166 lines
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166 lines
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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e10095" ID-PMC="PMC5018111" ID-Pensoft-Pub="1314-2828-4-10095" ID-PubMed="27660520" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e10095" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="A new spider species, Heserstoevi sp. nov., from Turkmenistan (Araneae: Gnaphosidae)" checkinTime="1472674796337" checkinUser="pensoft" docAuthor="Deltshev, Christo" docDate="2016" docId="09F9333A92A3B03EC963E9CE06A6F8AC" docLanguage="en" docName="BiodivDatJour 4: e10095" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e10095" docTitle="Heser stoevi Deltshev, sp. n." docType="treatment" docVersion="3" lastPageNumber="10095" masterDocId="FFCAFFE8FF902330FFA3FFD41436C77E" masterDocTitle="A new spider species, Heserstoevi sp. nov., from Turkmenistan (Araneae: Gnaphosidae)" masterLastPageNumber="10095" masterPageNumber="10095" pageNumber="10095" updateTime="1668126770021" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new spider species, Heserstoevi sp. nov., from Turkmenistan (Araneae: Gnaphosidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Deltshev, Christo</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Biodiversity Data Journal</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>10095</mods:start>
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<mods:end>10095</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/BDJ.4.e10095</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.4.e10095</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2828-4-10095</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:09F9333A92A3B03EC963E9CE06A6F8AC" httpUri="http://treatment.plazi.org/id/09F9333A92A3B03EC963E9CE06A6F8AC" lastPageNumber="10095" pageId="0" pageNumber="10095">
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<subSubSection pageId="0" pageNumber="10095" type="nomenclature">
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<paragraph pageId="0" pageNumber="10095">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:E4D7D5A0-D649-4F5E-9360-D0488D73EEE8" authority="Deltshev" class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser stoevi" order="Araneae" pageId="0" pageNumber="10095" phylum="Arthropoda" rank="species" species="stoevi">Heser stoevi Deltshev</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="10095">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="10095" type="materials_examined">
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<paragraph pageId="0" pageNumber="10095">Materials</paragraph>
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<paragraph pageId="0" pageNumber="10095">
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<materialsCitation collectingDate="2015-05-25" collectingMethod="hand collecting" collectorName="P. Stoev" country="Turkmenistan" location="v. Hojeypil, around the Dinosaurs tracks Site" pageId="0" pageNumber="10095" specimenCount="1" specimenCount-male="1" typeStatus="Holotype">
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Type status:
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<typeStatus pageId="0" pageNumber="10095">Holotype</typeStatus>
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. Occurrence: recordedBy:
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<collectorName pageId="0" pageNumber="10095">P. Stoev</collectorName>
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; individualCount:
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<specimenCount pageId="0" pageNumber="10095">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="10095">male</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="10095">adult</specimenType>
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; Taxon: scientificName: Heserstoevi; family: Gnaphosidae; taxonRank: species; scientificNameAuthorship: Deltshev; Location: continent: Asia; country:
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<collectingCountry pageId="0" pageNumber="10095">Turkmenistan</collectingCountry>
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; stateProvince: Lebap; county: Koytendag District; locality:
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<location LSID="urn:lsid:plazi:treatment:09F9333A92A3B03EC963E9CE06A6F8AC:904D06A85A734B4DE1F8EA469CFC0937" country="Turkmenistan" name="v. Hojeypil, around the Dinosaurs tracks Site" pageId="0" pageNumber="10095">v. Hojeypil, around the Dinosaurs tracks Site</location>
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; verbatimElevation:
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<elevation pageId="0" pageNumber="10095">1150</elevation>
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; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="10095" value="37 56.443">37 56.443</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="10095" value="66 37.597">66 37.597</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="10095">Christo Deltshev</determinerName>
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; dateIdentified: 2015; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="10095">hand collecting</collectingMethod>
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; eventDate:
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<collectingDate pageId="0" pageNumber="10095" value="2015-05-25">05/25/2015</collectingDate>
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; habitat: under stones; Record Level: institutionID: NMNHS; collectionID: Archnida; basisOfRecord: PreservedSpecimen
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</materialsCitation>
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<materialsCitation collectingDate="2015-05-25" collectingMethod="hand collecting" collectorName="P. Stoev" country="Turkmenistan" location="v. Hojeypil, around the Dinosaurs tracks Site" pageId="0" pageNumber="10095" specimenCount="1" specimenCount-female="1" typeStatus="Paratype">
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Type status:
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<typeStatus pageId="0" pageNumber="10095">Paratype</typeStatus>
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. Occurrence: recordedBy:
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<collectorName pageId="0" pageNumber="10095">P. Stoev</collectorName>
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; individualCount:
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<specimenCount pageId="0" pageNumber="10095">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="10095">female</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="10095">adult</specimenType>
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; Taxon: scientificName: Heserstoevi; family: Gnaphosidae; taxonRank: species; scientificNameAuthorship: Deltshev; Location: continent: Asia; country:
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<collectingCountry pageId="0" pageNumber="10095">Turkmenistan</collectingCountry>
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; stateProvince: Lebap; county: Koytendag District; locality:
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<location LSID="urn:lsid:plazi:treatment:09F9333A92A3B03EC963E9CE06A6F8AC:32E8FDAD5594C142F221C1EBF93D9F6C" country="Turkmenistan" name="v. Hojeypil, around the Dinosaurs tracks Site" pageId="0" pageNumber="10095">v. Hojeypil, around the Dinosaurs tracks Site</location>
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; verbatimElevation:
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<elevation pageId="0" pageNumber="10095">1150</elevation>
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; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="10095" value="37 56.443">37 56.443</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="10095" value="66 37.597">66 37.597</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="10095">Christo Deltshev</determinerName>
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; dateIdentified: 2015; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="10095">hand collecting</collectingMethod>
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; eventDate:
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<collectingDate pageId="0" pageNumber="10095" value="2015-05-25">05/25/2015</collectingDate>
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; habitat: under stones; Record Level: institutionID: NMNHS; collectionID: Archnida; basisOfRecord: PreservedSpecimen
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</materialsCitation>
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<materialsCitation collectingDate="2015-05-28" collectingMethod="hand collecting" collectorName="P. Stoev" country="Turkmenistan" location="v. Garlyk, around cave Gulshirin (= Geofizicheskaya)" pageId="0" pageNumber="10095" specimenCount="1" specimenCount-female="1" typeStatus="Paratype">
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Type status:
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<typeStatus pageId="0" pageNumber="10095">Paratype</typeStatus>
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. Occurrence: recordedBy:
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<collectorName pageId="0" pageNumber="10095">P. Stoev</collectorName>
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; individualCount:
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<specimenCount pageId="0" pageNumber="10095">1</specimenCount>
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; sex:
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<specimenType pageId="0" pageNumber="10095">female</specimenType>
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; lifeStage:
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<specimenType pageId="0" pageNumber="10095">adult</specimenType>
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; Taxon: scientificName: Heserstoevi; family: Gnaphosidae; taxonRank: species; scientificNameAuthorship: Deltshev; Location: continent: Asia; country:
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<collectingCountry pageId="0" pageNumber="10095">Turkmenistan</collectingCountry>
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; stateProvince: Lebap; county: Koytendag District; locality:
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<location LSID="urn:lsid:plazi:treatment:09F9333A92A3B03EC963E9CE06A6F8AC:0B2C09AFED35D638BDA12752FBB5A94B" country="Turkmenistan" name="v. Garlyk, around cave Gulshirin (= Geofizicheskaya)" pageId="0" pageNumber="10095">v. Garlyk, around cave Gulshirin (= Geofizicheskaya)</location>
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; verbatimElevation:
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<elevation pageId="0" pageNumber="10095">860</elevation>
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; decimalLatitude:
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<geoCoordinate orientation="latitude" pageId="0" pageNumber="10095" value="37 40.394">37 40.394</geoCoordinate>
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; decimalLongitude:
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<geoCoordinate orientation="longitude" pageId="0" pageNumber="10095" value="66 23.698">66 23.698</geoCoordinate>
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; Identification: identifiedBy:
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<determinerName pageId="0" pageNumber="10095">Christo Deltshev</determinerName>
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; dateIdentified: 2015; Event: samplingProtocol:
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<collectingMethod pageId="0" pageNumber="10095">hand collecting</collectingMethod>
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; eventDate:
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<collectingDate pageId="0" pageNumber="10095" value="2015-05-28">05/28/2015</collectingDate>
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; habitat: under stones; Record Level: institutionID: NMNHS; collectionID: Archnida; basisOfRecord: PreservedSpecimen
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="10095" type="description">
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<paragraph pageId="0" pageNumber="10095">Description</paragraph>
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<paragraph pageId="0" pageNumber="10095">Male holotype: Total length 5.25; prosoma, length 2.55, width 1.80; sternum length 1.43, width 1.05; chelicerae, length 0.75, width 0.30; opisthosoma, length 5.25. Carapace uniformly brown, with a fovea in the posterior half (Fig. 1a, c). All eyes subequal, AME circular, separated from each other by their own diameter, ALE oval, touching AME. PME oval to subtriangular, touching, larger than AME. PLE oval, slightly smaller than ALE, separated from PME by more of one PLE diameter. Clypeus vertical, large one diameter of AME. Chelicerae brown, with a few scattered thin setae on anterior surface, promarginal rim with three very small teeth spaced closely to fang base, retromarginal rim spineless. Sternum smooth, yellow brown, shield-shaped with a thin border (Fig. 1b). Abdomen grey with frontal row of curved hairs and a scutum covering about 10% of abdominal dorsal surface area (Fig. 1a, c). Legs yellowish to yellow-brown (Fig. 1a, c), leg formula 4123, measurements as in Table 1.</paragraph>
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<paragraph pageId="0" pageNumber="10095">
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Male palp (Fig. 2a, b, c, Fig. 5a, c, e): Distinctive retrolateral tibial apophysis is not presented, hence distal part of tibia retrolateral is strongly chitinized and dark colored. The embolus is slender, basally prolaterally inserted, extends across the broad and flat conductor (sensu
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<bibRefCitation author="Bosselaers, Jan" journalOrPublisher="ZooKeys" pageId="0" pageNumber="10095" pagination="1 - 11" title="A new species of Heser (Araneae, Gnaphosidae) from the south of India" volume="73" year="2010">Bosselaers 2010</bibRefCitation>
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), circling more than half of the tegulum. Median apophysis large, hook-shaped.
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</paragraph>
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<paragraph pageId="0" pageNumber="10095">Female paratype: Total length 6.00; prosoma, length 2.63, width 1.80; sternum length 1.50, width 1.05; opisthosoma, length 3.38 (Fig. 3a, c). All characters as described for male. Leg measurements as in Table 2.</paragraph>
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<paragraph pageId="0" pageNumber="10095">Epigyne and vulva (Figs 4a, b, 6a, b): characterized by well-developed copulatory openings, leading to coiled insemination ducts, connected with coiled spermathecae.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="10095" type="diagnosis">
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<paragraph pageId="0" pageNumber="10095">Diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="10095">
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The somatic characters of the new species correspond to those of the genus
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Heser" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heser" order="Araneae" pageId="0" pageNumber="10095" phylum="Arthropoda" rank="genus">Heser</taxonomicName>
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, but the genitalia are distinctive and separate well the new species from all other congeners. Morphologically, the new species resembles
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<taxonomicName lsidName="H. aradensis" pageId="0" pageNumber="10095" rank="species" species="aradensis">H. aradensis</taxonomicName>
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(Fig. 1b, d) but the male has longer embolus, and smaller and flattened conductor (sensu
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<bibRefCitation author="Bosselaers, Jan" journalOrPublisher="ZooKeys" pageId="0" pageNumber="10095" pagination="1 - 11" title="A new species of Heser (Araneae, Gnaphosidae) from the south of India" volume="73" year="2010">Bosselaers 2010</bibRefCitation>
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), as well as lacks distinctive retrolateral tibial apophysis (Fig. 2a, c, e). Female has distinctive epygine and vulva, characterized by well-defined copulatory openings leading to coiled insemination ducts, connected with coiled spermathecae (see Figs 4c, d, 6c, d for comparison).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="10095" type="etymology">
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<paragraph pageId="0" pageNumber="10095">Etymology</paragraph>
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<paragraph pageId="0" pageNumber="10095">Named in honour of the Bulgarian zoologist Pavel Stoev, who collected the species; name in genitive case.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="10095" type="distribution">
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<paragraph pageId="0" pageNumber="10095">Distribution</paragraph>
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<paragraph pageId="0" pageNumber="10095">The species is hitherto known only from two closely situated localities in the western slope of Koytentag Mts, southeastern Turkmenistan (Fig. 7).</paragraph>
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</subSubSection>
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</treatment>
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</document>
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