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<document id="3289E7B7D97A67C9A3461A3B6F155E56" ID-DOI="10.1080/00222933.2011.565130" ID-ISSN="1464-5262" ID-Zenodo-Dep="5204055" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629051536029" checkinUser="felipe" docAuthor="Monné, M. A. &amp; Monné, M. L." docDate="2011" docId="03DC4412575D2B3F6075F9DB50F8F7F7" docLanguage="en" docName="JNatHist.45.29-30.1859-1866.pdf" docOrigin="Journal of Natural History 45 (29 - 30)" docSource="http://dx.doi.org/10.1080/00222933.2011.565130" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Atrocolus guarani Monné &amp; Monné 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="1861" masterDocId="FFE53C6A575F2B3B6251FFCB5478F259" masterDocTitle="A new genus and species of Neotropical Anacolini and new distributional records (Insecta, Coleoptera, Cerambycidae)" masterLastPageNumber="1866" masterPageNumber="1859" pageNumber="1859" updateTime="1699058251373" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="4308EDFFC76BA111D5FA84EDB221F23F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="93EE4CFC97A05AC772873C401B5746F6">
<mods:title id="C702F1797FE343CFB6D6379B45F56952">A new genus and species of Neotropical Anacolini and new distributional records (Insecta, Coleoptera, Cerambycidae)</mods:title>
</mods:titleInfo>
<mods:name id="0FF0CEE68197B0C975C11143383651EB" type="personal">
<mods:role id="4E8ECF48AD5F972CAFDD8A5D1268FC98">
<mods:roleTerm id="50E644D7669A75BF998974C6202842C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="82C510472A952A51A6AF2400F32C635E">Monné, M. A.</mods:namePart>
</mods:name>
<mods:name id="7FEC4D0940871CB617D2C87FE46DCFED" type="personal">
<mods:role id="D5207AF380B9576C8F616309758EC9BC">
<mods:roleTerm id="BBA8CB25126D1E6169754FFD91D9974B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2016C26995C54C3EB5CC4AE9B309DB0">Monné, M. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="23D66F73CCF48AE4E3ABFB2D84BC7627">text</mods:typeOfResource>
<mods:relatedItem id="0A78290ADC99DD7BFEAF332AFA542C50" type="host">
<mods:titleInfo id="0E7DB7B75598E7E6E67820C5B8D02DC2">
<mods:title id="1A2DBB1B7898EB2E40D8DC7FC8C017EA">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="3BDF1AB10DAEF3E8C10EE997424B7D63">
<mods:date id="0E8B275DCF633942CDFD72959E2BF03E">2011</mods:date>
<mods:detail id="1E134F3A764C27D6B1216DBF5CF71AE9" type="pubDate">
<mods:number id="141811AE48138B03DDA0F5BEFB4975AA">2011-08-31</mods:number>
</mods:detail>
<mods:detail id="895EE00ECDFC0FEEA233C4BBBE6532DE" type="volume">
<mods:number id="0F081D400050ED7F83BD6AA1E2685068">45</mods:number>
</mods:detail>
<mods:detail id="623C7D36C542D4D791CE896918365912" type="issue">
<mods:number id="C3F89B311B216E2D41651C870E9C44C1">29 - 30</mods:number>
</mods:detail>
<mods:extent id="1E18BC15A6FD8E0CBDB692E4F76E12B8" unit="page">
<mods:start id="F5EA5850AA13FF5229C54D0896852D56">1859</mods:start>
<mods:end id="2BC13718919536800110E3476DEB6309">1866</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="2E6D9AFC6DB58328A0AC722807A7445F">
<mods:url id="988B3E109E005C8F057C0D5532DF0369">http://dx.doi.org/10.1080/00222933.2011.565130</mods:url>
</mods:location>
<mods:classification id="24FEA232EA6836D3F77763C7475484EF">journal article</mods:classification>
<mods:identifier id="8727882B119DE62B2D1D92012769300E" type="DOI">10.1080/00222933.2011.565130</mods:identifier>
<mods:identifier id="FA9436B48E7FF60E6D801985C6302D48" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="4D349890F7DC82525AA1F6E26C3F3230" type="Zenodo-Dep">5204055</mods:identifier>
</mods:mods>
<treatment id="03DC4412575D2B3F6075F9DB50F8F7F7" LSID="urn:lsid:plazi:treatment:03DC4412575D2B3F6075F9DB50F8F7F7" httpUri="http://treatment.plazi.org/id/03DC4412575D2B3F6075F9DB50F8F7F7" lastPageId="4" lastPageNumber="1861" pageId="2" pageNumber="1859">
<subSubSection id="C36FA68F575D2B396075F9DB573CF47E" box="[548,836,1552,1576]" pageId="2" pageNumber="1859" type="nomenclature">
<paragraph id="8BCAF504575D2B396075F9DB573CF47E" blockId="2.[548,836,1552,1607]" box="[548,836,1552,1576]" pageId="2" pageNumber="1859">
<heading id="D0824268575D2B396075F9DB573CF47E" box="[548,836,1552,1576]" centered="true" fontSize="36" level="2" pageId="2" pageNumber="1859" reason="7">
<taxonomicName id="4C758E87575D2B396075F9DB5690F471" ID-CoL="JLVV" authority="Monné &amp; Monné, 2011" authorityName="Monné &amp; Monné" authorityYear="2011" box="[548,744,1552,1576]" class="Insecta" family="Cerambycidae" genus="Atrocolus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1859" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
<emphasis id="B9012916575D2B396075F9DB5690F471" bold="true" box="[548,744,1552,1576]" italics="true" pageId="2" pageNumber="1859">Atrocolus guarani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232946D575D2B3960A1F9DB573CF47E" box="[752,836,1552,1575]" pageId="2" pageNumber="1859" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36FA68F575D2B3F600DF9E45766F0A0" lastPageId="4" lastPageNumber="1861" pageId="2" pageNumber="1859" type="description">
<paragraph id="8BCAF504575D2B39600DF9E45772F41E" blockId="2.[548,836,1552,1607]" box="[604,778,1583,1607]" pageId="2" pageNumber="1859">
(
<figureCitation id="134EE981575D2B396035F9E4577AF41E" box="[612,770,1583,1607]" captionStart="Figure 1" captionStartId="3.[144,211,1705,1727]" captionTargetBox="[145,1100,161,1672]" captionTargetId="figure-6@3.[144,1101,160,1672]" captionTargetPageId="3" captionText="Figure 1. (A) Atrocolus guarani sp. nov., male holotype; (B) Atrocolus guarani sp. nov., female paratype; (C) Acanthocornis flavus sp. nov., female holotype; (D) Myzomorphus gounellei Lameere, female." figureDoi="http://doi.org/10.5281/zenodo.5204057" httpUri="https://zenodo.org/record/5204057/files/figure.png" pageId="2" pageNumber="1859">Figures 1A, B</figureCitation>
)
</paragraph>
<caption id="DF0AA58C575C2B3862C1F9625545F4A0" ID-DOI="http://doi.org/10.5281/zenodo.5204057" ID-Zenodo-Dep="5204057" httpUri="https://zenodo.org/record/5204057/files/figure.png" pageId="3" pageNumber="1860" startId="3.[144,211,1705,1727]" targetBox="[145,1100,161,1672]" targetPageId="3">
<paragraph id="8BCAF504575C2B3862C1F9625545F4A0" blockId="3.[144,1101,1705,1785]" pageId="3" pageNumber="1860">
Figure 1. (A)
<taxonomicName id="4C758E87575C2B386370F96255ABF4E6" authority="Monné &amp; Monné, 2011" authorityName="Monné &amp; Monné" authorityYear="2011" box="[289,467,1705,1727]" class="Insecta" family="Cerambycidae" genus="Atrocolus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1860" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
<emphasis id="B9012916575C2B386370F96255ABF4E6" box="[289,467,1705,1727]" italics="true" pageId="3" pageNumber="1860">Atrocolus guarani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232946D575C2B38638BF9615651F4E6" box="[474,553,1706,1727]" pageId="3" pageNumber="1860" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype; (B)
<taxonomicName id="4C758E87575C2B3860A9F96257D1F4E6" authority="Monné &amp; Monné, 2011" authorityName="Monné &amp; Monné" authorityYear="2011" box="[760,937,1705,1727]" class="Insecta" family="Cerambycidae" genus="Atrocolus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1860" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
<emphasis id="B9012916575C2B3860A9F96257D1F4E6" box="[760,937,1705,1727]" italics="true" pageId="3" pageNumber="1860">Atrocolus guarani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232946D575C2B3861E1F9615078F4E6" box="[944,1024,1706,1727]" pageId="3" pageNumber="1860" rank="species">sp. nov.</taxonomicNameLabel>
, female paratype; (C)
<taxonomicName id="4C758E87575C2B38637BF90C5586F485" authority="Monné &amp; Monné, 2011" authorityName="Monné &amp; Monné" authorityYear="2011" box="[298,510,1735,1756]" class="Insecta" family="Cerambycidae" genus="Acanthocornis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1860" phylum="Arthropoda" rank="species" species="flavus" status="sp. nov.">
<emphasis id="B9012916575C2B38637BF90C5586F485" box="[298,510,1735,1756]" italics="true" pageId="3" pageNumber="1860">Acanthocornis flavus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A232946D575C2B38605BF90C5627F485" box="[522,607,1735,1756]" pageId="3" pageNumber="1860" rank="species">sp. nov.</taxonomicNameLabel>
, female holotype; (D)
<taxonomicName id="4C758E87575C2B38610AF90C5492F4A0" authority="Lameere" authorityName="Lameere" authorityYear="1912" class="Insecta" family="Cerambycidae" genus="Myzomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1860" phylum="Arthropoda" rank="species" species="gounellei">
<emphasis id="B9012916575C2B38610AF90C5034F485" box="[859,1100,1735,1756]" italics="true" pageId="3" pageNumber="1860">Myzomorphus gounellei</emphasis>
Lameere
</taxonomicName>
, female.
</paragraph>
</caption>
<paragraph id="8BCAF504575B2B3F6284FF6D552CF2E7" blockId="4.[213,340,166,190]" box="[213,340,166,190]" pageId="4" pageNumber="1861">
<emphasis id="B9012916575B2B3F6284FF6D552CF2E7" bold="true" box="[213,340,166,190]" italics="true" pageId="4" pageNumber="1861">Description</emphasis>
</paragraph>
<paragraph id="8BCAF504575B2B3F6284FF055577F2BF" blockId="4.[213,271,206,230]" box="[213,271,206,230]" pageId="4" pageNumber="1861">
<emphasis id="B9012916575B2B3F6284FF055577F2BF" box="[213,271,206,230]" italics="true" pageId="4" pageNumber="1861">Male</emphasis>
</paragraph>
<paragraph id="8BCAF504575B2B3F6284FF3C55F7F074" blockId="4.[213,1170,246,557]" pageId="4" pageNumber="1861">Integument dark brown. Head with sparse punctures, concentrated in the internal sides of the upper lobes of the eyes. Antennae hairless, the apex attaining the basal third of the elytra; scape with fine and sparse punctures, antennomeres 3–11 with a few superficial punctures. Pronotum with regular surface and sparse punctures more concentrated on the sides. Prosternum densely rugose-punctate; mesosternum and metasternum very finely and sparsely punctate. Elytra hairless, with eight longitudinal carinae from base to apex, finely punctate in the dull interspace between carinae. Femora and tibiae finely and sparsely punctate. Ventrites I–IV hairless, very finely and sparsely punctate. Ventrite V rounded in the apex and provided with a tuft of relatively short rigid hairs.</paragraph>
<paragraph id="8BCAF504575B2B3F6284FD91555CF02B" blockId="4.[213,292,602,626]" box="[213,292,602,626]" pageId="4" pageNumber="1861">
<emphasis id="B9012916575B2B3F6284FD91555CF02B" box="[213,292,602,626]" italics="true" pageId="4" pageNumber="1861">Female</emphasis>
</paragraph>
<paragraph id="8BCAF504575B2B3F6284FD485766F0A0" blockId="4.[213,1170,642,762]" pageId="4" pageNumber="1861">Integument mostly dark brown; prothorax and basal half of the elytra chestnut-orange coloured, in the elytra this colour extended backwards by the external sides reaching the beginning of the apical third. Antennae shorter, reaching the basal fifth of the elytra. Ventrite V transversely truncated in the apex.</paragraph>
</subSubSection>
<subSubSection id="C36FA68F575B2B3F6284FCEC5650F1DE" pageId="4" pageNumber="1861" type="etymology">
<paragraph id="8BCAF504575B2B3F6284FCEC5535F166" blockId="4.[213,333,807,831]" box="[213,333,807,831]" pageId="4" pageNumber="1861">
<emphasis id="B9012916575B2B3F6284FCEC5535F166" bold="true" box="[213,333,807,831]" italics="true" pageId="4" pageNumber="1861">Etymology</emphasis>
</paragraph>
<paragraph id="8BCAF504575B2B3F6284FC845650F1DE" blockId="4.[213,1170,847,903]" pageId="4" pageNumber="1861">
The species epithet refers to the people that inhabit
<collectingCountry id="F362B594575B2B3F6170FC9B57F3F131" box="[801,907,848,872]" name="Paraguay" pageId="4" pageNumber="1861">Paraguay</collectingCountry>
,
<collectingCountry id="F362B594575B2B3F61C7FC9B5783F131" box="[918,1019,848,872]" name="Uruguay" pageId="4" pageNumber="1861">Uruguay</collectingCountry>
, and parts of
<collectingCountry id="F362B594575B2B3F6284FCA45532F1DE" box="[213,330,879,903]" name="Argentina" pageId="4" pageNumber="1861">Argentina</collectingCountry>
,
<collectingCountry id="F362B594575B2B3F6304FCA455E0F1DE" box="[341,408,879,903]" name="Brazil" pageId="4" pageNumber="1861">Brazil</collectingCountry>
and
<collectingCountry id="F362B594575B2B3F6381FCA4565CF1DE" box="[464,548,879,903]" name="Bolivia" pageId="4" pageNumber="1861">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36FA68F575B2B3F6284FC7E56E5F64D" pageId="4" pageNumber="1861" type="description">
<paragraph id="8BCAF504575B2B3F6284FC7E575DF195" blockId="4.[213,805,948,972]" box="[213,805,948,972]" pageId="4" pageNumber="1861">
<emphasis id="B9012916575B2B3F6284FC7E5610F195" bold="true" box="[213,616,948,972]" italics="true" pageId="4" pageNumber="1861">
Measurements in mm,
<typeStatus id="54CE4BA6575B2B3F639FFC7F5654F195" box="[462,556,948,972]" pageId="4" pageNumber="1861" type="holotype">holotype</typeStatus>
male
</emphasis>
/
<emphasis id="B9012916575B2B3F6024FC7E575DF195" bold="true" box="[629,805,948,972]" italics="true" pageId="4" pageNumber="1861">
<typeStatus id="54CE4BA6575B2B3F6024FC7E56ADF195" box="[629,725,949,972]" pageId="4" pageNumber="1861" type="paratype">paratype</typeStatus>
female
</emphasis>
</paragraph>
<paragraph id="8BCAF504575B2B3F6284FC1756E5F64D" blockId="4.[213,1170,988,1044]" pageId="4" pageNumber="1861">
Total length 16.0
<emphasis id="B9012916575B2B3F63CCFC1755D1F1AD" bold="true" box="[413,425,988,1012]" italics="true" pageId="4" pageNumber="1861">/</emphasis>
19.9; prothorax length 2.8
<emphasis id="B9012916575B2B3F608CFC175691F1AD" bold="true" box="[733,745,988,1012]" italics="true" pageId="4" pageNumber="1861">/</emphasis>
3.0; prothorax width 5.0
<emphasis id="B9012916575B2B3F6658FC17506DF1AD" bold="true" box="[1033,1045,988,1012]" italics="true" pageId="4" pageNumber="1861">/</emphasis>
5.4; elytral length 11.0
<emphasis id="B9012916575B2B3F6300FC375525F64D" bold="true" box="[337,349,1020,1044]" italics="true" pageId="4" pageNumber="1861">/</emphasis>
13.5; humeral width 5.8
<emphasis id="B9012916575B2B3F6039FC37560CF64D" bold="true" box="[616,628,1020,1044]" italics="true" pageId="4" pageNumber="1861">/</emphasis>
6.5.
</paragraph>
</subSubSection>
<subSubSection id="C36FA68F575B2B3F6284FB89567EF698" pageId="4" pageNumber="1861" type="materials_examined">
<paragraph id="8BCAF504575B2B3F6284FB895516F600" blockId="4.[213,366,1089,1113]" box="[213,366,1089,1113]" pageId="4" pageNumber="1861">
<emphasis id="B9012916575B2B3F6284FB895516F600" bold="true" box="[213,366,1089,1113]" italics="true" pageId="4" pageNumber="1861">Type material</emphasis>
</paragraph>
<paragraph id="8BCAF504575B2B3F6284FBA2567EF698" blockId="4.[213,1170,1129,1217]" pageId="4" pageNumber="1861">
Male
<typeStatus id="54CE4BA6575B2B3F6344FBA25502F6D8" box="[277,378,1129,1153]" pageId="4" pageNumber="1861" type="holotype">holotype</typeStatus>
,
<collectingCountry id="F362B594575B2B3F63D5FBA15596F6DB" box="[388,494,1130,1154]" name="Paraguay" pageId="4" pageNumber="1861">Paraguay</collectingCountry>
,
<collectingRegion id="49B13BE6575B2B3F63A6FBA25616F6D8" box="[503,622,1129,1153]" country="Paraguay" name="Cordillera" pageId="4" pageNumber="1861">Cordillera</collectingRegion>
: San Bernardino,
<date id="FFCBD3C4575B2B3F6163FBA2579EF6D8" box="[818,998,1129,1153]" pageId="4" pageNumber="1861" value="1930-11">November 1930</date>
, K. Fiebrig col. (
<collectionCode id="ED646DC1575B2B3F628CFB42554FF6F8" box="[221,311,1161,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="1861" type="Museum">USNM</collectionCode>
).
<typeStatus id="54CE4BA6575B2B3F631AFB4155D7F6FB" box="[331,431,1162,1186]" pageId="4" pageNumber="1861" type="paratype">Paratype</typeStatus>
female, same locality and collector, in
<taxonomicName id="4C758E87575B2B3F6139FB4255E9F698" authority="Casar." authorityName="Casar." class="Magnoliopsida" family="Euphorbiaceae" genus="Alchornea" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="1861" phylum="Tracheophyta" rank="species" species="iricurana">
<emphasis id="B9012916575B2B3F6139FB42503BF6F8" box="[872,1091,1161,1185]" italics="true" pageId="4" pageNumber="1861">Alchornea iricurana</emphasis>
Casar. (Euphorbiaceae)
</taxonomicName>
(
<collectionCode id="ED646DC1575B2B3F63F1FB625582F698" box="[416,506,1193,1217]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="1861" type="Museum">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36FA68F575B2B3F6284FB2550F8F7F7" pageId="4" pageNumber="1861" type="discussion">
<paragraph id="8BCAF504575B2B3F6284FB255532F75F" blockId="4.[213,330,1262,1286]" box="[213,330,1262,1286]" pageId="4" pageNumber="1861">
<emphasis id="B9012916575B2B3F6284FB255532F75F" bold="true" box="[213,330,1262,1286]" italics="true" pageId="4" pageNumber="1861">Comments</emphasis>
</paragraph>
<paragraph id="8BCAF504575B2B3F6284FADD50F8F7F7" blockId="4.[213,1170,1302,1454]" pageId="4" pageNumber="1861">
<taxonomicName id="4C758E87575B2B3F6284FADD55E5F777" authorityName="Monné &amp; Monné" authorityYear="2011" box="[213,413,1302,1326]" class="Insecta" family="Cerambycidae" genus="Atrocolus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1861" phylum="Arthropoda" rank="species" species="guarani">
<emphasis id="B9012916575B2B3F6284FADD55E5F777" box="[213,413,1302,1326]" italics="true" pageId="4" pageNumber="1861">Atrocolus guarani</emphasis>
</taxonomicName>
is very similar in aspect to
<taxonomicName id="4C758E87575B2B3F60B9FADC5576F717" authority="Monne and Monne, 2008" authorityName="Monne and Monne" authorityYear="2008" class="Insecta" family="Cerambycidae" genus="Atrocolus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1861" phylum="Arthropoda" rank="species" species="mariahelenae">
<emphasis id="B9012916575B2B3F60B9FADC57E6F777" box="[744,926,1302,1326]" italics="true" pageId="4" pageNumber="1861">A. mariahelenae</emphasis>
<bibRefCitation id="EFE488F5575B2B3F61F8FADD5576F717" author="Monne MA &amp; Monne ML" pageId="4" pageNumber="1861" pagination="215 - 217" refId="ref2401" refString="Monne MA, Monne ML. 2008. Atrocolus mariahelenae, novo genero e especie de Anacolini (Coleoptera, Cerambycidae). Iheringia Ser Zool. 98 (2): 215 - 217." type="journal article" year="2008">Monné and Monné, 2008</bibRefCitation>
</taxonomicName>
. It differs by the sparse punctures in the occiput and pronotum, by the dull interspaces between elytral carinae and by the hairless abdomen. In
<taxonomicName id="4C758E87575B2B3F618CFA9C50E9F737" authorityName="Monne and Monne" authorityYear="2008" box="[989,1169,1366,1390]" class="Insecta" family="Cerambycidae" genus="Atrocolus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1861" phylum="Arthropoda" rank="species" species="mariahelenae">
<emphasis id="B9012916575B2B3F618CFA9C50E9F737" box="[989,1169,1366,1390]" italics="true" pageId="4" pageNumber="1861">A. mariahelenae</emphasis>
</taxonomicName>
the occiput and pronotum are densely and coarsely punctate, the interspace between carinae is shining and the abdomen is hairy in the posterior margin of each ventrite.
</paragraph>
</subSubSection>
</treatment>
</document>
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