y</emphasis>
and
<emphasis id="B920EAF05919FFBEFBFBF948FBE2F96D" box="[1033,1044,1686,1709]" italics="true" pageId="4" pageNumber="361">z</emphasis>
) of about the same length (33 and 30‒32) inserted near the tip (
<figureCitation id="136F2A675919FFBEFE28F962FDDBF915" box="[474,557,1724,1749]" captionStart="FIGURE 15" captionStartId="7.[151,250,1962,1984]" captionTargetBox="[167,1430,851,1910]" captionTargetId="figure@7.[151,1436,844,1920]" captionTargetPageId="7" captionText="FIGURE 15. Eukoenenia indalica sp. nov., female genitalia, ventral view. Scale bar 50 µm." httpUri="https://zenodo.org/record/829125/files/figure.png" pageId="4" pageNumber="361">Fig. 15</figureCitation>
).
</paragraph>
<caption id="DF2B666A591FFFB8FF65F84FFDDEF824" httpUri="https://zenodo.org/record/829115/files/figure.png" pageId="2" pageNumber="359" targetBox="[272,1307,200,1908]" targetPageId="2">
<paragraph id="8BEB36E2591FFFB8FF65F84FFDDEF824" blockId="2.[151,1436,1937,2020]" pageId="2" pageNumber="359">
<emphasis id="B920EAF0591FFFB8FF65F84FFEDAF866" bold="true" box="[151,300,1937,1959]" pageId="2" pageNumber="359">FIGURE 1–7.</emphasis>
<emphasis id="B920EAF0591FFFB8FEC6F84FFDE6F866" box="[308,528,1937,1958]" italics="true" pageId="2" pageNumber="359">
Eukoeneneia
<taxonomicName id="4C544D61591FFFB8FE4FF84FFDE6F866" box="[445,528,1937,1958]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="indalica">indalica</taxonomicName>
</emphasis>
<emphasis id="B920EAF0591FFFB8FDE5F84FFD94F866" bold="true" box="[535,610,1937,1958]" pageId="2" pageNumber="359">sp nov.</emphasis>
1, frontal organ, dorsal view; 2, metapeltidial setae, dorsal view; 3, left pedipalp coxa; 4, left coxa I; 5, right coxa II; 6, left coxa III; 7, left coxa IV. Thickened setae of coxae drawn in solid black. Scale bars 100 µm (Fig. 1 to same scale as Fig. 2).
</paragraph>
</caption>
<caption id="DF2B666A591EFFB9FF65F814FB15F820" box="[151,1251,1994,2016]" httpUri="https://zenodo.org/record/829117/files/figure.png" pageId="3" pageNumber="360" targetBox="[197,1405,202,1966]" targetPageId="3">
<paragraph id="8BEB36E2591EFFB9FF65F814FB15F820" blockId="3.[151,1251,1994,2016]" box="[151,1251,1994,2016]" pageId="3" pageNumber="360">
<emphasis id="B920EAF0591EFFB9FF65F814FEDAF81F" bold="true" box="[151,300,1994,2016]" pageId="3" pageNumber="360">FIGURE 8–9.</emphasis>
<taxonomicName id="4C544D61591EFFB9FEC0F814FDF5F81F" box="[306,515,1994,2015]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="indalica" status="sp. nov.">
<emphasis id="B920EAF0591EFFB9FEC0F814FDF5F81F" box="[306,515,1994,2015]" italics="true" pageId="3" pageNumber="360">Eukoenenia indalica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213578B591EFFB9FDFBF814FDACF820" box="[521,602,1994,2016]" pageId="3" pageNumber="360" rank="species">
<emphasis id="B920EAF0591EFFB9FDFBF814FDD7F81F" bold="true" box="[521,545,1994,2015]" pageId="3" pageNumber="360">sp</emphasis>
.
<emphasis id="B920EAF0591EFFB9FDDFF814FDA5F81F" bold="true" box="[557,595,1994,2015]" pageId="3" pageNumber="360">nov</emphasis>
.
</taxonomicNameLabel>
8, ta3
<emphasis id="B920EAF0591EFFB9FD6AF814FD53F81F" bold="true" box="[664,677,1994,2015]" pageId="3" pageNumber="360">–</emphasis>
bta1 of left palp; 9, ti
<emphasis id="B920EAF0591EFFB9FC89F814FC71F81F" bold="true" box="[891,903,1994,2015]" pageId="3" pageNumber="360">–</emphasis>
tc of right palp. Scale bar 100 µm.
</paragraph>
</caption>
<caption id="DF2B666A5919FFBEFF65F9EFFAADF987" box="[151,1371,1585,1607]" httpUri="https://zenodo.org/record/829119/files/figure.png" pageId="4" pageNumber="361" targetBox="[180,1415,204,1561]" targetPageId="4">
<paragraph id="8BEB36E25919FFBEFF65F9EFFAADF987" blockId="4.[151,1371,1585,1607]" box="[151,1371,1585,1607]" pageId="4" pageNumber="361">
<emphasis id="B920EAF05919FFBEFF65F9EFFEB2F986" bold="true" box="[151,324,1585,1607]" pageId="4" pageNumber="361">FIGURE 10–11.</emphasis>
<taxonomicName id="4C544D615919FFBEFEB8F9EFFDEDF986" box="[330,539,1585,1606]" class="Arachnida" family="Eukoeneniidae" genus="Eukoenenia" kingdom="Animalia" order="Palpigradi" pageId="4" pageNumber="361" phylum="Arthropoda" rank="species" species="indalica" status="sp. nov.">
<emphasis id="B920EAF05919FFBEFEB8F9EFFDEDF986" box="[330,539,1585,1606]" italics="true" pageId="4" pageNumber="361">Eukoenenia indalica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213578B5919FFBEFDD3F9EFFD84F987" box="[545,626,1585,1607]" pageId="4" pageNumber="361" rank="species">
<emphasis id="B920EAF05919FFBEFDD3F9EFFDCFF986" bold="true" box="[545,569,1585,1606]" pageId="4" pageNumber="361">sp</emphasis>
.
<emphasis id="B920EAF05919FFBEFDB4F9EFFD9AF986" bold="true" box="[582,620,1585,1606]" pageId="4" pageNumber="361">nov</emphasis>
.
</taxonomicNameLabel>
10, ta3
<emphasis id="B920EAF05919FFBEFD4FF9EFFD3FF986" bold="true" box="[701,713,1585,1606]" pageId="4" pageNumber="361">–</emphasis>
bta4 of left leg I; 11, bta3
<emphasis id="B920EAF05919FFBEFC39F9EFFC2EF986" bold="true" box="[971,984,1585,1606]" pageId="4" pageNumber="361">–</emphasis>
bta1+2 of left leg I. Scale bar 100 µm.
</paragraph>
</caption>
<paragraph id="8BEB36E25919FFBEFF35F93FFB10F8A5" blockId="4.[151,1437,1646,2037]" pageId="4" pageNumber="361">
<emphasis id="B920EAF05919FFBEFF35F93FFE9FF938" box="[199,361,1761,1784]" italics="true" pageId="4" pageNumber="361">Measurements</emphasis>
.
<emphasis id="B920EAF05919FFBEFE8AF901FE40F938" bold="true" box="[376,438,1759,1784]" pageId="4" pageNumber="361">Body</emphasis>
: B 460,
<emphasis id="B920EAF05919FFBEFDEBF901FDB9F938" bold="true" box="[537,591,1759,1784]" pageId="4" pageNumber="361">Palp</emphasis>
: Pcx 140, Ptc 203, Pfe 238, Pti 213, Pbta1 80, Pbta2 83, Pta1 45, Pta2 50, Pta3 70.
<emphasis id="B920EAF05919FFBEFEF3F8DAFEB2F8DD" bold="true" box="[257,324,1796,1821]" pageId="4" pageNumber="361">Leg I</emphasis>
: Icx 192, Itc 278, Itfe 238, Ipa 238, Iti 213, Ibta1+2 180, Ibta3 88, Ibta4 70, Ita1 43, Ita2 53, Ita3 178.
<emphasis id="B920EAF05919FFBEFEF0F8F9FEA2F880" bold="true" box="[258,340,1831,1856]" pageId="4" pageNumber="361">Leg IV</emphasis>
: IVcx 150, IVtc 212, IVfe 330, IVpa 232, IVti 313, IVbta 270, IVta1 80, IVta2 85, a 20, er 125, grt 153, gla 120, r 73, esd 78‒80, esp 93‒95, t/r 3.69, t/er 2.16, gla/grt 0.78, B/bta 1.70, bta/ti 0.86.
</paragraph>
<paragraph id="8BEB36E25919FFBEFF35F8B1FE70F848" blockId="4.[151,1437,1646,2037]" box="[199,390,1903,1928]" pageId="4" pageNumber="361">
<emphasis id="B920EAF05919FFBEFF35F8B1FEFFF848" bold="true" box="[199,265,1903,1928]" italics="true" pageId="4" pageNumber="361">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34E65695919FFBEFF35F84AFD7AF835" pageId="4" pageNumber="361" type="etymology">
<paragraph id="8BEB36E25919FFBEFF35F84AFD7AF835" blockId="4.[151,1437,1646,2037]" pageId="4" pageNumber="361">
<emphasis id="B920EAF05919FFBEFF35F84AFEB8F86B" bold="true" box="[199,334,1940,1965]" pageId="4" pageNumber="361">Etymology.</emphasis>
This species is named after the Indalo, a Neolithic representation of an anthropomorphic figure holding a rainbow/arch, depicted at the entrance of a few caves in the northern part of the province of Almeria (
<collectingCountry id="F34376725919FFBEFF6DF802FF12F835" box="[159,228,2012,2037]" name="Spain" pageId="4" pageNumber="361">Spain</collectingCountry>
), in which the
<typeStatus id="54EF88405919FFBEFE7BF802FE1DF835" box="[393,491,2012,2037]" pageId="4" pageNumber="361" type="holotype">holotype</typeStatus>
was captured.
</paragraph>
</subSubSection>
</treatment>
</document>
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