<mods:titleid="A16A4E16FDB88D4558A80952A25D14F8">Diversity underfoot of agromyzids (Agromyzidae, Diptera) mining thalli of liverworts and hornworts</mods:title>
<figureCitationid="CE7410D5F2401D3310E39264F0647287"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">Fig. 13.</figureCitation>
A medium-sized species (wing length 1.4-1.8 mm) having subshiny brown scutum with brownish yellow pattern extending from mid-posterior margin to scutellum, black 1st flagellomere, brown maxillary palpus, brown halteres, and brownish yellow legs. Male epandrium inner-laterally with a comb comprising five long fused tubercle-like setae. Larva mines the thallus of
<figureCitationid="96571E0011A8D020AE4153B4171BAFF9"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13A-D</figureCitation>
<figureCitationid="E0BEA7898FF2D324DFDEDAA54C7D8A62"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13C</figureCitation>
<figureCitationid="E92CE5A7072929CCEF66076CCCAD4B13"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13B</figureCitation>
<figureCitationid="9C9ED8FEBAAB7D1C33C451253ED6B18E"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13D</figureCitation>
). Scutellum yellow with lateral corner narrowly brown. Subscutellum yellow. Mediotergite, katatergite and anatergite brown. Pleuron yellow, but propleuron, meron, anepimeron centrally pale brown (Fig.
<figureCitationid="E92551632D672A7ED375F2C68943A695"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13B</figureCitation>
<figureCitationid="9CA81A9A1055AA1F9C204149D3BD4351"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13D</figureCitation>
<figureCitationid="9263507B909CFFA05C9C22C897923A56"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13D</figureCitation>
<figureCitationid="B531F7E1249DC8D6800D1620A99911A1"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13A</figureCitation>
<figureCitationid="F8828FA46B87F3A5F0EF834418505A92"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13D</figureCitation>
<figureCitationid="C6F661F1A7D4AA853389CEB24A3FB470"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13F-I</figureCitation>
) Epandrium dark brown, rounded apically; posterior end of inner margin with a stout tubercle-like seta; inner-ventral surface of epandrium with a comb comprising five fused long tubercle-like setae; inner-ventral margin with a row of several short tubercle-like setae (Fig.
<figureCitationid="3D02022F6881E18C2D3FDC767422CBBA"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13I</figureCitation>
). Surstylus rounded, directed inwards, setose apically but without tubercle-like seta. Cercus narrow, setose. Subepandrial sclerite with one pair of posteriorly directed plate-like arms (Fig.
<figureCitationid="3BBA26926C493143FEDEC97493A1595B"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13I</figureCitation>
<figureCitationid="4BA24505E31318E9224947D2F0766222"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13G</figureCitation>
<figureCitationid="649D91BA20A90BCCE80B8949FF5AE2E2"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13H</figureCitation>
<figureCitationid="B56B1531AF5B749376A69CEDBBA285EC"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13G</figureCitation>
<figureCitationid="C8CC4E117F093BE654F36ABFADB9485B"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13H</figureCitation>
). Basiphallus composing of a left anterodorsal sclerite and a right L-shaped lateral sclerite; former narrow, protruding rightward, the latter protruding ventrally (Fig.
<figureCitationid="B47001E5CB3868DA9352CF43F9AF69A7"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13G-H</figureCitation>
<figureCitationid="A6FE70093E1D1BB9A499AD9230700E44"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13G, H</figureCitation>
<figureCitationid="BBDB7266E9D2D61A5D016A8CA5669450"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13H</figureCitation>
). Distiphallus consists of a pair of moderately long tubules slightly diverging apically; basal half composed of dark narrow lobate sclerite and weaker medial region; distal half cylindrical, dorsally pigmented, with truncated, unpigmented apex (Fig.
<figureCitationid="748137BEEC0DBD5A5A922829B995B793"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13H</figureCitation>
<figureCitationid="98852BAEF41046DB4FB64417E481484E"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13F</figureCitation>
<figureCitationid="236E87DB279A320DB14D6786FE44DED8"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13E</figureCitation>
<figureCitationid="CBF53BCADC058EC45DF689FA904DBAFB"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13E</figureCitation>
<figureCitationid="DFED9A8010C52CCF4132B6B1ADE27D2F"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13K, L</figureCitation>
<figureCitationid="9193199CDEFC2019FCA431C8E586E612"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13K</figureCitation>
<figureCitationid="C23016D3F25747759B15E797F3CD3354"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13L</figureCitation>
<figureCitationid="CBDC102AA2F68C4B9427EB0775F627C6"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13L</figureCitation>
<figureCitationid="B310F142E831153EF524F78CDD07D8C9"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13K</figureCitation>
<figureCitationid="6769CBD5F8DBB137E05E770B669B7985"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13J</figureCitation>
<figureCitationid="18D44889FB07D406C2150317395518F6"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13N-P</figureCitation>
) Larvae mine the midrib of the thallus in early instars, later construct radiate mines by coming out from the midrib, and pupate in the mines. Puparium is smooth and pale brown, posterior spiracles projecting (Fig.
<figureCitationid="E5212FA9B94625EEC7740D3E835F8FAD"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13J</figureCitation>
<figureCitationid="C5D46FD7DD56F1CF03D04EA3C1541362"captionStart="Figure 13"captionStartId="F13"captionText="Figure 13. Phytoliriomyza crepusculum sp. nov. A-C holotype male A habitus B lateral C frontal D paratype male (MK-AG-a 309), dorsal E paratype female (MK-AG- 274), dorsal F-I male genitalia F ejaculatory apodeme, dorsal G whole genitalia, ventral H phallic complex, lateral I epandrium, ventral J puparium K-L female postabdomen K oviscape and spermatheca L tergite 10 M habitat of type locality N-P mined thalli of Dumortiera hirsuta (N type locality O Kiyosumi P Mt. Nabejiri). An arrow in N indicate an puparium."figureDoi="10.3897/zookeys.1133.94530.figure13"httpUri="https://binary.pensoft.net/fig/775386"pageId="0"pageNumber="1">13M</figureCitation>
<figureCitationid="65B29D722F399D0A53139D2325743792"captionStart="Figure 12"captionStartId="F12"captionText="Figure 12. Locality records of three Phytoliriomyza species associated with Dumortiera spp.: P. dumortierae and P. crepusculum."figureDoi="10.3897/zookeys.1133.94530.figure12"httpUri="https://binary.pensoft.net/fig/775385"pageId="0"pageNumber="1">12</figureCitation>
in having a pair of brown lateral bands on the scutum; it is distinguished from them by the brownish yellow color of the medial mark on scutum (light yellow in the other species).
