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<document id="08E79EA8F3F2737C9B4D68F2EDB96AC3" ID-DOI="10.5281/zenodo.156671" ID-GBIF-Dataset="ad8505f5-c05a-4567-9f98-7618610ae888" ID-ISSN="1175­5326" ID-Zenodo-Dep="156671" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459595750768" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="03C287EB1A32FF8BFECEB668A3BB722A" docLanguage="en" docName="zt00165.pdf" docOrigin="Zootaxa 165" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Mayatyphlus carltoni Gusarov, sp. n." docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FFFBFF931A31FF8FFFC6B774A1097114" masterDocTitle="Mayatyphlus carltoni Gusarov, a new genus and species of leptotyphline staphylinid beetle from Belize (Coleoptera: Staphylinidae: Leptotyphlinae)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="4" updateTime="1698202588245" updateUser="plazi">
<mods:mods id="DBD86E0E3FB8E24659A0126BB3A72D3B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1F5EA36C5F0B83756FC7FF1A0911A238">Mayatyphlus carltoni Gusarov, a new genus and species of leptotyphline staphylinid beetle from Belize (Coleoptera: Staphylinidae: Leptotyphlinae)</mods:title>
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<mods:namePart id="3864F8840ABB8F137A27F6C87F26E13D">Gusarov, Vladimir I.</mods:namePart>
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<mods:typeOfResource id="9C3BF8A84E03C68101E87338BE7699B8">text</mods:typeOfResource>
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<mods:date id="CEF4117DAE58E9B4A18A329DCBE4B496">2003</mods:date>
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<mods:identifier id="ADABFA5C21D059B2FF27A96E6C59A117" type="GBIF-Dataset">ad8505f5-c05a-4567-9f98-7618610ae888</mods:identifier>
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<treatment id="03C287EB1A32FF8BFECEB668A3BB722A" ID-DOI="http://doi.org/10.5281/zenodo.6277099" ID-GBIF-Taxon="119331235" ID-Zenodo-Dep="6277099" LSID="urn:lsid:plazi:treatment:03C287EB1A32FF8BFECEB668A3BB722A" httpUri="http://treatment.plazi.org/id/03C287EB1A32FF8BFECEB668A3BB722A" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="C37165761A32FF8CFECEB668A2567021" box="[264,863,283,310]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BD436FD1A32FF8CFECEB668A2567021" blockId="3.[264,863,283,310]" box="[264,863,283,310]" pageId="3" pageNumber="4">
<heading id="D09C81911A32FF8CFECEB668A2567021" bold="true" box="[264,863,283,310]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="B91FEAEF1A32FF8CFECEB668A2567021" bold="true" box="[264,863,283,310]" pageId="3" pageNumber="4">
<taxonomicName id="4C6B4D7E1A32FF8CFECEB668A3747021" ID-CoL="3YH6Y" authority="Gusarov" authorityName="Gusarov" box="[264,637,283,310]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="carltoni" status="sp. nov.">
<emphasis id="B91FEAEF1A32FF8CFECEB668A3027022" bold="true" box="[264,523,284,310]" italics="true" pageId="3" pageNumber="4">Mayatyphlus carltoni</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="A22C57941A32FF8CFD4CB66FA3C57021" box="[650,716,283,309]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="13502A781A32FF8CFD1DB66FA25F7021" box="[731,854,283,309]" captionStart-0="FIGURES 1 ­ 12" captionStart-1="FIGURES 13 ­ 18" captionStartId-0="2.[264,383,1668,1692]" captionStartId-1="3.[264,383,1768,1792]" captionTargetBox-0="[321,1264,618,1634]" captionTargetBox-1="[352,1236,931,1733]" captionTargetId-0="figure@2.[312,1275,609,1641]" captionTargetId-1="figure@3.[345,1242,924,1742]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." captionText-1="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri-0="https://zenodo.org/record/156672/files/figure.png" httpUri-1="https://zenodo.org/record/156673/files/figure.png" pageId="3" pageNumber="4">Figs. 1­18</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37165761A32FF8CFECEB612A54C70ED" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BD436FD1A32FF8CFECEB612A26570C5" blockId="3.[264,1324,358,825]" pageId="3" pageNumber="4">
<emphasis id="B91FEAEF1A32FF8CFECEB612A0BF7094" bold="true" box="[264,438,358,384]" pageId="3" pageNumber="4">
<typeStatus id="54D0885F1A32FF8CFECEB612A04A7094" box="[264,323,358,384]" pageId="3" pageNumber="4">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54D0885F1A32FF8CFE17B613A34A7095" box="[465,579,359,385]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
:,
<emphasis id="B91FEAEF1A32FF8CFDBAB612A3E07094" bold="true" box="[636,745,358,384]" pageId="3" pageNumber="4">
<collectingCountry id="F37C766D1A32FF8CFDBAB612A3E07094" box="[636,745,358,384]" name="Belize" pageId="3" pageNumber="4">BELIZE</collectingCountry>
</emphasis>
: Orange Walk Distr., Rio Bravo Conservation / Management Area, La Milpa Station, Chiche Trail,
<geoCoordinate id="EE5F503A1A32FF8CFCA0B6FBA2E470BD" box="[870,1005,399,425]" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="15" value="17.839445">17º50'22&quot;N</geoCoordinate>
<geoCoordinate id="EE5F503A1A32FF8CFC32B6FBA58B70BD" box="[1012,1154,399,425]" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="15" value="-89.020004">89º01'12&quot;W</geoCoordinate>
, lot #119, berlese (C.E.Carlton &amp; D.Murray),
<date id="FFD5103D1A32FF8CFD43B6C3A3F170C5" box="[645,760,439,465]" pageId="3" pageNumber="4" value="1997-05-18">18.v.1997</date>
(
<collectionCode id="ED7AAE381A32FF8CFCCEB6C3A26970C5" box="[776,864,439,465]" pageId="3" pageNumber="4">KSEM</collectionCode>
).
</paragraph>
<paragraph id="8BD436FD1A32FF8CFEFEB6ABA54C70ED" blockId="3.[264,1324,358,825]" box="[312,1093,478,505]" pageId="3" pageNumber="4">
<typeStatus id="54D0885F1A32FF8CFEFEB6ABA0A770ED" box="[312,430,479,505]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B91FEAEF1A32FF8CFE06B6AAA32470EC" bold="true" box="[448,557,478,504]" pageId="3" pageNumber="4">
<collectingCountry id="F37C766D1A32FF8CFE06B6AAA32470EC" box="[448,557,478,504]" name="Belize" pageId="3" pageNumber="4">BELIZE</collectingCountry>
</emphasis>
:, 2, same data as the
<typeStatus id="54D0885F1A32FF8CFCB7B6ABA2D170ED" box="[881,984,479,505]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED7AAE381A32FF8CFC21B6ABA53170ED" box="[999,1080,479,505]" pageId="3" pageNumber="4">LSAC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C37165761A32FF8CFEFEB572A2CD738D" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BD436FD1A32FF8CFEFEB572A2CD738D" blockId="3.[264,1324,358,825]" pageId="3" pageNumber="4">
<emphasis id="B91FEAEF1A32FF8CFEFEB572A0B97334" bold="true" box="[312,432,518,544]" pageId="3" pageNumber="4">Diagnosis</emphasis>
.
<taxonomicName id="4C6B4D7E1A32FF8CFE03B573A3C97334" box="[453,704,519,544]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B91FEAEF1A32FF8CFE03B573A3C97334" box="[453,704,519,544]" italics="true" pageId="3" pageNumber="4">Mayatyphlus carltoni</emphasis>
</taxonomicName>
is the only known species of the genus
<taxonomicName id="4C6B4D7E1A32FF8CFB4AB573A42A7334" box="[1164,1315,519,544]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAEF1A32FF8CFB4AB573A42A7334" box="[1164,1315,519,544]" italics="true" pageId="3" pageNumber="4">Mayatyphlus</emphasis>
</taxonomicName>
. If any additional species of
<taxonomicName id="4C6B4D7E1A32FF8CFD97B55BA3E0735C" box="[593,745,559,584]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B91FEAEF1A32FF8CFD97B55BA3E0735C" box="[593,745,559,584]" italics="true" pageId="3" pageNumber="4">Mayatyphlus</emphasis>
</taxonomicName>
are ever discovered,
<taxonomicName id="4C6B4D7E1A32FF8CFC20B55BA563735C" box="[998,1130,559,584]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B91FEAEF1A32FF8CFC20B55BA563735C" box="[998,1130,559,584]" italics="true" pageId="3" pageNumber="4">M. carltoni</emphasis>
</taxonomicName>
could be distinguished by the shape of the aedeagus, especially the apex of the median lobe (
<figureCitation id="13502A781A32FF8CFB52B523A4147365" box="[1172,1309,599,625]" captionStart="FIGURES 13 ­ 18" captionStartId="3.[264,383,1768,1792]" captionTargetBox="[352,1236,931,1733]" captionTargetId="figure@3.[345,1242,924,1742]" captionTargetPageId="3" captionText="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156673/files/figure.png" pageId="3" pageNumber="4">Figs. 13­15</figureCitation>
), and by the shape of the female accessory sclerites (
<figureCitation id="13502A781A32FF8CFC99B50BA2BF738D" box="[863,950,639,665]" captionStart="FIGURES 13 ­ 18" captionStartId="3.[264,383,1768,1792]" captionTargetBox="[352,1236,931,1733]" captionTargetId="figure@3.[345,1242,924,1742]" captionTargetPageId="3" captionText="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156673/files/figure.png" pageId="3" pageNumber="4">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37165761A32FF8BFEFEB5D2A41873D2" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="description">
<paragraph id="8BD436FD1A32FF8CFEFEB5D2A3C073D5" blockId="3.[264,1324,358,825]" box="[312,713,678,705]" pageId="3" pageNumber="4">
<emphasis id="B91FEAEF1A32FF8CFEFEB5D2A0D973D4" bold="true" box="[312,464,678,704]" pageId="3" pageNumber="4">Description.</emphasis>
Length
<quantity id="4C939B181A32FF8CFDFFB5D3A3CD73D5" box="[569,708,679,705]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="3" pageNumber="4" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7­0.8 mm</quantity>
.
</paragraph>
<paragraph id="8BD436FD1A32FF8CFEFEB5BBA419722D" blockId="3.[264,1324,358,825]" pageId="3" pageNumber="4">
Head with short setae arranged as in
<figureCitation id="13502A781A32FF8CFD34B5BBA25773FD" box="[754,862,719,745]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="3" pageNumber="4">Figs. 1­2</figureCitation>
. Second and third antennal articles as long as wide and twice as wide as article 3, articles 3­9 strongly transverse, 1.6­2.0 times as wide as long, article 10 1.5 times as wide as long, last article as long as wide (
<figureCitation id="13502A781A32FF8CFB7CB46BA40B722D" box="[1210,1282,799,825]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
).
</paragraph>
<caption id="DF1466751A32FF8CFECEB19CA50E7651" httpUri="https://zenodo.org/record/156673/files/figure.png" pageId="3" pageNumber="4" targetBox="[352,1236,931,1733]" targetPageId="3">
<paragraph id="8BD436FD1A32FF8CFECEB19CA50E7651" blockId="3.[264,1323,1768,1861]" pageId="3" pageNumber="4">
<emphasis id="B91FEAEF1A32FF8CFECEB19CA0C37614" bold="true" box="[264,458,1768,1792]" pageId="3" pageNumber="4">FIGURES 13­18.</emphasis>
Details of
<taxonomicName id="4C6B4D7E1A32FF8CFD80B19CA28477EB" authority="Gusarov" authorityName="Gusarov" box="[582,909,1768,1791]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="carltoni" status="gen. nov.">
<emphasis id="B91FEAEF1A32FF8CFD80B19CA22277EB" box="[582,811,1768,1791]" italics="true" pageId="3" pageNumber="4">Mayatyphlus carltoni</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="B91FEAEF1A32FF8CFC51B19CA2E87614" bold="true" box="[919,993,1768,1792]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="A22C57941A32FF8CFC51B19CA2E87614" box="[919,993,1768,1792]" pageId="3" pageNumber="4" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B91FEAEF1A32FF8CFC28B19CA5237614" bold="true" box="[1006,1066,1768,1792]" pageId="3" pageNumber="4">sp. n.</emphasis>
(paratypes). 13 aedeagus, parameral view; 14­15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9­10, ventral view; 18 female sternum 8. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BD436FD1A35FF8BFEFEB668A5AA7092" blockId="4.[264,1324,284,830]" pageId="4" pageNumber="5">
Pronotum with short setae arranged as in
<figureCitation id="13502A781A35FF8BFCD9B668A26C7022" box="[799,869,284,310]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
, narrowed posteriad, 1.1 times as long as wide, as wide as head and elytra (
<figureCitation id="13502A781A35FF8BFD79B630A20C704A" box="[703,773,324,350]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
). Elytra with short setae, with parallel lateral sides (
<figureCitation id="13502A781A35FF8BFE95B618A09E7092" box="[339,407,364,390]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
), 1.1 times as long as wide (length measured from humeral level).
</paragraph>
<paragraph id="8BD436FD1A35FF8BFEFEB6E0A21570BA" blockId="4.[264,1324,284,830]" box="[312,796,404,430]" pageId="4" pageNumber="5">
Abdominal terga with long setae (
<figureCitation id="13502A781A35FF8BFD00B6E0A20270BA" box="[710,779,404,430]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BD436FD1A35FF8BFEFEB6C8A3267332" blockId="4.[264,1324,284,830]" pageId="4" pageNumber="5">
Male protarsus with adhesive setae (
<figureCitation id="13502A781A35FF8BFD27B6C8A23070C2" box="[737,825,444,470]" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
). Male mesotrochanter with denticle (
<figureCitation id="13502A781A35FF8BFB3AB6C8A02E70EA" captionStart="FIGURES 1 ­ 12" captionStartId="2.[264,383,1668,1692]" captionTargetBox="[321,1264,618,1634]" captionTargetId="figure@2.[312,1275,609,1641]" captionTargetPageId="2" captionText="FIGURES 1 ­ 12. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (male paratype). 1 forebody; 2 head, ventral view (microsculpture shown on one side only); 3 prothorax, ventral view; 4 meso­ and metathorax, ventral view (microsculpture shown on one side only); 5 left mandible, dorsal view; 6 right mandible, dorsal view; 7 labrum, dorsal view; 8 right maxilla, ventral view; 9 right antenna, dorsal view; 10 left anterior leg, dorsal view; 11 left middle leg, ventral view; 12 left posterior leg, ventral view. Scale bar 0.1 mm (1 ­ 6, 8 ­ 12), 0.05 mm (7)." httpUri="https://zenodo.org/record/156672/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
). Male abdominal sternum 8 with broad apical emargination and narrow medial zone devoid of setae (
<figureCitation id="13502A781A35FF8BFE0CB578A3287332" box="[458,545,524,550]" captionStart="FIGURES 13 ­ 18" captionStartId="3.[264,383,1768,1792]" captionTargetBox="[352,1236,931,1733]" captionTargetId="figure@3.[345,1242,924,1742]" captionTargetPageId="3" captionText="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156673/files/figure.png" pageId="4" pageNumber="5">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8BD436FD1A35FF8BFEFEB540A0A1738A" blockId="4.[264,1324,284,830]" pageId="4" pageNumber="5">
Aedeagus with short and narrow parameres, apex of median lobe in lateral view Sshaped, in parameral view asymmetrical, with two subapical processes pointed to the left (
<figureCitation id="13502A781A35FF8BFED7B5F0A093738A" box="[273,410,644,670]" captionStart="FIGURES 13 ­ 18" captionStartId="3.[264,383,1768,1792]" captionTargetBox="[352,1236,931,1733]" captionTargetId="figure@3.[345,1242,924,1742]" captionTargetPageId="3" captionText="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156673/files/figure.png" pageId="4" pageNumber="5">Figs. 13­15</figureCitation>
).
</paragraph>
<paragraph id="8BD436FD1A35FF8BFEFEB5D8A41873D2" blockId="4.[264,1324,284,830]" box="[312,1297,684,710]" pageId="4" pageNumber="5">
Female abdominal sternum 8 as in
<figureCitation id="13502A781A35FF8BFD17B5D8A22173D2" box="[721,808,684,710]" captionStart="FIGURES 13 ­ 18" captionStartId="3.[264,383,1768,1792]" captionTargetBox="[352,1236,931,1733]" captionTargetId="figure@3.[345,1242,924,1742]" captionTargetPageId="3" captionText="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156673/files/figure.png" pageId="4" pageNumber="5">Fig. 18</figureCitation>
. Female accessory sclerites as in
<figureCitation id="13502A781A35FF8BFB71B5D8A40473D2" box="[1207,1293,684,710]" captionStart="FIGURES 13 ­ 18" captionStartId="3.[264,383,1768,1792]" captionTargetBox="[352,1236,931,1733]" captionTargetId="figure@3.[345,1242,924,1742]" captionTargetPageId="3" captionText="FIGURES 13 ­ 18. Details of Mayatyphlus carltoni Gusarov, gen. n., sp. n. (paratypes). 13 aedeagus, parameral view; 14 ­ 15 aedeagus, lateral view; 16 male sternum 8; 17 female abdominal segments 9 ­ 10, ventral view; 18 female sternum 8. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156673/files/figure.png" pageId="4" pageNumber="5">Fig. 17</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37165761A35FF8BFEFEB5A7A3BB722A" pageId="4" pageNumber="5" type="distribution">
<paragraph id="8BD436FD1A35FF8BFEFEB5A7A2BE73FA" blockId="4.[264,1324,284,830]" box="[312,951,723,750]" pageId="4" pageNumber="5">
<emphasis id="B91FEAEF1A35FF8BFEFEB5A7A0D173F9" bold="true" box="[312,472,723,749]" pageId="4" pageNumber="5">Distribution.</emphasis>
Known from a single locality in
<collectingCountry id="F37C766D1A35FF8BFCA2B5A0A2BA73FA" box="[868,947,724,750]" name="Belize" pageId="4" pageNumber="5">Belize</collectingCountry>
.
</paragraph>
<paragraph id="8BD436FD1A35FF8BFEFEB58FA3BB722A" blockId="4.[264,1324,284,830]" pageId="4" pageNumber="5">
<emphasis id="B91FEAEF1A35FF8BFEFEB58FA30D7201" bold="true" box="[312,516,763,789]" pageId="4" pageNumber="5">Natural History.</emphasis>
<taxonomicName id="4C6B4D7E1A35FF8BFDD4B588A2047201" box="[530,781,764,789]" class="Insecta" family="Staphylinidae" genus="Mayatyphlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="carltoni">
<emphasis id="B91FEAEF1A35FF8BFDD4B588A2047201" box="[530,781,764,789]" italics="true" pageId="4" pageNumber="5">Mayatyphlus carltoni</emphasis>
</taxonomicName>
presumably inhabits soil and/or leaf litter, and was collected using Berlese funnels.
</paragraph>
</subSubSection>
</treatment>
</document>