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141 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.400.6611" ID-GBIF-Dataset="a1a4245d-cd0a-431f-8556-fdb7816910c4" ID-PMC="PMC4023243" ID-Pensoft-Pub="1313-2970-400-67" ID-PubMed="24843256" ID-ZBK="91DC4784F49A4353B4C8DC0F67B1EF92" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-400-67" ModsDocOrigin="ZooKeys 400" ModsDocTitle="A new species of the paper wasp genus Polistes (Hymenoptera, Vespidae, Polistinae) in Europe revealed by morphometrics and molecular analyses" checkinTime="1451245992475" checkinUser="pensoft" docAuthor="Neumeyer, Rainer, Baur, Hannes, Guex, Gaston-Denis & Praz, Christophe" docDate="2014" docId="456F92E1EA0749A046427681DD0EDDB4" docLanguage="en" docName="ZooKeys 400: 67-118" docOrigin="ZooKeys 400" docSource="http://dx.doi.org/10.3897/zookeys.400.6611" docTitle="Polistes hellenicus Arens 2011" docType="treatment" docVersion="4" lastPageNumber="93" masterDocId="F67BFF80B122D151FFD22016FFC45C1E" masterDocTitle="A new species of the paper wasp genus Polistes (Hymenoptera, Vespidae, Polistinae) in Europe revealed by morphometrics and molecular analyses" masterLastPageNumber="118" masterPageNumber="67" pageNumber="92" updateTime="1668158214865" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of the paper wasp genus Polistes (Hymenoptera, Vespidae, Polistinae) in Europe revealed by morphometrics and molecular analyses</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neumeyer, Rainer</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Baur, Hannes</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Guex, Gaston-Denis</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Praz, Christophe</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>400</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>67</mods:start>
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<mods:end>118</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.400.6611</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.400.6611</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-400-67</mods:identifier>
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<mods:identifier type="ZBK">91DC4784F49A4353B4C8DC0F67B1EF92</mods:identifier>
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<mods:identifier type="ZooBank">91DC4784F49A4353B4C8DC0F67B1EF92</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127857278" LSID="urn:lsid:plazi:treatment:456F92E1EA0749A046427681DD0EDDB4" httpUri="http://treatment.plazi.org/id/456F92E1EA0749A046427681DD0EDDB4" lastPageId="26" lastPageNumber="93" pageId="25" pageNumber="92">
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<subSubSection pageId="25" pageNumber="92" type="nomenclature">
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<paragraph pageId="25" pageNumber="92">
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<taxonomicName LSID="http://species-id.net/wiki/Polistes_hellenicus" authority="Arens, 2011" authorityName="Arens" authorityYear="2011" class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes hellenicus" order="Hymenoptera" pageId="25" pageNumber="92" phylum="Arthropoda" rank="species" species="hellenicus">Polistes hellenicus Arens, 2011</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="92" type="reference_group">
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<paragraph pageId="25" pageNumber="92">
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes hellenicus" order="Hymenoptera" pageId="25" pageNumber="92" phylum="Arthropoda" rank="species" species="hellenicus">Polistes hellenicus</taxonomicName>
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Arens, 2011: 464 - Holotype male (WA), type locality Ano Kotili, Greece (examined by RN)
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="26" lastPageNumber="93" pageId="25" pageNumber="92" type="diagnosis">
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<paragraph pageId="25" pageNumber="92">Diagnosis.</paragraph>
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<paragraph pageId="25" pageNumber="92">Relatively small, bright species with flagellum bright yellow on upper side in both sexes; pedicel and extreme base of flagellomere 1 always black on upper side.</paragraph>
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<paragraph lastPageId="26" lastPageNumber="93" pageId="25" pageNumber="92">
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Females: Epicnemial carina distinct. Hypopygium often orange and yellow at the tip. Metacoxa spotted yellow on upper side. Mesoscutum sometimes with pair of yel
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<pageBreakToken pageId="26" pageNumber="93" start="start">low</pageBreakToken>
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spots. Propodeum laterally often with yellow spot on each side. Clypeus breadth: malar space 3.46-4.46; head breadth: malar space 7.75-10.65; malar space: lateral ocelli distance 0.85-1.24; metatibia length: malar space 9.00-11.13; terminal flagellomere length: malar space 0.74-1.11.
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</paragraph>
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<paragraph pageId="26" pageNumber="93">Males: Gena in dorsal view immediately narrowing behind eye. Epicnemium and mesosternum black (Fig. 8). Head breadth: head height 1.15-1.21; lower face: clypeus breadth 1.48-1.62; terminal flagellomere length: lateral ocelli distance 1.21-1.4; terminal flagellomere length: malar space 1.44-2.16; terminal flagellomere length: terminal flagellomere breadth 2.12-2.76.</paragraph>
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<caption pageId="26" pageNumber="93">
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<paragraph pageId="26" pageNumber="93">
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Figure 8. Lateral aspect of the holotype (individual RN0242; male) of
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes hellenicus" order="Hymenoptera" pageId="26" pageNumber="93" phylum="Arthropoda" rank="species" species="hellenicus">Polistes hellenicus</taxonomicName>
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Arens, 2011. Unique for males of European
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes" order="Hymenoptera" pageId="26" pageNumber="93" phylum="Arthropoda" rank="genus">Polistes</taxonomicName>
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is the black ventral side of the mesosoma, especially epicnemium and mesosternum.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="93" type="comments">
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<paragraph pageId="26" pageNumber="93">Comments.</paragraph>
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<paragraph pageId="26" pageNumber="93">
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The male holotype of
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes hellenicus" order="Hymenoptera" pageId="26" pageNumber="93" phylum="Arthropoda" rank="species" species="hellenicus">Polistes hellenicus</taxonomicName>
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can be easily distinguished from
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes bischoffi" order="Hymenoptera" pageId="26" pageNumber="93" phylum="Arthropoda" rank="species" species="bischoffi">Polistes bischoffi</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes helveticus" order="Hymenoptera" pageId="26" pageNumber="93" phylum="Arthropoda" rank="species" species="helveticus">Polistes helveticus</taxonomicName>
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sp. n. based on its black epicnemium and mesosternum (Fig. 8). From a morphometric point of view the holotype (
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes hellenicus" order="Hymenoptera" pageId="26" pageNumber="93" phylum="Arthropoda" rank="species" species="hellenicus">Polistes hellenicus</taxonomicName>
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) lies at the border with
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes bischoffi" order="Hymenoptera" pageId="26" pageNumber="93" phylum="Arthropoda" rank="species" species="bischoffi">Polistes bischoffi</taxonomicName>
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, but clearly outside the range of
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<taxonomicName class="Insecta" family="Eumenidae" genus="Polistes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polistes helveticus" order="Hymenoptera" pageId="26" pageNumber="93" phylum="Arthropoda" rank="species" species="helveticus">Polistes helveticus</taxonomicName>
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sp. n. (Fig. 3b, c [L]).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="93" type="material examined">
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<paragraph pageId="26" pageNumber="93">Material examined.</paragraph>
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<paragraph pageId="26" pageNumber="93">Holotype ♂ (RN0242): Greece, Arcadia, Ano Kotili, peak of Lykaion, 1100-1400 m, 07 Jul 2010, Werner Arens leg., WA coll.</paragraph>
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<paragraph pageId="26" pageNumber="93">Paratypes: 1 ♀ (RN0174): Greece, Arcadia, Andritsena, 1997, Werner Arens leg., ZSM coll.; 1 ♂ (RN0125): Ano Kotili, peak of Lykaion, 1100-1400 m, 07 Jul 2010, Werner Arens leg., WA coll.; 1 ♂ (RN0179): Argolis, Drepano, 06 Jul 2008, Werner Arens leg., WA coll.</paragraph>
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<paragraph pageId="26" pageNumber="93">
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Further material: 4 ♂ (RN0220, RN0221, RN0222, RN0223): Croatia, Istria,
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<normalizedToken originalValue="Ližnjan">Liznjan</normalizedToken>
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,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="44.816666">44°49'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="13.983334">13°59'E</geoCoordinate>
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, 27 Aug 2005, Christian Schmid-Egger leg., CSE coll.; 2 ♀ (RN0008, RN0088): Rovinj,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="45.10197">45°06'07.1"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="13.652333">13°39'08.4"E</geoCoordinate>
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, 22 m, 22 Jul 2011, Rainer Neumeyer leg., RN coll.; 1 ♀ (RN0097): Rovinj, Cesta za Mondelaco,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="45.09933">45°05'57.6"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="13.643556">13°38'36.8"E</geoCoordinate>
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, 15 m, 24 Jul 2012, Rainer Neumeyer leg., CSE coll.; 1 ♀ (RN0098): 28 Jul 2012, Rainer Neumeyer leg., CSE coll.; 1 ♀ (RN0193): Rainer Neumeyer leg., RN coll.; 2 ♀ (RN0010, RN0011): Rovinj, Cesta za Valaltu-Lim,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="45.104664">45°06'16.8"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="13.641222">13°38'28.4"E</geoCoordinate>
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, 26 m, 26 Jul 2011, Rainer Neumeyer leg., RN coll.; 1 ♀ (RN0191): 21 Jul 2012, Rainer Neumeyer leg., RN coll.; 1 ♀ (RN0193): Rovinj, near Valalta,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="45.11725">45°07'02.1"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="13.631667">13°37'54.0"E</geoCoordinate>
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, 14 m, 21 Jul 2012, Rainer Neumeyer leg., RN coll.; 2 ♂ (RN0101, RN0102): Vela
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<normalizedToken originalValue="Učka">Ucka</normalizedToken>
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,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="45.307137">45°18'25.7"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="14.194555">14°11'40.4"E</geoCoordinate>
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, 824 m, 27 Jul 2012, karst mountain range, Rainer Neumeyer leg., RN coll.; 1 ♀ (RN0183): Greece, Arcadia, Andritsena, 31 May 2011, Werner Arens leg., WA coll.; 1 ♀ (RN0132): Mantinea, 08 Jun 2011, Werner Arens leg., WA coll.; 1 ♀ (RN0181): Elis, Olympia, 29 May 2011, Werner Arens leg., WA coll.; 1 ♀ (RN0131): 08 Jun 2011, Werner Arens leg., WA coll.; 1 ♀ (RN0182): Zacharo, Lake Kaiapha, 30 May 2011, Werner Arens leg., WA coll.; 1 ♀ (RN0096): Zacharo, Neochori, 30 May 2011, Werner Arens leg., RN coll.; 1 ♀ (RN0175): Ionian Islands, Cephalonia, 03 Sep 1992, Peter Hartmann leg., ZSM coll.; 1 ♀ (RN0127): Messenia, Avia near Kalamata, 01 Jun 2011, Werner Arens leg., WA coll.; 3 ♀ (RN0133, RN0185, RN0186): 02 Jun 2011, Werner Arens leg., WA coll.; 1 ♀ (RN0176): Kalamata, 1997, Werner Arens leg., ZSM coll.; 1 ♀ (RN0184): Mavromati, Ithome mountain, 01 Jun 2011, Werner Arens leg., WA coll.; 1 ♀ (RN0177): 1997, Werner Arens leg., ZSM coll.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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