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<document id="E1F9784535DFA321018A1D97BBDD5C0D" ID-DOI="10.11646/zootaxa.5367.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10145106" ID-ZooBank="5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700167363318" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="03FD878BFFE8FFCDFCBEFDE358D23E56" docLanguage="en" docName="zootaxa.5367.1.1.pdf" docOrigin="Zootaxa 5367 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ochthebius (Asiobates) cascadeus Perkins 2023, new species" docType="treatment" docVersion="2" lastPageNumber="61" masterDocId="FFC4FFF3FFD0FFF1FC29FFA5596F3C37" masterDocTitle="New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)" masterLastPageNumber="86" masterPageNumber="1" pageNumber="57" updateTime="1700168056660" updateUser="ExternalLinkService">
<mods:mods id="64EB31674E9B0B9B29813F07506D4B27" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9711CA3B42F255E1585EF152A7E2F2C4">New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)</mods:title>
</mods:titleInfo>
<mods:name id="024250FDDE575A3742F24DB3C61A75DA" type="personal">
<mods:role id="5050423EAECF69362BB5C4B4724CFDCA">
<mods:roleTerm id="3E9B7F7E3033C864F4929F28C4FA54A0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0075816D3D13A8826E7686DD9C401264">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="80AF681A2173328B3A91CEFFC53946BE">text</mods:typeOfResource>
<mods:relatedItem id="F719DC999BD1D47C1891B38158EA450B" type="host">
<mods:titleInfo id="DC88A4ABCA02F10F56A89FDC51667EB2">
<mods:title id="1E8A824C5FB890D012939A96801846CA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D1F79A41ACBF5636FCF368487A6BF145">
<mods:date id="89DD25FAE94FC1B45AA7D626C645FF45">2023</mods:date>
<mods:detail id="E69BA665F6F5C5DB1201E1368BFBF14C" type="pubDate">
<mods:number id="E71E269044F0C703ABFA1CBC57B7E768">2023-11-07</mods:number>
</mods:detail>
<mods:detail id="04845E2BCF16651F53CBCF9100896E92" type="volume">
<mods:number id="26AEAD513E83D9B40984BD5A9F89B9D4">5367</mods:number>
</mods:detail>
<mods:detail id="0E7DFC200BBC46FF6E9075E49DC20CFF" type="issue">
<mods:number id="E8BB820C220210DACD38520E43D44DF7">1</mods:number>
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<mods:start id="81E0919DE1C1D30D1C8F760F9FC3B2E1">1</mods:start>
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<mods:location id="A7EC63CE9B77B947E3858C7D44BEDEC2">
<mods:url id="4FB4ADA9FFBDF1D96C3A772C52EEE339">https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225</mods:url>
</mods:location>
<mods:classification id="17C7FD81B50D283AB6EAE4A84B918828">journal article</mods:classification>
<mods:identifier id="6A81E427FBA8C4600AA9793D61885862" type="DOI">10.11646/zootaxa.5367.1.1</mods:identifier>
<mods:identifier id="C580EC360A3125163F26C8EE7E92FDEB" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="814A723BB52BA2C22DB0416E043E8DA7" type="Zenodo-Dep">10145106</mods:identifier>
<mods:identifier id="677C77E1E3AB2AC30C554536A5D812C8" type="ZooBank">5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723</mods:identifier>
</mods:mods>
<treatment id="03FD878BFFE8FFCDFCBEFDE358D23E56" LSID="urn:lsid:plazi:treatment:03FD878BFFE8FFCDFCBEFDE358D23E56" httpUri="http://treatment.plazi.org/id/03FD878BFFE8FFCDFCBEFDE358D23E56" lastPageId="60" lastPageNumber="61" pageId="56" pageNumber="57">
<subSubSection id="C34E6516FFE8FFC9FCBEFDE35BBF3E57" box="[151,720,582,609]" pageId="56" pageNumber="57" type="nomenclature">
<paragraph id="8BEB369DFFE8FFC9FCBEFDE35BBF3E57" blockId="56.[151,778,582,645]" box="[151,720,582,609]" pageId="56" pageNumber="57">
<heading id="D0A381F1FFE8FFC9FCBEFDE35BBF3E57" bold="true" box="[151,720,582,609]" fontSize="11" level="2" pageId="56" pageNumber="57" reason="4">
<emphasis id="B920EA8FFFE8FFC9FCBEFDE35BBF3E57" bold="true" box="[151,720,582,609]" pageId="56" pageNumber="57">
<taxonomicName id="4C544D1EFFE8FFC9FCBEFDE35B5D3E56" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[151,562,582,609]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="cascadeus" status="sp. nov." subGenus="Asiobates">
<emphasis id="B920EA8FFFE8FFC9FCBEFDE35B5D3E56" bold="true" box="[151,562,582,609]" italics="true" pageId="56" pageNumber="57">Ochthebius (Asiobates) cascadeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21357F4FFE8FFC9FE68FDE25BBF3E57" box="[577,720,582,609]" pageId="56" pageNumber="57" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFE8FFC9FCBEFDCE5A653EB2" box="[151,778,619,645]" pageId="56" pageNumber="57" type="description">
<paragraph id="8BEB369DFFE8FFC9FCBEFDCE5A653EB2" blockId="56.[151,778,582,645]" box="[151,778,619,645]" pageId="56" pageNumber="57">
<figureCitation id="136F2A18FFE8FFC9FCBEFDCE598C3EB2" box="[151,227,619,645]" captionStart="FIGURE 4" captionStartId="8.[151,250,1867,1891]" captionTargetBox="[214,1374,181,1844]" captionTargetId="figure-35@8.[214,1374,181,1844]" captionTargetPageId="8" captionText="FIGURE 4. Dorsal habitus of species of Ochthebius." figureDoi="http://doi.org/10.5281/zenodo.10145114" httpUri="https://zenodo.org/record/10145114/files/figure.png" pageId="56" pageNumber="57">Figs. 4</figureCitation>
(habitus comparison), 33 (habitus and aedeagus)
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFE8FFC9FCBEFD165A993F6A" pageId="56" pageNumber="57" type="materials_examined">
<paragraph id="8BEB369DFFE8FFC9FCBEFD165A993F6A" blockId="56.[151,1437,690,1941]" pageId="56" pageNumber="57">
<emphasis id="B920EA8FFFE8FFC9FCBEFD1658283EFB" bold="true" box="[151,327,690,717]" pageId="56" pageNumber="57">Type Material.</emphasis>
<materialsCitation id="3B3C3CC0FFE8FFC9FD64FD1659933F22" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" country="United States of America" county="Pierce Co." location="West Side Rd." municipality="Mt. Rainier" pageId="56" pageNumber="57" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<typeStatus id="54EF883FFFE8FFC9FD64FD1658DA3EFA" box="[333,437,691,717]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B920EA8FFFE8FFC9FE3BFD175B0F3EFB" bold="true" box="[530,608,690,716]" pageId="56" pageNumber="57">
<collectingCountry id="F343760DFFE8FFC9FE3BFD175B0F3EFB" box="[530,608,690,716]" name="United States of America" pageId="56" pageNumber="57">U.S.A.</collectingCountry>
</emphasis>
:
<collectingRegion id="4990F87FFFE8FFC9FE44FD165B983EFA" box="[621,759,691,717]" country="United States of America" name="Washington" pageId="56" pageNumber="57">Washington</collectingRegion>
: “WASH:
<collectingCounty id="628A4E11FFE8FFC9FF40FD165AB03EFA" box="[873,991,691,717]" pageId="56" pageNumber="57">Pierce Co.</collectingCounty>
,
<collectingMunicipality id="6B8FACE7FFE8FFC9FFC3FD165D053EFA" box="[1002,1130,691,717]" pageId="56" pageNumber="57">Mt. Rainier</collectingMunicipality>
N.P.,
<location id="8E8B6046FFE8FFC9F885FD165C243EFA" LSID="urn:lsid:plazi:treatment:03FD878BFFE8FFCDFCBEFDE358D23E56:8E8B6046FFE8FFC9F885FD165C243EFA" box="[1196,1355,691,717]" country="United States of America" county="Pierce Co." municipality="Mt. Rainier" name="West Side Rd." pageId="56" pageNumber="57" stateProvince="Washington">West Side Rd.</location>
,
<quantity id="4CAC9B78FFE8FFC9F97FFD165CF23EFA" box="[1366,1437,691,717]" metricMagnitude="3" metricUnit="m" metricValue="5.3108352" pageId="56" pageNumber="57" unit="mi" value="3.3">3.3 mi</quantity>
N jct Wash 706, 2800’ VII.19.1975 // muddy and gravelly banks, lg. stream //
<collectorName id="26A1534BFFE8FFC9F82CFD725DEE3EC6" box="[1029,1153,727,753]" pageId="56" pageNumber="57">A. Newton</collectorName>
&amp;
<collectorName id="26A1534BFFE8FFC9F88CFD725C743EC6" box="[1189,1307,727,753]" pageId="56" pageNumber="57">M. Thayer</collectorName>
collectors” (
<collectionCode id="ED45AE58FFE8FFC9FCB6FD5E599D3F22" box="[159,242,763,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="56" pageNumber="57" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3C3CC0FFE8FFC9FD23FD5E5A3A3F22" box="[266,853,762,789]" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" country="United States of America" county="Pierce Co." location="West Side Rd." municipality="Mt. Rainier" pageId="56" pageNumber="57" specimenCount="1" stateProvince="Washington" typeStatus="paratype">
<emphasis id="B920EA8FFFE8FFC9FD23FD5E58D73F23" bold="true" box="[266,440,762,789]" pageId="56" pageNumber="57">
<typeStatus id="54EF883FFFE8FFC9FD23FD5E58ED3F22" box="[266,386,763,789]" pageId="56" pageNumber="57" type="paratype">Paratypes</typeStatus>
(15)
</emphasis>
: Same data as holotype (11
<collectionCode id="ED45AE58FFE8FFC9FED0FD5E5A253F22" box="[761,842,763,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="56" pageNumber="57" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CC0FFE8FFC9FF4AFD5F5DE23F0E" collectingDate="1975-07-11" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" country="United States of America" county="Clackamas Co." location="Ore" pageId="56" pageNumber="57" specimenCount="1" stateProvince="Oregon" typeStatus="paratype">
<emphasis id="B920EA8FFFE8FFC9FF4AFD5F5AD23F23" bold="true" box="[867,957,762,788]" pageId="56" pageNumber="57">
<collectingRegion id="4990F87FFFE8FFC9FF4AFD5F5AD23F23" box="[867,957,762,788]" country="United States of America" name="Oregon" pageId="56" pageNumber="57">Oregon</collectingRegion>
</emphasis>
,
<collectingCounty id="628A4E11FFE8FFC9FFE2FD5E5D173F22" box="[971,1144,763,789]" pageId="56" pageNumber="57">Clackamas Co.</collectingCounty>
,
<quantity id="4CAC9B78FFE8FFC9F8ADFD5F5DBB3F22" box="[1156,1236,762,789]" metricMagnitude="3" metricUnit="m" metricValue="2.414016" pageId="56" pageNumber="57" unit="mi" value="1.5">1.5 mi.</quantity>
S. jct. US 26 and
<location id="8E8B6046FFE8FFC9FCBEFCBA59AE3F0E" LSID="urn:lsid:plazi:treatment:03FD878BFFE8FFCDFCBEFDE358D23E56:8E8B6046FFE8FFC9FCBEFCBA59AE3F0E" box="[151,193,799,825]" country="United States of America" county="Clackamas Co." name="Ore" pageId="56" pageNumber="57" stateProvince="Oregon">Ore</location>
35, flood debris, forest stream,
<date id="FFEA105DFFE8FFC9FE19FCBB5BDE3F0E" box="[560,689,798,825]" pageId="56" pageNumber="57" value="1975-07-11">
<collectingDate id="EFAEE9B5FFE8FFC9FE19FCBB5BDE3F0E" box="[560,689,798,825]" pageId="56" pageNumber="57" value="1975-07-11">11.vii.1975</collectingDate>
</date>
, leg.
<collectorName id="26A1534BFFE8FFC9FEC7FCBA5A043F0E" box="[750,875,799,825]" pageId="56" pageNumber="57">A. Newton</collectorName>
&amp;
<collectorName id="26A1534BFFE8FFC9FFB8FCBA5D673F0E" box="[913,1032,799,825]" pageId="56" pageNumber="57">M. Thayer</collectorName>
(3
<collectionCode id="ED45AE58FFE8FFC9F818FCBA5DED3F0E" box="[1073,1154,799,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="56" pageNumber="57" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CC0FFE8FFC9F8B2FCBB5A9D3F6A" collectingDate="1968-05-15" collectionCode="MCZ" collectorName="Olympic Nat. Pk. &amp; Campbell &amp; Smetana" country="United States of America" location="Washington" pageId="56" pageNumber="57" specimenCount="1" stateProvince="Washington" typeStatus="paratype">
<emphasis id="B920EA8FFFE8FFC9F8B2FCBB5C453F0F" bold="true" box="[1179,1322,798,824]" pageId="56" pageNumber="57">
<collectingRegion id="4990F87FFFE8FFC9F8B2FCBB5C453F0F" box="[1179,1322,798,824]" country="United States of America" name="Washington" pageId="56" pageNumber="57">Washington</collectingRegion>
</emphasis>
,
<collectorName id="26A1534BFFE8FFC9F913FCBA599A3F6A" pageId="56" pageNumber="57">Olympic Nat. Pk.</collectorName>
, Queets Ranger Sta.,
<date id="FFEA105DFFE8FFC9FDCFFCE75B393F6A" box="[486,598,834,861]" pageId="56" pageNumber="57" value="1968-05-15">
<collectingDate id="EFAEE9B5FFE8FFC9FDCFFCE75B393F6A" box="[486,598,834,861]" pageId="56" pageNumber="57" value="1968-05-15">15.v.1968</collectingDate>
</date>
, leg.
<collectorName id="26A1534BFFE8FFC9FEBBFCE65B903F6A" box="[658,767,835,861]" pageId="56" pageNumber="57">Campbell</collectorName>
&amp;
<collectorName id="26A1534BFFE8FFC9FF0AFCE65AEC3F6A" box="[803,899,835,861]" pageId="56" pageNumber="57">Smetana</collectorName>
(1
<collectionCode id="ED45AE58FFE8FFC9FF81FCE65A853F6A" box="[936,1002,835,861]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="56" pageNumber="57" type="Museum">MCZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFE8FFC9FCEEFCC35C743FFE" pageId="56" pageNumber="57" type="diagnosis">
<paragraph id="8BEB369DFFE8FFC9FCEEFCC35C743FFE" blockId="56.[151,1437,690,1941]" pageId="56" pageNumber="57">
<emphasis id="B920EA8FFFE8FFC9FCEEFCC358BA3FB7" bold="true" box="[199,469,870,896]" pageId="56" pageNumber="57">Differential Diagnosis.</emphasis>
This reddish, very convex species will key to
<taxonomicName id="4C544D1EFFE8FFC9FFDDFCC25DA33FB6" authority="Brown" authorityName="Brown" box="[1012,1228,871,897]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="mimicus">
<emphasis id="B920EA8FFFE8FFC9FFDDFCC25D193FB7" box="[1012,1142,871,897]" italics="true" pageId="56" pageNumber="57">O. mimicus</emphasis>
Brown
</taxonomicName>
in
<bibRefCitation id="EFC54B6CFFE8FFC9F8DCFCC259A23F92" author="Perkins, P. D." pageId="56" pageNumber="57" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980: 303)</bibRefCitation>
. The male genitalia clearly show the relationship of the two species, while differing significantly from each other in the shape of the distal piece (
<figureCitation id="136F2A18FFE8FFC9FE13FC0A5BE23FFE" box="[570,653,943,969]" captionStart="FIGURE 33" captionStartId="57.[151,250,1792,1816]" captionTargetBox="[323,1263,181,1768]" captionTargetId="figure-18@57.[323,1263,181,1768]" captionTargetPageId="57" captionText="FIGURE 33. Ochthebius (Asiobates) cascadeus, new species, holotype habitus and aedeagus (inset:aedeagal apex of Ochthebius (Asiobates) mimicus Brown)." figureDoi="http://doi.org/10.5281/zenodo.10145172" httpUri="https://zenodo.org/record/10145172/files/figure.png" pageId="56" pageNumber="57">Fig. 33</figureCitation>
). Reliable determinations will require dissection of males.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFE8FFCAFCEEFC775A733BA1" lastPageId="59" lastPageNumber="60" pageId="56" pageNumber="57" type="description">
<paragraph id="8BEB369DFFE8FFC9FCEEFC775ABC3962" blockId="56.[151,1437,690,1941]" pageId="56" pageNumber="57">
<emphasis id="B920EA8FFFE8FFC9FCEEFC77583E3FDB" bold="true" box="[199,337,978,1004]" pageId="56" pageNumber="57">Description</emphasis>
. Size:
<typeStatus id="54EF883FFFE8FFC9FD8DFC765B693FDA" box="[420,518,979,1005]" pageId="56" pageNumber="57" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.67/0.88; head width 0.52; pronotum 0.48/0.79, PA 0.62, PB 0.53 (PB = sclerotized portion only); elytra 1.00/0.88. Habitus as illustrated (
<figureCitation id="136F2A18FFE8FFC9F946FC5259D93802" captionStart="FIGURE 33" captionStartId="57.[151,250,1792,1816]" captionTargetBox="[323,1263,181,1768]" captionTargetId="figure-18@57.[323,1263,181,1768]" captionTargetPageId="57" captionText="FIGURE 33. Ochthebius (Asiobates) cascadeus, new species, holotype habitus and aedeagus (inset:aedeagal apex of Ochthebius (Asiobates) mimicus Brown)." figureDoi="http://doi.org/10.5281/zenodo.10145172" httpUri="https://zenodo.org/record/10145172/files/figure.png" pageId="56" pageNumber="57">Fig. 33</figureCitation>
). Form ovate, very convex. Pronotum and elytra dark reddish brown, dorsum of head dark brown; legs and palpi light reddish brown. Frons coarsely densely punctate, some punctures confluent, interstices shining; interocular foveae deep and large, width of each slightly greater than distance between them; interocular tuberculi large, large depression between interocular tuberculi and eye; basomedial fovea smaller and shallower than interocular foveae. Frontoclypeal suture evenly arcuate. Clypeus length 0.50 width; medially very finely very sparsely punctulate, shining, laterally more densely punctulate and microreticulate. Labroclypeal suture weakly bisinuate. Labrum length 0.33 width; very finely sparsely punctate, very finely pubescent; anterior margin with deep apicomedian notch, edge very weakly upturned. Maxillary palpus with palpomere 3 moderately wide; palpomere 4 distinctly less than 0.50 length of penultimate. Lateral margin of mandibles (male) without spines.
</paragraph>
<paragraph id="8BEB369DFFE8FFC9FCEEFAFA5A583AD6" blockId="56.[151,1437,690,1941]" pageId="56" pageNumber="57">Pronotum with anterior hyaline border moderately wide in front of disc, slightly wider in front of lateral fossulae, then tapered to anterior angles. Lateral hyaline border origin at anterior angles, extended narrowly along lateral depressions, broad in region of posterior excavation, very narrow around posterior margin. Anterior margin of pronotum trisinuate, with emargination in front of each lateral depression; anterior angles rounded, obtuse. Lateral depressions not appearing inflated, broad, with very shallow, sparse punctures; margins arcuate, irregular; posterior extremes excavate and with small tooth; pronotum markedly constricted behind lateral depressions. Lateral fossulae deeply impressed; inner margin abrupt, posterior extreme ended before lateral hyaline border. Pronotal disc very convex, punctures in interfoveal area large, dense, irregularly sized, separated at most by 1xpd, usually separated by ca. 0.5xpd or thin walls; each puncture with very short seta; surface between punctures smooth and shiny. Median groove deeply impressed, micropunctulate; widened at anterior and posterior. Anterior foveae moderately large. Posterior foveae very large, deep, with diagonal orientation.</paragraph>
<paragraph id="8BEB369DFFE8FFC9FCEEF94E5A8C3BA2" blockId="56.[151,1437,690,1941]" pageId="56" pageNumber="57">Elytral disc without depression in midregion, shiny, with six rows of punctures between suture and humeri. Longitudinally and transversely very convex, appearing humped at posterior declivity, at midlength; punctures round, ca. 1xpd of largest pronotal punctures; intervals rounded, slightly elevated, width ca. 2–3xpd, with very shallow surface irregularities; interstices between punctures ca. 1–2xpd; each puncture with very short decumbent seta. Explanate margin moderately wide. Elytral apices separately rounded.</paragraph>
<caption id="DF2B6615FFE9FFC8FCBEF8A558AF3B0B" ID-DOI="http://doi.org/10.5281/zenodo.10145172" ID-Zenodo-Dep="10145172" httpUri="https://zenodo.org/record/10145172/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1792,1816]" targetBox="[323,1263,181,1768]" targetPageId="57" targetType="figure">
<paragraph id="8BEB369DFFE9FFC8FCBEF8A558AF3B0B" blockId="57.[151,1436,1792,1852]" pageId="57" pageNumber="58">
<emphasis id="B920EA8FFFE9FFC8FCBEF8A558713B2F" bold="true" box="[151,286,1792,1816]" pageId="57" pageNumber="58">FIGURE 33.</emphasis>
<taxonomicName id="4C544D1EFFE9FFC8FD0BF8A55B133B2F" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[290,636,1792,1816]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="cascadeus" status="sp. nov." subGenus="Asiobates">
<emphasis id="B920EA8FFFE9FFC8FD0BF8A558FA3B2F" box="[290,405,1792,1816]" italics="true" pageId="57" pageNumber="58">Ochthebius</emphasis>
(
<emphasis id="B920EA8FFFE9FFC8FD8BF8A55B6A3B2F" box="[418,517,1792,1816]" italics="true" pageId="57" pageNumber="58">Asiobates</emphasis>
)
<emphasis id="B920EA8FFFE9FFC8FE38F8A55B133B2F" box="[529,636,1792,1816]" italics="true" pageId="57" pageNumber="58">cascadeus</emphasis>
</taxonomicName>
<emphasis id="B920EA8FFFE9FFC8FE55F8A55BEF3B2F" box="[636,640,1792,1816]" italics="true" pageId="57" pageNumber="58">,</emphasis>
<taxonomicNameLabel id="A21357F4FFE9FFC8FEADF8A45B933B2F" box="[644,764,1792,1816]" pageId="57" pageNumber="58" rank="species">new species</taxonomicNameLabel>
, holotype habitus and aedeagus (inset: aedeagal apex of
<taxonomicName id="4C544D1EFFE9FFC8F900F8A558DA3B0B" authority="Brown" authorityName="Brown" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="mimicus" subGenus="Asiobates">
<emphasis id="B920EA8FFFE9FFC8F900F8A55CF33B2F" box="[1321,1436,1792,1816]" italics="true" pageId="57" pageNumber="58">Ochthebius</emphasis>
(
<emphasis id="B920EA8FFFE9FFC8FC89F881586C3B0B" box="[160,259,1828,1852]" italics="true" pageId="57" pageNumber="58">Asiobates</emphasis>
)
<emphasis id="B920EA8FFFE9FFC8FD38F880580A3B0B" box="[273,357,1829,1852]" italics="true" pageId="57" pageNumber="58">mimicus</emphasis>
Brown
</taxonomicName>
).
</paragraph>
</caption>
<caption id="DF2B6615FFEAFFCBFCBEF8A558213B0B" ID-DOI="http://doi.org/10.5281/zenodo.10145174" ID-Zenodo-Dep="10145174" httpUri="https://zenodo.org/record/10145174/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,264,1792,1816]" targetBox="[186,1395,184,1753]" targetPageId="58" targetType="figure">
<paragraph id="8BEB369DFFEAFFCBFCBEF8A558213B0B" blockId="58.[151,1436,1792,1852]" pageId="58" pageNumber="59">
<emphasis id="B920EA8FFFEAFFCBFCBEF8A5583E3B2F" bold="true" box="[151,337,1792,1816]" pageId="58" pageNumber="59">FIGURES 34–35.</emphasis>
Habitus and male genitalia.34.
<taxonomicName id="4C544D1EFFEAFFCBFEAAF8A55D2C3B2F" authority="(Perkins)" baseAuthorityName="Perkins" box="[643,1091,1792,1816]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
<emphasis id="B920EA8FFFEAFFCBFEAAF8A55B993B2F" box="[643,758,1792,1816]" italics="true" pageId="58" pageNumber="59">Ochthebius</emphasis>
(
<emphasis id="B920EA8FFFEAFFCBFF28F8A55AC43B2F" box="[769,939,1792,1816]" italics="true" pageId="58" pageNumber="59">Gymnochthebius</emphasis>
)
<emphasis id="B920EA8FFFEAFFCBFF9EF8A55A8F3B2F" box="[951,992,1792,1816]" italics="true" pageId="58" pageNumber="59">falli</emphasis>
(Perkins)
</taxonomicName>
.35.
<taxonomicName id="4C544D1EFFEAFFCBF842F8A558213B0B" authority="LeConte." authorityName="LeConte." class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
<emphasis id="B920EA8FFFEAFFCBF842F8A55DB13B2F" box="[1131,1246,1792,1816]" italics="true" pageId="58" pageNumber="59">Ochthebius</emphasis>
(
<emphasis id="B920EA8FFFEAFFCBF8C3F8A55CFB3B2F" box="[1258,1428,1792,1816]" italics="true" pageId="58" pageNumber="59">Gymnochthebius</emphasis>
)
<emphasis id="B920EA8FFFEAFFCBFCBEF88159863B0B" box="[151,233,1828,1852]" italics="true" pageId="58" pageNumber="59">fossatus</emphasis>
LeConte.
</taxonomicName>
</paragraph>
</caption>
<caption id="DF2B6615FFEBFFCAFCBEF9F85D753A42" ID-DOI="http://doi.org/10.5281/zenodo.10145176" ID-Zenodo-Dep="10145176" box="[151,1050,1629,1653]" httpUri="https://zenodo.org/record/10145176/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,250,1629,1653]" targetBox="[151,1436,181,1605]" targetPageId="59" targetType="figure">
<paragraph id="8BEB369DFFEBFFCAFCBEF9F85D753A42" blockId="59.[151,1050,1629,1653]" box="[151,1050,1629,1653]" pageId="59" pageNumber="60">
<emphasis id="B920EA8FFFEBFFCAFCBEF9F8584F3A42" bold="true" box="[151,288,1629,1653]" pageId="59" pageNumber="60">FIGURE 36.</emphasis>
Geographical distribution of
<taxonomicName id="4C544D1EFFEBFFCAFE62F9F85D793A42" authority="(Perkins)" baseAuthorityName="Perkins" box="[587,1046,1629,1653]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
<emphasis id="B920EA8FFFEBFFCAFE62F9F85BD13A42" box="[587,702,1629,1653]" italics="true" pageId="59" pageNumber="60">Ochthebius</emphasis>
(
<emphasis id="B920EA8FFFEBFFCAFEE7F9F85A173A42" box="[718,888,1629,1653]" italics="true" pageId="59" pageNumber="60">Gymnochthebius</emphasis>
)
<emphasis id="B920EA8FFFEBFFCAFFAEF9F85ADF3A42" box="[903,944,1629,1653]" italics="true" pageId="59" pageNumber="60">falli</emphasis>
(Perkins)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BEB369DFFEBFFCAFCEEF96D5A733BA1" blockId="59.[151,1437,1735,2014]" pageId="59" pageNumber="60">Venter: Mentum moderately shining, with moderately dense moderately fine setigerous punctures, setae short, inconspicuous except longer and denser at anterolateral angles. Prosternum hydrofuge pubescent, with strong median carina. Metaventrite hydrofuge pubescent laterally, with large suboval slightly angulate anteriorly shining with glabrous area on disc. Abdominal ventrites 1–5 hydrofuge pubescent. Ventrite 6 moderately shining, anterior ½ effacedly microreticulate, posterior ½ with with sparce, short setae. Last tergite rounded apically, with short setae. Protarsomeres 1–3 (male) with large pad of suction setae.</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFEBFFCAFCEEF83A5CF23B8E" box="[199,1437,1951,1978]" pageId="59" pageNumber="60" type="distribution">
<paragraph id="8BEB369DFFEBFFCAFCEEF83A5CF23B8E" blockId="59.[151,1437,1735,2014]" box="[199,1437,1951,1978]" pageId="59" pageNumber="60">
<emphasis id="B920EA8FFFEBFFCAFCEEF83A580E3B8E" bold="true" box="[199,353,1951,1977]" pageId="59" pageNumber="60">Distribution.</emphasis>
Currently known only from several localities near the
<typeStatus id="54EF883FFFEBFFCAFFECF8055A9A3B8D" box="[965,1013,1952,1978]" pageId="59" pageNumber="60">type</typeStatus>
locality at Mt. Rainier National Park.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFEBFFCAFCEEF8665DDD3BEA" box="[199,1202,1987,2014]" pageId="59" pageNumber="60" type="etymology">
<paragraph id="8BEB369DFFEBFFCAFCEEF8665DDD3BEA" blockId="59.[151,1437,1735,2014]" box="[199,1202,1987,2014]" pageId="59" pageNumber="60">
<emphasis id="B920EA8FFFEBFFCAFCEEF86658213BEA" bold="true" box="[199,334,1987,2013]" pageId="59" pageNumber="60">Etymology.</emphasis>
Named in reference to the geographical distribution, the Cascade Mountains.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFECFFCDFCEEFF3258D23E56" pageId="60" pageNumber="61" type="discussion">
<paragraph id="8BEB369DFFECFFCDFCEEFF325C553CE2" blockId="60.[151,1436,151,213]" pageId="60" pageNumber="61">
<emphasis id="B920EA8FFFECFFCDFCEEFF3258543C86" bold="true" box="[199,315,151,177]" pageId="60" pageNumber="61">Remarks.</emphasis>
The aedeagal distal piece of
<taxonomicName id="4C544D1EFFECFFCDFEBFFF325A1F3C86" authority="Brown" authorityName="Brown" box="[662,880,151,177]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="mimicus">
<emphasis id="B920EA8FFFECFFCDFEBFFF325A763C87" box="[662,793,151,177]" italics="true" pageId="60" pageNumber="61">O. mimicus</emphasis>
Brown
</taxonomicName>
is illustrated herein (
<figureCitation id="136F2A18FFECFFCDF845FF325DAC3C86" box="[1132,1219,151,177]" captionStart="FIGURE 33" captionStartId="57.[151,250,1792,1816]" captionTargetBox="[323,1263,181,1768]" captionTargetId="figure-18@57.[323,1263,181,1768]" captionTargetPageId="57" captionText="FIGURE 33. Ochthebius (Asiobates) cascadeus, new species, holotype habitus and aedeagus (inset:aedeagal apex of Ochthebius (Asiobates) mimicus Brown)." figureDoi="http://doi.org/10.5281/zenodo.10145172" httpUri="https://zenodo.org/record/10145172/files/figure.png" pageId="60" pageNumber="61">Fig. 33</figureCitation>
), from a specimen collected in
<collectingRegion id="4990F87FFFECFFCDFD09FF1E58183CE2" box="[288,375,187,213]" country="United States of America" name="Oregon" pageId="60" pageNumber="61">Oregon</collectingRegion>
, Wasco County. A line drawing of the entire aedeagus is given in
<bibRefCitation id="EFC54B6CFFECFFCDF87CFF1E5C593CE2" author="Perkins, P. D." box="[1109,1334,187,213]" pageId="60" pageNumber="61" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980: 387)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEB369DFFECFFCDFCBEFE8258BF3D76" blockId="60.[151,464,294,321]" box="[151,464,294,321]" pageId="60" pageNumber="61">
<emphasis id="B920EA8FFFECFFCDFCBEFE8258BF3D76" bold="true" box="[151,464,294,321]" pageId="60" pageNumber="61">New distribution records</emphasis>
</paragraph>
<paragraph id="8BEB369DFFECFFCDFCBEFECA5D9C3DE6" blockId="60.[151,1437,367,609]" pageId="60" pageNumber="61">New distribution records are provided for species that have range extensions, and/or for species that were previously known from very few specimens, to document the new collection records and to record new museum depositions. All new records below are represented by at least one, and usually more, dissected male specimens.</paragraph>
<paragraph id="8BEB369DFFECFFCDFCEEFE7E58D23E56" blockId="60.[151,1437,367,609]" pageId="60" pageNumber="61">
Previously published collection/depository and distribution information presented below are summerized from
<bibRefCitation id="EFC54B6CFFECFFCDFCBEFE5A582F3E2E" author="Perkins, P. D." box="[151,320,511,537]" pageId="60" pageNumber="61" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980)</bibRefCitation>
. The citations for original publication of names, which are presented in the new distribution records section, are given in
<bibRefCitation id="EFC54B6CFFECFFCDFD56FD865B483E0A" author="Perkins, P. D." box="[383,551,547,573]" pageId="60" pageNumber="61" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980)</bibRefCitation>
. Specimen records in
<bibRefCitation id="EFC54B6CFFECFFCDFF34FD865AAC3E0A" author="Perkins, P. D." box="[797,963,547,573]" pageId="60" pageNumber="61" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980)</bibRefCitation>
with the depository given as PDP have all been donated to the MCZ.
</paragraph>
</subSubSection>
</treatment>
</document>
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