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<mods:title id="34ACEC1CA9193E1FDA321205B10B75AE">Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species</mods:title>
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<mods:namePart id="108FD335EA5CC1F26D11C46F6EE87360">Moctezuma, Victor</mods:namePart>
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<mods:affiliation id="332672BC402B8A915357C9B66FB92CC0">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:affiliation id="89BEAF9933189DE0527DC074BBF696AC">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:title id="74DEC8CC519C0721A266367D2882CCA5">European Journal of Taxonomy</mods:title>
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<subSubSection id="C379656FFFD0FFEFFD9740BDBC8638EA" box="[575,1012,694,721]" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="8BDC36E4FFD0FFEFFD9740BDBC8638EA" blockId="46.[575,1012,694,758]" box="[575,1012,694,721]" pageId="46" pageNumber="47">
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<heading id="D0948188FFD0FFEFFD9740BDBC8638EA" box="[575,1012,694,721]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
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<taxonomicName id="4C634D67FFD0FFEFFD9740BDBC8638EA" ID-CoL="4FSK2" authority="Edmonds, 2006" authorityName="Edmonds" authorityYear="2006" box="[575,1012,694,721]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="zapotecus">
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<emphasis id="B917EAF6FFD0FFEFFD9740BDBC4638EB" bold="true" box="[575,820,694,721]" italics="true" pageId="46" pageNumber="47">Phanaeus zapotecus</emphasis>
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<bibRefCitation id="EFF24B15FFD0FFEFFC9440BDBC8638EA" author="Edmonds W. D." box="[828,1012,694,720]" pageId="46" pageNumber="47" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds, 2006</bibRefCitation>
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<subSubSection id="C379656FFFD0FFEFFDC240D0BCBB38CF" box="[618,969,731,758]" pageId="46" pageNumber="47" type="description">
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<paragraph id="8BDC36E4FFD0FFEFFDC240D0BCBB38CF" blockId="46.[575,1012,694,758]" box="[618,969,731,758]" pageId="46" pageNumber="47">
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<figureCitation id="13582A61FFD0FFEFFDC240D0BDB338CF" box="[618,705,731,757]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="46" pageNumber="47">Figs 1P</figureCitation>
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,
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<figureCitation id="13582A61FFD0FFEFFD6640D0BD8038CF" box="[718,754,731,757]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="46" pageNumber="47">2L</figureCitation>
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,
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<figureCitation id="13582A61FFD0FFEFFD5640D0BC3838CC" box="[766,842,731,758]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="48.[189,232,1877,1903]" captionStartId-1="50.[189,232,1890,1916]" captionTargetBox-0="[228,1363,265,1833]" captionTargetBox-1="[193,1392,927,1854]" captionTargetId-0="figure-40@48.[189,1399,265,1833]" captionTargetId-1="figure-235@50.[189,1398,924,1857]" captionTargetPageId-0="48" captionTargetPageId-1="50" captionText-0="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." captionText-1="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi-0="http://doi.org/10.5281/zenodo.4836794" figureDoi-1="http://doi.org/10.5281/zenodo.4836796" httpUri-0="https://zenodo.org/record/4836794/files/figure.png" httpUri-1="https://zenodo.org/record/4836796/files/figure.png" pageId="46" pageNumber="47">14–15</figureCitation>
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,
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<figureCitation id="13582A61FFD0FFEFFCFE40D0BCF938CF" box="[854,907,731,757]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="46" pageNumber="47">18Q</figureCitation>
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,
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<figureCitation id="13582A61FFD0FFEFFC3F40D0BCBB38CF" box="[919,969,731,757]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="46" pageNumber="47">19Q</figureCitation>
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</paragraph>
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<paragraph id="8BDC36E4FFD0FFEFFF154115BCB93903" blockId="46.[189,1383,798,861]" box="[189,971,798,825]" pageId="46" pageNumber="47">
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<taxonomicName id="4C634D67FFD0FFEFFF154115BDB53902" ID-CoL="4FSK2" authority="Edmonds, 2006: 31 - 32" authorityName="Edmonds" authorityPageNumber="31 - 32" authorityYear="2006" box="[189,711,798,824]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="zapotecus">
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<emphasis id="B917EAF6FFD0FFEFFF154115BEDE3902" box="[189,428,798,824]" italics="true" pageId="46" pageNumber="47">Phanaeus zapotecus</emphasis>
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<bibRefCitation id="EFF24B15FFD0FFEFFE1B4115BDB53902" author="Edmonds W. D." box="[435,711,798,824]" pageId="46" pageNumber="47" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds, 2006: 31–32</bibRefCitation>
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, 35–37, figs 1, 3, 5, 7.
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</paragraph>
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|
<taxonomicName id="4C634D67FFD0FFEFFF154148BC633967" ID-CoL="856BD" authority="Kohlmann et al., 2018: 67 - 70" authorityName="Kohlmann" authorityPageNumber="67 - 70" authorityYear="2018" box="[189,785,835,861]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF154148BED53967" box="[189,423,835,861]" italics="true" pageId="46" pageNumber="47">Phanaeus dionysius</emphasis>
|
|||
|
|
<bibRefCitation id="EFF24B15FFD0FFEFFE064148BC633967" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[430,785,835,861]" pageId="46" pageNumber="47" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFD994148BD173967" box="[561,613,835,861]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
|
|||
|
|
., 2018: 67–70
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
, 76, 78, 82, 88–89, figs 1b, 3a–b, d, 4–7.
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFB5E4149BA153967" bold="true" box="[1270,1383,834,861]" pageId="46" pageNumber="47">Syn. nov.</emphasis>
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF15418DBCC639FF" blockId="46.[189,1399,901,1038]" pageId="46" pageNumber="47">
|
|||
|
|
<treatmentCitationGroup id="AB7311CAFFD0FFEFFF15418DBCC639FF" pageId="46" pageNumber="47">
|
|||
|
|
<taxonomicName id="4C634D67FFD0FFEFFF15418DBEDD39A5" ID-CoL="4FSK2" authorityName="Edmonds" authorityYear="2006" box="[189,431,902,928]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF15418DBEDD39A5" box="[189,431,902,928]" italics="true" pageId="46" pageNumber="47">Phanaeus zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
–
|
|||
|
|
<treatmentCitation id="0AC210F5FFD0FFEFFE7A418DBC65399A" author="Edmonds W. D. & Zidek J." box="[466,791,902,928]" page="1" pageId="46" pageNumber="47" year="2012">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD0FFEFFE7A418DBC65399A" author="Edmonds W. D. & Zidek J." box="[466,791,902,928]" pageId="46" pageNumber="47" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek 2012: 1</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
, 7–8. —
|
|||
|
|
<treatmentCitation id="0AC210F5FFD0FFEFFC25418DBA65399A" author="Moctezuma V. & Halffter G." box="[909,1303,901,928]" page="52" pageId="46" pageNumber="47" year="2017">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD0FFEFFC25418DBA65399A" author="Moctezuma V. & Halffter G." box="[909,1303,901,928]" pageId="46" pageNumber="47" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter 2017: 52</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
, 54–55, fig. 23. — Moctezuma
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFE5241A0BD5C39FE" box="[506,558,938,964]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
|
|||
|
|
. 2017: 114, 130, 132; 2019: 253.
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF1541C4BA2B39D0" blockId="46.[189,1399,901,1038]" box="[189,1369,975,1002]" pageId="46" pageNumber="47">
|
|||
|
|
<treatmentCitationGroup id="AB7311CAFFD0FFEFFF1541C4BA2B39D0" box="[189,1369,975,1002]" pageId="46" pageNumber="47">
|
|||
|
|
<taxonomicName id="4C634D67FFD0FFEFFF1541C4BD0A39D3" ID-CoL="4FSK2" authorityName="Edmonds" authorityYear="2006" box="[189,632,975,1002]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="zapotecus" subGenus="Notiophanaeus">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF1541C4BE4339D3" box="[189,305,975,1001]" italics="true" pageId="46" pageNumber="47">Phanaeus</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFEEA41C4BE8639D3" box="[322,500,975,1001]" italics="true" pageId="46" pageNumber="47">Notiophanaeus</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFDAC41DBBD0A39D3" box="[516,632,976,1001]" italics="true" pageId="46" pageNumber="47">zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
–
|
|||
|
|
<treatmentCitation id="0AC210F5FFD0FFEFFD3E41C4BCA239D3" author="Edmonds W. D. & Zidek J." box="[662,976,975,1001]" page="3" pageId="46" pageNumber="47" year="2012">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD0FFEFFD3E41C4BCA239D3" author="Edmonds W. D. & Zidek J." box="[662,976,975,1001]" pageId="46" pageNumber="47" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek 2012: 3</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
, 13, figs 139, 141, 143, 148–151.
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF1541FFBC833E34" blockId="46.[189,1399,901,1038]" box="[189,1009,1012,1038]" pageId="46" pageNumber="47">
|
|||
|
|
<treatmentCitationGroup id="AB7311CAFFD0FFEFFF1541FFBC833E34" box="[189,1009,1012,1038]" pageId="46" pageNumber="47">
|
|||
|
|
<taxonomicName id="4C634D67FFD0FFEFFF1541FFBD013E34" ID-CoL="856BD" box="[189,627,1012,1038]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="dionysius" subGenus="Notiophanaeus">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF1541FFBE433E34" box="[189,305,1012,1038]" italics="true" pageId="46" pageNumber="47">Phanaeus</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFEEA41FFBE863E34" box="[322,500,1012,1038]" italics="true" pageId="46" pageNumber="47">Notiophanaeus</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFDAC41FFBD013E34" box="[516,627,1012,1038]" italics="true" pageId="46" pageNumber="47">dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
–
|
|||
|
|
<treatmentCitation id="0AC210F5FFD0FFEFFD3941FFBCB23E34" LSID="urn:lsid:zoobank.org:act:B4F5DBB5-6802-44F0-87FF-199895A96C80" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[657,960,1012,1038]" httpUri="http://treatment.plazi.org/id/355A9EEBAD9D2DDA6EACCCBF6A512440" page="68" pageId="46" pageNumber="47" year="2018">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD0FFEFFD3941FFBCB23E34" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[657,960,1012,1038]" pageId="46" pageNumber="47" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFCBC41FEBC353E34" box="[788,839,1012,1038]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
|
|||
|
|
. 2018: 68
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
, 70.
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF15463CBBE83E68" blockId="46.[189,1178,1079,1142]" box="[189,1178,1079,1106]" pageId="46" pageNumber="47">
|
|||
|
|
<treatmentCitationGroup id="AB7311CAFFD0FFEFFF15463CBBE83E68" box="[189,1178,1079,1106]" pageId="46" pageNumber="47">
|
|||
|
|
Non
|
|||
|
|
<taxonomicName id="4C634D67FFD0FFEFFF5F463CBE963E6B" ID-CoL="4FSJL" baseAuthorityName="Edmonds" baseAuthorityYear="1994" box="[247,484,1079,1105]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="endymion">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF5F463CBE963E6B" box="[247,484,1079,1105]" italics="true" pageId="46" pageNumber="47">Phanaeus endymion</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
–
|
|||
|
|
<treatmentCitation id="0AC210F5FFD0FFEFFDA9463CBDAD3E6B" author="Edmonds W. D." box="[513,735,1079,1105]" page="44" pageId="46" pageNumber="47" year="1994">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD0FFEFFDA9463CBDAD3E6B" author="Edmonds W. D." box="[513,735,1079,1105]" pageId="46" pageNumber="47" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds 1994: 44</bibRefCitation>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(referred to as “
|
|||
|
|
<collectingRegion id="49A7F806FFD0FFEFFC34463CBC8B3E6B" box="[924,1017,1079,1105]" country="Mexico" name="Oaxaca" pageId="46" pageNumber="47">Oaxaca</collectingRegion>
|
|||
|
|
” population).
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF154657BB1A3E4C" blockId="46.[189,1178,1079,1142]" box="[189,1128,1115,1142]" pageId="46" pageNumber="47">
|
|||
|
|
<treatmentCitationGroup id="AB7311CAFFD0FFEFFF154657BB1A3E4C" box="[189,1128,1115,1142]" pageId="46" pageNumber="47">
|
|||
|
|
Non
|
|||
|
|
<taxonomicName id="4C634D67FFD0FFEFFF5F4650BD453E4C" ID-CoL="63ZKT" authority="" authorityName="Edmonds" authorityYear="1994" box="[247,567,1115,1142]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="subGenus" subGenus="Notiophanaeus">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF5F4650BE193E4F" box="[247,363,1115,1141]" italics="true" pageId="46" pageNumber="47">Phanaeus</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFED44650BD5C3E4F" box="[380,558,1115,1141]" italics="true" pageId="46" pageNumber="47">Notiophanaeus</emphasis>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
“
|
|||
|
|
<collectingRegion id="49A7F806FFD0FFEFFDE14657BDD43E4C" box="[585,678,1116,1142]" country="Mexico" name="Oaxaca" pageId="46" pageNumber="47">Oaxaca</collectingRegion>
|
|||
|
|
”
|
|||
|
|
<taxonomicName id="4C634D67FFD0FFEFFD104650BC583E4F" ID-CoL="4FSJL" baseAuthorityName="Edmonds" baseAuthorityYear="1994" box="[696,810,1115,1141]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="endymion">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFD104650BC583E4F" box="[696,810,1115,1141]" italics="true" pageId="46" pageNumber="47">endymion</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
–
|
|||
|
|
<treatmentCitation id="0AC210F5FFD0FFEFFCEF4650BB1A3E4C" box="[839,1128,1115,1142]" pageId="46" pageNumber="47">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD0FFEFFCEF4650BC8E3E4C" author="Edmonds W. D." box="[839,1020,1115,1142]" pageId="46" pageNumber="47" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds 1994</bibRefCitation>
|
|||
|
|
: fig. 221.
|
|||
|
|
</treatmentCitation>
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C379656FFFD0FFEEFF154694BC173B68" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="materials_examined">
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF154694BCE43E83" blockId="46.[189,918,1182,1209]" box="[189,918,1182,1209]" pageId="46" pageNumber="47">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF154694BD9A3E82" bold="true" box="[189,744,1182,1209]" pageId="46" pageNumber="47">
|
|||
|
|
Type material of
|
|||
|
|
<taxonomicName id="4C634D67FFD0FFEFFE3B4695BDFA3E83" authorityName="Edmonds" authorityYear="2006" box="[403,648,1182,1209]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFE3B4695BDFA3E83" bold="true" box="[403,648,1182,1209]" italics="true" pageId="46" pageNumber="47">Phanaeus zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
revised
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFD514694BC4F3E83" box="[761,829,1183,1209]" pageId="46" pageNumber="47" type="male">3 ♂♂</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFCE24695BCF83E83" box="[842,906,1182,1209]" pageId="46" pageNumber="47" type="female">5 ♀♀</specimenCount>
|
|||
|
|
)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF4246C4BDB03ED3" blockId="46.[189,1292,1230,1294]" box="[234,706,1230,1257]" pageId="46" pageNumber="47">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF4246C4BE2E3ED3" bold="true" box="[234,348,1231,1257]" pageId="46" pageNumber="47">
|
|||
|
|
<typeStatus id="54D88846FFD0FFEFFF4246C4BE2E3ED3" box="[234,348,1231,1257]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
(not studied, temporarily lost)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF1546F8BA7E3F37" blockId="46.[189,1292,1230,1294]" box="[189,1292,1267,1294]" pageId="46" pageNumber="47">
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFF1546F8BA723F37" ID-GBIF-Occurrence="3093980542" box="[189,1280,1267,1294]" collectionCode="IEXA" country="Mexico" location="Oaxaca" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
|
|||
|
|
<collectingCountry id="F3747674FFD0FFEFFF1546F8BE433F37" box="[189,305,1267,1293]" name="Mexico" pageId="46" pageNumber="47">MEXICO</collectingCountry>
|
|||
|
|
–
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFEE646F8BEDF3F37" bold="true" box="[334,429,1267,1293]" pageId="46" pageNumber="47">
|
|||
|
|
<collectingRegion id="49A7F806FFD0FFEFFEE646F8BEDF3F37" box="[334,429,1267,1293]" country="Mexico" name="Oaxaca" pageId="46" pageNumber="47">Oaxaca</collectingRegion>
|
|||
|
|
</emphasis>
|
|||
|
|
•
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFE6D46FFBEAF3F34" box="[453,477,1268,1294]" pageId="46" pageNumber="47" type="male">♂</specimenCount>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFF24B15FFD0FFEFFE4246F8BDB93F37" author="Edmonds W. D." box="[490,715,1267,1293]" pageId="46" pageNumber="47" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds 2006: 32</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<locationDeviation id="9A6E2FC6FFD0FFEFFD7F46F8BBDF3F34" box="[727,1197,1267,1294]" pageId="46" pageNumber="47">
|
|||
|
|
<quantity id="4C9B9B01FFD0FFEFFD7F46F8BC603F37" box="[727,786,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="46" pageNumber="47" unit="km" value="8.0">8 km</quantity>
|
|||
|
|
south of San Miguel Sola de Vega
|
|||
|
|
</locationDeviation>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFB1046F8BA723F37" box="[1208,1280,1267,1293]" pageId="46" pageNumber="47">IEXA</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
?
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF42473DBE153F6A" blockId="46.[189,1399,1334,1508]" box="[234,359,1334,1360]" pageId="46" pageNumber="47">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF42473DBE153F6A" bold="true" box="[234,359,1334,1360]" pageId="46" pageNumber="47">
|
|||
|
|
<typeStatus id="54D88846FFD0FFEFFF42473DBE153F6A" box="[234,359,1334,1360]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF154750BC413FD9" blockId="46.[189,1399,1334,1508]" pageId="46" pageNumber="47">
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFF154750BD403FA0" ID-GBIF-Occurrence="3093980526" collectedFrom="Pine oak forest" collectingDate="2005-09-07" collectingDateMax="2005-09-17" collectingDateMin="2005-09-07" collectingMethod="Mushroom-baited trap" collectionCode="CEMT" collectorName="D. Curoe" country="Mexico" elevation="1850" location="Oaxaca" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
|
|||
|
|
<collectingCountry id="F3747674FFD0FFEFFF154750BE433F4F" box="[189,305,1371,1397]" name="Mexico" pageId="46" pageNumber="47">MEXICO</collectingCountry>
|
|||
|
|
–
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFEE44750BED93F4F" bold="true" box="[332,427,1371,1397]" pageId="46" pageNumber="47">
|
|||
|
|
<collectingRegion id="49A7F806FFD0FFEFFEE44750BED93F4F" box="[332,427,1371,1397]" country="Mexico" name="Oaxaca" pageId="46" pageNumber="47">Oaxaca</collectingRegion>
|
|||
|
|
</emphasis>
|
|||
|
|
•
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFE6B4750BE833F4F" box="[451,497,1371,1397]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFE554750BD5B3F4F" box="[509,553,1371,1397]" pageId="46" pageNumber="47" type="female">1 ♀</specimenCount>
|
|||
|
|
; “
|
|||
|
|
<locationDeviation id="9A6E2FC6FFD0FFEFFDEC4750BC533F4F" box="[580,801,1371,1397]" pageId="46" pageNumber="47">
|
|||
|
|
<quantity id="4C9B9B01FFD0FFEFFDEC4750BD0D3F4F" box="[580,639,1371,1397]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="46" pageNumber="47" unit="km" value="8.0">8 km</quantity>
|
|||
|
|
Sola de Vega
|
|||
|
|
</locationDeviation>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9B9B01FFD0FFEFFC844750BCF43F4F" box="[812,902,1371,1398]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="46" pageNumber="47" unit="m" value="1850.0">
|
|||
|
|
<elevation id="004ED1D7FFD0FFEFFC844750BCF43F4F" box="[812,902,1371,1398]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="46" pageNumber="47" unit="m" value="1850.0">1850 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="FFDD1024FFD0FFEFFC3A4750BB633F4C" box="[914,1041,1371,1398]" pageId="46" pageNumber="47" value="2005-09-07" valueMax="2005-09-17" valueMin="2005-09-07">
|
|||
|
|
<collectingDate id="EF99E9CCFFD0FFEFFC3A4750BB633F4C" box="[914,1041,1371,1398]" pageId="46" pageNumber="47" value="2005-09-07" valueMax="2005-09-17" valueMin="2005-09-07">7–17 ix 05</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectedFrom id="387F7519FFD0FFEFFBB44750BBBD3F4F" box="[1052,1231,1371,1397]" pageId="46" pageNumber="47">Pine oak forest</collectedFrom>
|
|||
|
|
,
|
|||
|
|
<collectorName id="26965332FFD0FFEFFB714750BA373F4F" box="[1241,1349,1371,1397]" pageId="46" pageNumber="47">D. Curoe</collectorName>
|
|||
|
|
col.
|
|||
|
|
<collectingMethod id="52224EF3FFD0FFEFFF15478BBEB83FA0" box="[189,458,1408,1434]" pageId="46" pageNumber="47">Mushroom-baited trap</collectingMethod>
|
|||
|
|
”;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFE49478BBD403FA0" box="[481,562,1408,1434]" collectionName="CEMT" pageId="46" pageNumber="47">CEMT</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
•
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFDE3478BBBAA3FA0" ID-GBIF-Occurrence="3093980506" box="[587,1240,1408,1434]" collectedFrom="Pine oak forest" collectingDate="2005-09-07" collectingDateMax="2005-09-17" collectingDateMin="2005-09-07" collectingMethod="Mushroom-baited trap" collectionCode="TAMU" collectorName="D. Curoe" country="Mexico" elevation="1850" location="8 km Sola de Vega" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFDE3478BBD0B3FA0" box="[587,633,1408,1434]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFD2E478BBDC13FA0" box="[646,691,1408,1434]" pageId="46" pageNumber="47" type="female">1 ♀</specimenCount>
|
|||
|
|
; same collection data as for preceding;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFB2C478BBBAA3FA0" box="[1156,1240,1408,1434]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="46" pageNumber="47">TAMU</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
•
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFB59478BBDBB3F84" ID-GBIF-Occurrence="3093980570" collectedFrom="Pine oak forest" collectingDate="2005-09-07" collectingDateMax="2005-09-17" collectingDateMin="2005-09-07" collectingMethod="Mushroom-baited trap" collectionCode="VMC" collectorName="D. Curoe" country="Mexico" elevation="1850" location="8 km Sola de Vega" pageId="46" pageNumber="47" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFB59478BBA6D3FA0" box="[1265,1311,1408,1434]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFA84478BBA1D3FA0" box="[1324,1391,1408,1434]" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
|
|||
|
|
; same collection data as for preceding;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFD2E47AFBDBB3F84" box="[646,713,1444,1470]" pageId="46" pageNumber="47">VMC</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
•
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFD4C47AFBC5D3FD9" ID-GBIF-Occurrence="3093980362" collectedFrom="Pine oak forest" collectingDate="2005-07-04" collectingDateMax="2005-07-09" collectingDateMin="2005-07-04" collectingMethod="Mushroom-baited trap" collectionCode="TAMU" collectorName="D. Curoe" country="Mexico" elevation="1850" location="8 km Sola de Vega" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca" typeStatus="paratype">
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFD4C47AFBC603F85" box="[740,786,1444,1471]" pageId="46" pageNumber="47" type="female">1 ♀</specimenCount>
|
|||
|
|
; “
|
|||
|
|
<locationDeviation id="9A6E2FC6FFD0FFEFFC8647AFBB633F85" box="[814,1041,1444,1471]" pageId="46" pageNumber="47">
|
|||
|
|
<quantity id="4C9B9B01FFD0FFEFFC8647AFBC183F84" box="[814,874,1444,1470]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="46" pageNumber="47" unit="km" value="8.0">8 km</quantity>
|
|||
|
|
Sola de Vega
|
|||
|
|
</locationDeviation>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9B9B01FFD0FFEFFBB547AFBB0B3F85" box="[1053,1145,1444,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="46" pageNumber="47" unit="m" value="1850.0">
|
|||
|
|
<elevation id="004ED1D7FFD0FFEFFBB547AFBB0B3F85" box="[1053,1145,1444,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="46" pageNumber="47" unit="m" value="1850.0">1850 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="FFDD1024FFD0FFEFFB2F47AFBA713F85" box="[1159,1283,1444,1471]" pageId="46" pageNumber="47" value="2005-07-04" valueMax="2005-07-09" valueMin="2005-07-04">
|
|||
|
|
<collectingDate id="EF99E9CCFFD0FFEFFB2F47AFBA713F85" box="[1159,1283,1444,1471]" pageId="46" pageNumber="47" value="2005-07-04" valueMax="2005-07-09" valueMin="2005-07-04">4–9 vii 05</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectedFrom id="387F7519FFD0FFEFFAB847AFBE713FD9" pageId="46" pageNumber="47">Pine oak forest</collectedFrom>
|
|||
|
|
,
|
|||
|
|
<collectorName id="26965332FFD0FFEFFEA647C1BE083FD9" box="[270,378,1481,1508]" pageId="46" pageNumber="47">D. Curoe</collectorName>
|
|||
|
|
col.
|
|||
|
|
<collectingMethod id="52224EF3FFD0FFEFFE1C47C2BDB23FD9" box="[436,704,1481,1507]" pageId="46" pageNumber="47">Mushroom-baited trap</collectingMethod>
|
|||
|
|
”;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFD7F47C2BC5D3FD9" box="[727,815,1481,1507]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="46" pageNumber="47">TAMU</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF154407BCE13C1D" blockId="46.[189,915,1547,1575]" box="[189,915,1547,1575]" pageId="46" pageNumber="47">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF154407BD973C1C" bold="true" box="[189,741,1547,1574]" pageId="46" pageNumber="47">
|
|||
|
|
Type material of
|
|||
|
|
<taxonomicName id="4C634D67FFD0FFEFFE3B4400BDF73C1C" box="[403,645,1547,1574]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFE3B4400BDF73C1C" bold="true" box="[403,645,1547,1574]" italics="true" pageId="46" pageNumber="47">Phanaeus dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
revised
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFD5D4407BC483C1D" box="[757,826,1548,1575]" pageId="46" pageNumber="47" type="male">2 ♂♂</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFCEF4407BCF53C1D" box="[839,903,1548,1575]" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
|
|||
|
|
)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF424437BE2E3C6C" blockId="46.[189,1398,1596,1696]" box="[234,348,1596,1622]" pageId="46" pageNumber="47">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF424437BE2E3C6C" bold="true" box="[234,348,1596,1622]" pageId="46" pageNumber="47">
|
|||
|
|
<typeStatus id="54D88846FFD0FFEFFF424437BE2E3C6C" box="[234,348,1596,1622]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF15446ABB013C9A" blockId="46.[189,1398,1596,1696]" pageId="46" pageNumber="47">
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFF15446ABB013C9A" ID-GBIF-Occurrence="3093980313" collectedFrom="bosque de Encino" collectingDate="2017-06-23" collectionCode="IEXA" collectorName="Arenas A. Col." country="Mexico" elevation="1976" latitude="17.1651" location="La mesita San Pablo Etla" longLatPrecision="1" longitude="96.73858" municipality="La" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
|
|||
|
|
<collectingCountry id="F3747674FFD0FFEFFF15446ABE433C41" box="[189,305,1633,1659]" name="Mexico" pageId="46" pageNumber="47">MEXICO</collectingCountry>
|
|||
|
|
–
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFEF1446BBECA3C40" bold="true" box="[345,440,1632,1658]" pageId="46" pageNumber="47">
|
|||
|
|
<collectingRegion id="49A7F806FFD0FFEFFEF1446BBECA3C40" box="[345,440,1632,1658]" country="Mexico" name="Oaxaca" pageId="46" pageNumber="47">Oaxaca</collectingRegion>
|
|||
|
|
</emphasis>
|
|||
|
|
• “
|
|||
|
|
<location id="8EBC603FFFD0FFEFFE4E446ABC553C41" LSID="urn:lsid:plazi:treatment:03CA87F2FFD0FFF0FD9740BDBE763F02:8EBC603FFFD0FFEFFE4E446ABC553C41" box="[486,807,1633,1659]" country="Mexico" latitude="17.1651" longLatPrecision="1" longitude="96.73858" municipality="La" name="La mesita San Pablo Etla" pageId="46" pageNumber="47" stateProvince="Oaxaca">La mesita San Pablo Etla</location>
|
|||
|
|
,
|
|||
|
|
<date id="FFDD1024FFD0FFEFFC91446ABCC03C41" box="[825,946,1633,1659]" pageId="46" pageNumber="47" value="2017-06-23">
|
|||
|
|
<collectingDate id="EF99E9CCFFD0FFEFFC91446ABCC03C41" box="[825,946,1633,1659]" pageId="46" pageNumber="47" value="2017-06-23">23–VI–17</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
, coprotrampa,
|
|||
|
|
<date id="FFDD1024FFD0FFEFFBC4446ABBC03C41" box="[1132,1202,1633,1659]" pageId="46" pageNumber="47" value="1996-10">
|
|||
|
|
x-
|
|||
|
|
<geoCoordinate id="EE575023FFD0FFEFFB39446ABBC03C41" box="[1169,1202,1633,1659]" degrees="96" direction="west" minutes="44" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="18.91" value="-96.73858">96</geoCoordinate>
|
|||
|
|
</date>
|
|||
|
|
°44’18.91’’O, y-
|
|||
|
|
<geoCoordinate id="EE575023FFD0FFEFFF15448EBE1B3C9A" box="[189,361,1669,1696]" degrees="17" direction="north" minutes="9" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="54.36" value="17.1651">17°9’54.36’’N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<collectedFrom id="387F7519FFD0FFEFFEDE448EBD3B3CA5" box="[374,585,1669,1695]" pageId="46" pageNumber="47">bosque de Encino</collectedFrom>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9B9B01FFD0FFEFFDFD448EBDC23C9A" box="[597,688,1669,1696]" metricMagnitude="3" metricUnit="m" metricValue="1.9759999999999998" pageId="46" pageNumber="47" unit="m" value="1976.0">
|
|||
|
|
<elevation id="004ED1D7FFD0FFEFFDFD448EBDC23C9A" box="[597,688,1669,1696]" metricMagnitude="3" metricUnit="m" metricValue="1.9759999999999998" pageId="46" pageNumber="47" unit="m" value="1976.0">1976 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
, Arriaga A. &
|
|||
|
|
<collectorName id="26965332FFD0FFEFFCF2448DBB7F3CA5" box="[858,1037,1669,1696]" pageId="46" pageNumber="47">Arenas A. Col.</collectorName>
|
|||
|
|
”;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFB8D448DBB1D3C9A" box="[1061,1135,1670,1696]" pageId="46" pageNumber="47">IEXA</collectionCode>
|
|||
|
|
.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF4244C2BE153CD9" blockId="46.[189,1399,1737,1873]" box="[234,359,1737,1763]" pageId="46" pageNumber="47">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF4244C2BE153CD9" bold="true" box="[234,359,1737,1763]" pageId="46" pageNumber="47">
|
|||
|
|
<typeStatus id="54D88846FFD0FFEFFF4244C2BE153CD9" box="[234,359,1737,1763]" pageId="46" pageNumber="47" type="paratype">Paratypes</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF1544E6BD643D6B" blockId="46.[189,1399,1737,1873]" pageId="46" pageNumber="47">
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFF1544E6BB6D3D3D" ID-GBIF-Occurrence="3093980385" box="[189,1055,1773,1800]" collectionCode="IEXA" collectorName="Arenas A. Col." country="Mexico" elevation="1976" latitude="17.1651" location="Oaxaca" longLatPrecision="1" longitude="96.73858" municipality="La" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="2" stateProvince="Oaxaca" typeStatus="paratype">
|
|||
|
|
<collectingCountry id="F3747674FFD0FFEFFF1544E6BE433D3D" box="[189,305,1773,1799]" name="Mexico" pageId="46" pageNumber="47">MEXICO</collectingCountry>
|
|||
|
|
–
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFEF944E6BEC23D3D" bold="true" box="[337,432,1773,1799]" pageId="46" pageNumber="47">
|
|||
|
|
<collectingRegion id="49A7F806FFD0FFEFFEF944E6BEC23D3D" box="[337,432,1773,1799]" country="Mexico" name="Oaxaca" pageId="46" pageNumber="47">Oaxaca</collectingRegion>
|
|||
|
|
</emphasis>
|
|||
|
|
•
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFE6144E6BD7F3D32" box="[457,525,1773,1800]" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
|
|||
|
|
; same collection data as for holotype;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFC7144E6BB6D3D3D" box="[985,1055,1773,1799]" pageId="46" pageNumber="47">IEXA</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
•
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFB9144E6BD603D6B" ID-GBIF-Occurrence="3093980603" collectedFrom="bosque de Encino" collectingDate="2017-07-14" collectionCode="IEXA" collectorName="Arriaga A. & Arenas A. Col." country="Mexico" elevation="1954" latitude="17.164875" location="La mesita San Pablo Etla" longLatPrecision="1" longitude="96.74821" municipality="La mesita San Pablo Etla" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca" typeStatus="paratype">
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFB9144E6BB1B3D32" box="[1081,1129,1773,1800]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
|
|||
|
|
; “
|
|||
|
|
<collectingMunicipality id="6BB8AC9EFFD0FFEFFB2944E6BF9D3D16" pageId="46" pageNumber="47">La mesita San Pablo Etla</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<date id="FFDD1024FFD0FFEFFF514519BE0E3D16" box="[249,380,1810,1836]" pageId="46" pageNumber="47" value="2017-07-14">
|
|||
|
|
<collectingDate id="EF99E9CCFFD0FFEFFF514519BE0E3D16" box="[249,380,1810,1836]" pageId="46" pageNumber="47" value="2017-07-14">14–VII–17</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
, coprotrampa,
|
|||
|
|
<date id="FFDD1024FFD0FFEFFD804519BD143D17" box="[552,614,1810,1837]" pageId="46" pageNumber="47" value="1996-10">
|
|||
|
|
x-
|
|||
|
|
<geoCoordinate id="EE575023FFD0FFEFFDEE4519BD143D17" box="[582,614,1810,1837]" degrees="96" direction="west" minutes="44" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="53.55" value="-96.74821">96</geoCoordinate>
|
|||
|
|
</date>
|
|||
|
|
°44’53.55’’O, y-
|
|||
|
|
<geoCoordinate id="EE575023FFD0FFEFFC824519BCA43D17" box="[810,982,1810,1837]" degrees="17" direction="north" minutes="9" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="53.55" value="17.164875">17°9’53.55’’N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<collectedFrom id="387F7519FFD0FFEFFC494519BBC23D16" box="[993,1200,1810,1836]" pageId="46" pageNumber="47">bosque de Encino</collectedFrom>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9B9B01FFD0FFEFFB124519BA663D16" box="[1210,1300,1810,1837]" metricMagnitude="3" metricUnit="m" metricValue="1.954" pageId="46" pageNumber="47" unit="m" value="1954.0">
|
|||
|
|
<elevation id="004ED1D7FFD0FFEFFB124519BA663D16" box="[1210,1300,1810,1837]" metricMagnitude="3" metricUnit="m" metricValue="1.954" pageId="46" pageNumber="47" unit="m" value="1954.0">1954 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<collectorName id="26965332FFD0FFEFFAB54519BFA83D6B" pageId="46" pageNumber="47">Arriaga A.</collectorName>
|
|||
|
|
&
|
|||
|
|
<collectorName id="26965332FFD0FFEFFF55453CBEC23D6A" box="[253,432,1846,1873]" pageId="46" pageNumber="47">Arenas A. Col.</collectorName>
|
|||
|
|
”;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFE60453CBD603D6B" box="[456,530,1847,1873]" pageId="46" pageNumber="47">IEXA</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF154571BD3B3DAE" blockId="46.[189,585,1913,1940]" box="[189,585,1913,1940]" pageId="46" pageNumber="47">
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFF154571BD703DA9" bold="true" box="[189,514,1913,1940]" pageId="46" pageNumber="47">Non-type material revised</emphasis>
|
|||
|
|
(
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFDBD4571BD323DAE" box="[533,576,1914,1940]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
|
|||
|
|
)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD0FFEFFF1545A1BC3C3DD2" blockId="46.[189,1398,1961,2025]" pageId="46" pageNumber="47">
|
|||
|
|
<materialsCitation id="3B0B3CB9FFD0FFEFFF1545A1BC393DD2" ID-GBIF-Occurrence="3093980470" collectingDate="2016-10-20" collectionCode="VMC" collectorName="Alfosina Arriaga" country="Mexico" elevation="1994" latitude="17.167007" location="San Pablo Etla" longLatPrecision="1" longitude="96.73729" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Oaxaca">
|
|||
|
|
<collectingCountry id="F3747674FFD0FFEFFF1545A1BE433DFE" box="[189,305,1962,1988]" name="Mexico" pageId="46" pageNumber="47">MEXICO</collectingCountry>
|
|||
|
|
–
|
|||
|
|
<emphasis id="B917EAF6FFD0FFEFFEF945A2BEC23DF9" bold="true" box="[337,432,1961,1987]" pageId="46" pageNumber="47">
|
|||
|
|
<collectingRegion id="49A7F806FFD0FFEFFEF945A2BEC23DF9" box="[337,432,1961,1987]" country="Mexico" name="Oaxaca" pageId="46" pageNumber="47">Oaxaca</collectingRegion>
|
|||
|
|
</emphasis>
|
|||
|
|
•
|
|||
|
|
<specimenCount id="9D65FD6DFFD0FFEFFE6245A1BE883DFE" box="[458,506,1962,1988]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
|
|||
|
|
; “
|
|||
|
|
<location id="8EBC603FFFD0FFEFFDBA45A1BDB43DFE" LSID="urn:lsid:plazi:treatment:03CA87F2FFD0FFF0FD9740BDBE763F02:8EBC603FFFD0FFEFFDBA45A1BDB43DFE" box="[530,710,1962,1988]" country="Mexico" latitude="17.167007" longLatPrecision="1" longitude="96.73729" name="San Pablo Etla" pageId="46" pageNumber="47" stateProvince="Oaxaca">San Pablo Etla</location>
|
|||
|
|
,
|
|||
|
|
<location id="8EBC603FFFD0FFEFFD7B45A1BC993DFE" LSID="urn:lsid:plazi:treatment:03CA87F2FFD0FFF0FD9740BDBE763F02:8EBC603FFFD0FFEFFD7B45A1BC993DFE" box="[723,1003,1962,1988]" country="Mexico" latitude="17.167007" longLatPrecision="1" longitude="96.73729" name="Reserva San Pablo Etla" pageId="46" pageNumber="47" stateProvince="Oaxaca">Reserva San Pablo Etla</location>
|
|||
|
|
,
|
|||
|
|
<date id="FFDD1024FFD0FFEFFC5045A2BB0B3DF9" box="[1016,1145,1961,1987]" pageId="46" pageNumber="47" value="2016-10-20">
|
|||
|
|
<collectingDate id="EF99E9CCFFD0FFEFFC5045A2BB0B3DF9" box="[1016,1145,1961,1987]" pageId="46" pageNumber="47" value="2016-10-20">20/X/2016</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
, C.D.
|
|||
|
|
<geoCoordinate id="EE575023FFD0FFEFFB6D45A1BA033DFE" box="[1221,1393,1962,1988]" degrees="17" direction="north" minutes="10" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="1.23" value="17.167007">17°10’1.23’’N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE575023FFD0FFEFFF1545C5BE0A3DD3" box="[189,376,1998,2025]" degrees="96" direction="east" minutes="44" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="14.25" value="96.73729">96°44’14.25’’O</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9B9B01FFD0FFEFFE2C45C5BEAD3DD3" box="[388,479,1998,2025]" metricMagnitude="3" metricUnit="m" metricValue="1.994" pageId="46" pageNumber="47" unit="m" value="1994.0">
|
|||
|
|
<elevation id="004ED1D7FFD0FFEFFE2C45C5BEAD3DD3" box="[388,479,1998,2025]" metricMagnitude="3" metricUnit="m" metricValue="1.994" pageId="46" pageNumber="47" unit="m" value="1994.0">1994 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<collectorName id="26965332FFD0FFEFFE4245C5BDDD3DD2" box="[490,687,1998,2024]" pageId="46" pageNumber="47">Alfosina Arriaga</collectorName>
|
|||
|
|
Col.”;
|
|||
|
|
<collectionCode id="ED72AE21FFD0FFEFFCAC45C5BC393DD2" box="[772,843,1998,2024]" pageId="46" pageNumber="47">VMC</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF154302BE2F3B18" blockId="47.[189,349,264,291]" box="[189,349,264,291]" pageId="47" pageNumber="48">
|
|||
|
|
<heading id="D0948188FFD1FFEEFF154302BE2F3B18" bold="true" box="[189,349,264,291]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFF154302BE2F3B18" bold="true" box="[189,349,264,291]" pageId="47" pageNumber="48">Type locality</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF15433CBC173B68" blockId="47.[189,869,311,338]" box="[189,869,311,338]" pageId="47" pageNumber="48">
|
|||
|
|
<collectingCountry id="F3747674FFD1FFEEFF15433CBE683B6B" box="[189,282,311,337]" name="Mexico" pageId="47" pageNumber="48">Mexico</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="49A7F806FFD1FFEEFE8D433CBEF23B6B" box="[293,384,311,337]" country="Mexico" name="Oaxaca" pageId="47" pageNumber="48">Oaxaca</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9B9B01FFD1FFEEFE24433CBEB53B6B" box="[396,455,311,337]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="47" pageNumber="48" unit="km" value="8.0">8 km</quantity>
|
|||
|
|
south of San Miguel Sola de Vega.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C379656FFFD1FFF0FF154373BBAA386D" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="description">
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF154373BEDD3BA9" blockId="47.[189,431,376,403]" box="[189,431,376,403]" pageId="47" pageNumber="48">
|
|||
|
|
<heading id="D0948188FFD1FFEEFF154373BEDD3BA9" bold="true" box="[189,431,376,403]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFF154373BEDD3BA9" authorityYear="2018" box="[189,431,376,403]" pageId="47" pageNumber="48">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFF154373BEDD3BA9" bold="true" box="[189,431,376,403]" italics="true" pageId="47" pageNumber="48">Phanaeus dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF1543A3BC27393B" blockId="47.[189,1399,424,770]" pageId="47" pageNumber="48">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFF1543A3BEA83BF8" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[189,474,424,450]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFEEB43A2BE083BF8" box="[323,378,424,450]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
recently described
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFD6043A2BC253BF8" box="[712,855,424,450]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFD6043A2BC253BF8" box="[712,855,424,450]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
based upon specimens from San Pablo Etla,
|
|||
|
|
<collectingRegion id="49A7F806FFD1FFEEFF1543C0BE6A3BDF" box="[189,280,459,485]" country="Mexico" name="Oaxaca" pageId="47" pageNumber="48">Oaxaca</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F3747674FFD1FFEEFE8143C0BEF13BDF" box="[297,387,459,485]" name="Mexico" pageId="47" pageNumber="48">Mexico</collectingCountry>
|
|||
|
|
and suggested that
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFDD743C7BC673BDF" authorityName="Edmonds" authorityYear="2006" box="[639,789,460,485]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFDD743C7BC673BDF" box="[639,789,460,485]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFCF043C7BC9B3BDF" box="[856,1001,459,485]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFCF043C7BC9B3BDF" box="[856,1001,459,485]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were sister taxa. The following character combination helps to separate both species according to
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFC6F43E4BB923833" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[967,1248,495,521]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFBE343E4BB0C3833" box="[1099,1150,495,521]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
:
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFB4343E4BA053833" box="[1259,1399,495,521]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFB4343E4BA053833" box="[1259,1399,495,521]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has long and slender pronotal posterolateral angles, whereas
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFC134018BB203816" authorityName="Edmonds" authorityYear="2006" box="[955,1106,531,556]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFC134018BB203816" box="[955,1106,531,556]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has short and rounded posterolateral angles. The basal border of the tergite VIII in
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFC3A403DBB523875" box="[914,1056,565,591]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFC3A403DBB523875" box="[914,1056,565,591]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
forms a small indentation at its middle, whereas it runs completely straight in
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFCAE4051BCE53849" authorityName="Edmonds" authorityYear="2006" box="[774,919,602,627]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFCAE4051BCE53849" box="[774,919,602,627]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Additionally, the apex of the parameres of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFF774076BE1F38AC" box="[223,365,636,662]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFF774076BE1F38AC" box="[223,365,636,662]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is more projected, than that from
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFCA64076BCEC38AC" authorityName="Edmonds" authorityYear="2006" box="[782,926,637,662]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFCA64076BCEC38AC" box="[782,926,637,662]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Moreover, the middle sinuation of the parameres in lateral view is much more pronounced in
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFCCC40AABC813880" box="[868,1011,672,698]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFCCC40AABC813880" box="[868,1011,672,698]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Nevertheless, we consider that the description and diagnosis of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFDE240CFBDA838E7" box="[586,730,707,733]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFDE240CFBDA838E7" box="[586,730,707,733]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
exemplify some of the frequent problems occurring in taxonomical work (
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFE0940ECBDB73938" author="Komarek A. & Beutel R. G." box="[417,709,743,770]" pageId="47" pageNumber="48" pagination="55 - 70" refId="ref39635" refString="Komarek A. & Beutel R. G. 2006. Problems in taxonomy and suggestions for a standardized description of new insect taxa. Entomological Problems 36: 55 - 70." type="journal article" year="2006">Komarek & Beutel 2006</bibRefCitation>
|
|||
|
|
) as follows.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF424123BA053997" blockId="47.[189,1399,808,977]" pageId="47" pageNumber="48">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFF424123BD5E3979" bold="true" box="[234,556,808,835]" pageId="47" pageNumber="48">Insufficient study of types</emphasis>
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFF154147BEAA395C" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[189,472,844,871]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFEEB4146BE0B395C" box="[323,377,844,870]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
did not indicate how many specimens of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFC7D4146BB1B395C" authorityName="Edmonds" authorityYear="2006" box="[981,1129,845,870]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFC7D4146BB1B395C" box="[981,1129,845,870]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
they revised, but they commented that the
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFE68417BBD4239B0" box="[448,560,880,906]" pageId="47" pageNumber="48" type="paratype">paratypes</typeStatus>
|
|||
|
|
housed at CNIN were checked by them. Taking into account this information and the original description of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFD60419FBB593994" authority="(Edmonds 2006)" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[712,1067,915,942]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFD60419FBC2A3997" box="[712,856,916,941]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFCC24198BB513994" author="Edmonds W. D." box="[874,1059,915,942]" pageId="47" pageNumber="48" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds 2006</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, we assume that
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFB534198BA053997" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[1275,1399,915,941]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">Kohlmann</bibRefCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF1741BCBDCB39EB" blockId="47.[189,1399,808,977]" box="[191,697,950,977]" pageId="47" pageNumber="48">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFF1741BCBF8339EA" box="[191,241,950,976]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018) revised
|
|||
|
|
<specimenCount id="9D65FD6DFFD1FFEEFE1941BCBD2639EB" box="[433,596,951,977]" pageId="47" pageNumber="48" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
|||
|
|
at most.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF4241F3BCCC3E28" blockId="47.[188,1400,1016,1326]" box="[234,958,1016,1042]" pageId="47" pageNumber="48">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFF4241F3BCCC3E28" bold="true" box="[234,958,1016,1042]" pageId="47" pageNumber="48">Insufficient assessment of the range of character variation</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF154610BEC73F14" blockId="47.[188,1400,1016,1326]" pageId="47" pageNumber="48">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFF154610BEA23E0C" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[189,464,1051,1078]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFEE84617BE063E0F" box="[320,372,1051,1077]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
were not able to adequately assess the morphological variation of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFB774617BA023E0F" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[1247,1392,1052,1077]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFB774617BA023E0F" box="[1247,1392,1052,1077]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, as a consequence of the reduced number of
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFD774634BC3D3E63" box="[735,847,1087,1113]" pageId="47" pageNumber="48" type="paratype">paratypes</typeStatus>
|
|||
|
|
studied by them. The pronotal posterolateral angles were found by us to be long and slender in some
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFCE84668BCD73E47" box="[832,933,1123,1149]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
|
|||
|
|
major males of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFBF64668BB9F3E46" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[1118,1261,1123,1148]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFBF64668BB9F3E46" box="[1118,1261,1123,1148]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13582A61FFD1FFEEFB554669BA1B3E47" box="[1277,1385,1122,1149]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="47" pageNumber="48">Fig. 14A</figureCitation>
|
|||
|
|
). The basal border of the tergite VIII is highly variable:
|
|||
|
|
<specimenCount id="9D65FD6DFFD1FFEEFCF3468DBB753E9A" box="[859,1031,1158,1184]" pageId="47" pageNumber="48" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFB9E468CBBB83E9A" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[1078,1226,1159,1184]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFB9E468CBBB83E9A" box="[1078,1226,1159,1184]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
revised by us show the small indentation at the middle of the basal border of the tergite VIII (
|
|||
|
|
<figureCitation id="13582A61FFD1FFEEFBD046A2BB953EF9" box="[1144,1255,1193,1219]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="47" pageNumber="48">Fig. 14D</figureCitation>
|
|||
|
|
), whereas it runs completely straight in
|
|||
|
|
<specimenCount id="9D65FD6DFFD1FFEEFDA146C6BDC03EDD" box="[521,690,1229,1255]" pageId="47" pageNumber="48" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
|||
|
|
. The small indentation is present in the basal border of the tergite VIII of
|
|||
|
|
<specimenCount id="9D65FD6DFFD1FFEEFEC346FBBD553F30" box="[363,551,1264,1290]" pageId="47" pageNumber="48" type="generic">three specimens</specimenCount>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFDE446FABDAA3F30" box="[588,728,1264,1290]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFDE446FABDAA3F30" box="[588,728,1264,1290]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
revised by us, whereas it runs completely straight in
|
|||
|
|
<specimenCount id="9D65FD6DFFD1FFEEFAE246FABE5F3F14" pageId="47" pageNumber="48" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13582A61FFD1FFEEFE96471FBEDA3F14" box="[318,424,1300,1326]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="47" pageNumber="48">Fig. 14E</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF15475EBB0A3CCF" blockId="47.[189,1400,1365,1782]" pageId="47" pageNumber="48">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFF15475EBE9D3F55" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[189,495,1365,1392]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFEE2475DBEFA3F55" box="[330,392,1365,1391]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
illustrated the endophallite copulatrix of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFBB9475DBBD53F55" box="[1041,1191,1365,1391]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFBB9475DBBD53F55" box="[1041,1191,1365,1391]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(available from https://zookeys.pensoft.net/article/23029/zoom/fig/13/) but they overlooked its comparison with
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFF574796BEE43F8C" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[255,406,1437,1462]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFF574796BEE43F8C" box="[255,406,1437,1462]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13582A61FFD1FFEEFE034797BD783F8C" box="[427,522,1436,1462]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="47" pageNumber="48">Fig. 1P</figureCitation>
|
|||
|
|
). We noticed that the morphology of both
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFB844796BBB13F8C" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[1068,1219,1437,1462]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFB844796BBB13F8C" box="[1068,1219,1437,1462]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13582A61FFD1FFEEFB714797BA473F8C" box="[1241,1333,1436,1462]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="47" pageNumber="48">Fig. 1P</figureCitation>
|
|||
|
|
) and
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFF1547CABE383FE0" box="[189,330,1472,1498]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFF1547CABE383FE0" box="[189,330,1472,1498]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is identical when we compared the endophallites under the microscope. Furthermore, the endophallite copulatrix of the
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFD8047E8BDFD3FC7" box="[552,655,1507,1533]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFD1247EFBC3A3FC7" box="[698,840,1507,1533]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFD1247EFBC3A3FC7" box="[698,840,1507,1533]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has suffered deformation and abrasion, maybe as a result of the preparation methods (soaking for 5 min in a solution of 5% boiling KOH) and/or storage (dry preservation in a plastic microvial). This structure was not found by us to be stored in a microvial with glycerol as mentioned by
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFD6F4446BC943C52" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[711,998,1613,1640]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFCE54445BCF13C5D" box="[845,899,1613,1639]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
. The morphology of the cephalic carina and the pronotal tubercles of the females of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFCEA4479BCAB3CB1" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[834,985,1650,1675]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFCEA4479BCAB3CB1" box="[834,985,1650,1675]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13582A61FFD1FFEEFC46447ABB2E3CB1" box="[1006,1116,1649,1675]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="47" pageNumber="48">Fig. 14B</figureCitation>
|
|||
|
|
) and
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFB014479BA483CB1" box="[1193,1338,1649,1675]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFB014479BA483CB1" box="[1193,1338,1649,1675]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see https://zookeys.pensoft.net/article/23029/zoom/fig/14/) did not differ. Moreover, the body colour in the
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFF1544B3BE5F3CE8" box="[189,301,1720,1746]" pageId="47" pageNumber="48" type="paratype">paratypes</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFEFE44B2BE9A3CE8" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[342,488,1721,1746]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFEFE44B2BE9A3CE8" box="[342,488,1721,1746]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
varied from jet black or bright dark blue to jet black with a green-blue sheen (
|
|||
|
|
<figureCitation id="13582A61FFD1FFEEFF6E44D0BE243CCF" box="[198,342,1755,1781]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="47" pageNumber="48">Fig. 14A–C</figureCitation>
|
|||
|
|
), falling within the variation of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFD6244D7BB063CCF" authority="(Kohlmann et al. 2018)" baseAuthorityName="Kohlmann" baseAuthorityYear="2018" box="[714,1140,1755,1782]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFD6244D7BC243CCF" box="[714,854,1755,1781]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFCCD44D0BB1E3CCF" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[869,1132,1755,1782]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFC4244D7BB6C3CCF" box="[1002,1054,1755,1781]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. 2018
|
|||
|
|
</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFD1FFEEFF154516BA6D3DD2" blockId="47.[187,1399,1821,2025]" pageId="47" pageNumber="48">
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFF154516BEBA3D0D" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[189,456,1821,1848]" pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFE954515BE1C3D0D" box="[317,366,1821,1847]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
commented that a major male was designated as
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFC5F4516BB123D0D" box="[1015,1120,1821,1847]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
|||
|
|
. Nevertheless, we found out that the
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFEE3454BBEC33D60" box="[331,433,1856,1882]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFE77454ABD1E3D60" box="[479,620,1856,1882]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFE77454ABD1E3D60" box="[479,620,1856,1882]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
housed at IEXA is the minor male illustrated by
|
|||
|
|
<bibRefCitation id="EFF24B15FFD1FFEEFB1F454BBE783D44" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." pageId="47" pageNumber="48" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFA94454ABA023D60" box="[1340,1392,1856,1882]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
in the original description (available from https://zookeys.pensoft.net/article/23029/zoom/fig/15/). As a consequence, we assume that the original description of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFC294583BB7E3D9B" box="[897,1036,1927,1953]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFC294583BB7E3D9B" box="[897,1036,1927,1953]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was not based on the
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFAA0458CBA013D9B" box="[1288,1395,1927,1953]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
|||
|
|
. We found that a
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFE2745A0BE863DFF" box="[399,500,1963,1989]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
|
|||
|
|
minor male of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFD1545A7BC203DFF" baseAuthorityName="Edmonds" baseAuthorityYear="2006" box="[701,850,1964,1989]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFD1545A7BC203DFF" box="[701,850,1964,1989]" italics="true" pageId="47" pageNumber="48">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13582A61FFD1FFEEFCCF45A0BCA73DFF" box="[871,981,1963,1989]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="47" pageNumber="48">Fig. 14C</figureCitation>
|
|||
|
|
) and the
|
|||
|
|
<typeStatus id="54D88846FFD1FFEEFBE745A0BBC43DFF" box="[1103,1206,1963,1989]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C634D67FFD1FFEEFB4E45A7BA053DFF" box="[1254,1399,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFD1FFEEFB4E45A7BA053DFF" box="[1254,1399,1963,1989]" italics="true" pageId="47" pageNumber="48">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(https://zookeys.pensoft.net/article/23029/zoom/fig/15/) were morphologically almost identical.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF1C666CFFCEFFF1FF15455EBEDB3D8C" ID-DOI="http://doi.org/10.5281/zenodo.4836794" ID-Zenodo-Dep="4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="48" pageNumber="49" startId="48.[189,232,1877,1903]" targetBox="[228,1363,265,1833]" targetPageId="48">
|
|||
|
|
<paragraph id="8BDC36E4FFCEFFF1FF15455EBEDB3D8C" blockId="48.[189,1406,1877,1974]" pageId="48" pageNumber="49">
|
|||
|
|
<emphasis id="B917EAF6FFCEFFF1FF15455EBE253D55" bold="true" box="[189,343,1877,1903]" pageId="48" pageNumber="49">Fig. 14. A–D</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C634D67FFCEFFF1FECD455EBC663D55" authority="Edmonds, 2006" authorityName="Edmonds" authorityYear="2006" box="[357,788,1877,1903]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFCEFFF1FECD455EBD213D55" box="[357,595,1877,1903]" italics="true" pageId="48" pageNumber="49">Phanaeus zapotecus</emphasis>
|
|||
|
|
<bibRefCitation id="EFF24B15FFCEFFF1FDF2455EBC663D55" author="Edmonds W. D." box="[602,788,1877,1903]" pageId="48" pageNumber="49" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds, 2006</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B917EAF6FFCEFFF1FCB7455EBC463D55" bold="true" box="[799,820,1877,1903]" pageId="48" pageNumber="49">A</emphasis>
|
|||
|
|
. ♂ (TAMU).
|
|||
|
|
<emphasis id="B917EAF6FFCEFFF1FC7D455EBC9B3D55" bold="true" box="[981,1001,1877,1903]" pageId="48" pageNumber="49">B</emphasis>
|
|||
|
|
. ♀ (TAMU).
|
|||
|
|
<emphasis id="B917EAF6FFCEFFF1FB21455EBBEC3D55" bold="true" box="[1161,1182,1877,1903]" pageId="48" pageNumber="49">C</emphasis>
|
|||
|
|
. Minor ♂ (VMC).
|
|||
|
|
<emphasis id="B917EAF6FFCEFFF1FF154573BFA03DA8" bold="true" box="[189,210,1912,1938]" pageId="48" pageNumber="49">D</emphasis>
|
|||
|
|
. Tergite VIII (TAMU).
|
|||
|
|
<emphasis id="B917EAF6FFCEFFF1FE5B4573BD773DA8" bold="true" box="[499,517,1912,1938]" pageId="48" pageNumber="49">E</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C634D67FFCEFFF1FDBD4573BB893DA8" authority="Kohlmann, Arriaga-Jimenez & Ros, 2018" authorityName="Kohlmann, Arriaga-Jimenez & Ros" authorityYear="2018" box="[533,1275,1912,1939]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFCEFFF1FDBD4573BC733DA8" box="[533,769,1912,1938]" italics="true" pageId="48" pageNumber="49">Phanaeus dionysius</emphasis>
|
|||
|
|
<bibRefCitation id="EFF24B15FFCEFFF1FCA44573BB893DA8" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[780,1275,1912,1939]" pageId="48" pageNumber="49" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">Kohlmann, Arriaga-Jiménez & Rös, 2018</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A224578DFFCEFFF1FAAC4572BA0A3DA9" box="[1284,1400,1913,1939]" pageId="48" pageNumber="49">syn. nov.</taxonomicNameLabel>
|
|||
|
|
, tergite VIII (IEXA).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BDC36E4FFCFFFF0FF424303BC993B19" blockId="49.[188,1398,264,430]" box="[234,1003,264,291]" pageId="49" pageNumber="50">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FF424303BC993B19" bold="true" box="[234,1003,264,291]" pageId="49" pageNumber="50">Characters not suitable for a study at a given taxonomic rank</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFCFFFF0FF154320BEE63B97" blockId="49.[188,1398,264,430]" pageId="49" pageNumber="50">
|
|||
|
|
The revision of a majority of species within the
|
|||
|
|
<taxonomicName id="4C634D67FFCFFFF0FCBA4327BCD73B7F" baseAuthorityName="Edmonds" baseAuthorityYear="1994" box="[786,933,299,325]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="endymion">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FCBA4327BCD73B7F" box="[786,933,299,325]" italics="true" pageId="49" pageNumber="50">P. endymion</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species group led us to conclude that the morphology of the aedeagus is not taxonomically informative to separate species, as suggested by previous authors (
|
|||
|
|
<bibRefCitation id="EFF24B15FFCFFFF0FE27437BBD363BB1" author="Edmonds W. D." box="[399,580,368,395]" pageId="49" pageNumber="50" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds 1994</bibRefCitation>
|
|||
|
|
; Price 1995;
|
|||
|
|
<bibRefCitation id="EFF24B15FFCFFFF0FD73437BBCF53BB0" author="Arnaud P." box="[731,903,368,394]" pageId="49" pageNumber="50" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFF24B15FFCFFFF0FC3A437ABB923BB1" author="Moctezuma V. & Halffter G." box="[914,1248,368,395]" pageId="49" pageNumber="50" pagination="47 - 56" refId="ref40548" refString="Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47" type="journal article" year="2017">Moctezuma & Halffter 2017</bibRefCitation>
|
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|
|
; Moctezuma
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FF14439FBF823B97" box="[188,240,403,429]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
|
|||
|
|
. 2017, 2019).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFCFFFF0FF1543DEBBAA386D" blockId="49.[189,1398,468,599]" pageId="49" pageNumber="50">
|
|||
|
|
In the light of the findings presented herein, we conclude that the description and diagnoses provided by
|
|||
|
|
<bibRefCitation id="EFF24B15FFCFFFF0FF4B43FCBE8B382B" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[227,505,503,529]" pageId="49" pageNumber="50" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">
|
|||
|
|
Kohlmann
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FEC043F3BEE9382B" box="[360,411,503,529]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
|
|||
|
|
. (2018)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
do not justify the splitting of
|
|||
|
|
<taxonomicName id="4C634D67FFCFFFF0FCF643F3BC99382B" authorityName="Kohlmann, Arriaga-Jimenez & Ros" authorityYear="2018" box="[862,1003,503,529]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FCF643F3BC99382B" box="[862,1003,503,529]" italics="true" pageId="49" pageNumber="50">P. dionysius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C634D67FFCFFFF0FB8E43F3BBCA382B" authorityName="Edmonds" authorityYear="2006" box="[1062,1208,504,529]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FB8E43F3BBCA382B" box="[1062,1208,504,529]" italics="true" pageId="49" pageNumber="50">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in two different species. As a consequence, a new junior subjective synonymy is recognized herein:
|
|||
|
|
<taxonomicName id="4C634D67FFCFFFF0FB204012BE07386C" authority="Edmonds, 2006" authorityName="Edmonds" authorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FB204012BA043809" box="[1160,1398,537,563]" italics="true" pageId="49" pageNumber="50">Phanaeus zapotecus</emphasis>
|
|||
|
|
<bibRefCitation id="EFF24B15FFCFFFF0FF154037BE07386C" author="Edmonds W. D." box="[189,373,572,598]" pageId="49" pageNumber="50" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds, 2006</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
=
|
|||
|
|
<taxonomicName id="4C634D67FFCFFFF0FE214037BB15386C" authority="Kohlmann, Arriaga-Jimenez & Ros, 2018" authorityName="Kohlmann, Arriaga-Jimenez & Ros" authorityYear="2018" box="[393,1127,572,599]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="dionysius">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FE214037BD01386C" box="[393,627,572,598]" italics="true" pageId="49" pageNumber="50">Phanaeus dionysius</emphasis>
|
|||
|
|
<bibRefCitation id="EFF24B15FFCFFFF0FDD64037BB15386C" author="Kohlmann B. & Arriaga-Jimenez A. & Ros M." box="[638,1127,572,599]" pageId="49" pageNumber="50" pagination="67 - 93" refId="ref39574" refString="Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029" type="journal article" year="2018">Kohlmann, Arriaga-Jiménez & Rös, 2018</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A224578DFFCFFFF0FBC64036BBAA386D" box="[1134,1240,573,599]" pageId="49" pageNumber="50">syn. nov.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C379656FFFCFFFF0FF154076BCDC38FC" box="[189,942,637,710]" pageId="49" pageNumber="50" type="distribution">
|
|||
|
|
<paragraph id="8BDC36E4FFCFFFF0FF154076BE2438AD" blockId="49.[189,342,637,663]" box="[189,342,637,663]" pageId="49" pageNumber="50">
|
|||
|
|
<heading id="D0948188FFCFFFF0FF154076BE2438AD" bold="true" box="[189,342,637,663]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FF154076BE2438AD" bold="true" box="[189,342,637,663]" pageId="49" pageNumber="50">Distribution</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFCFFFF0FF1540A0BCDC38FC" blockId="49.[189,942,683,710]" box="[189,942,683,710]" pageId="49" pageNumber="50">
|
|||
|
|
Sierra Madre del Sur and Sierra Norte, central
|
|||
|
|
<collectingRegion id="49A7F806FFCFFFF0FD4940A0BC4B38FF" box="[737,825,683,709]" country="Mexico" name="Oaxaca" pageId="49" pageNumber="50">Oaxaca</collectingRegion>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13582A61FFCFFFF0FCE140A0BCD238FC" box="[841,928,683,710]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="49" pageNumber="50">Fig. 15</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C379656FFFCFFFF0FF1540E7BE763F02" pageId="49" pageNumber="50" type="discussion">
|
|||
|
|
<paragraph id="8BDC36E4FFCFFFF0FF1540E7BE5D393C" blockId="49.[189,303,748,774]" box="[189,303,748,774]" pageId="49" pageNumber="50">
|
|||
|
|
<heading id="D0948188FFCFFFF0FF1540E7BE5D393C" bold="true" box="[189,303,748,774]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FF1540E7BE5D393C" bold="true" box="[189,303,748,774]" pageId="49" pageNumber="50">Remarks</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFCFFFF0FF154110BECE3E70" blockId="49.[188,1401,794,1098]" pageId="49" pageNumber="50">
|
|||
|
|
Mean length
|
|||
|
|
<quantity id="4C9B9B01FFCFFFF0FEC84111BEBE390F" box="[352,460,794,821]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="49" pageNumber="50" unit="mm" value="16.4">16.4 mm</quantity>
|
|||
|
|
(
|
|||
|
|
<quantity id="4C9B9B01FFCFFFF0FE774111BDE6390F" box="[479,660,794,821]" metricMagnitude="-2" metricUnit="m" metricValue="1.6649999999999998" metricValueMax="1.85" metricValueMin="1.48" pageId="49" pageNumber="50" unit="mm" value="16.65" valueMax="18.5" valueMin="14.8">14.8–18.5 mm</quantity>
|
|||
|
|
). To the original description of this species, we add that the elytral striae are impressed basally as distinct fossae (
|
|||
|
|
<figureCitation id="13582A61FFCFFFF0FCE54136BCD5396D" box="[845,935,829,855]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="49" pageNumber="50">Fig. 14</figureCitation>
|
|||
|
|
); right lobe of endophallite copulatrix more developed than left lobe; right lobe obtusely triangular in shape, strongly developed superiorly; left lobe strongly reduced, lobed, slightly concave inferiorly; central ridge more developed than central column (
|
|||
|
|
<figureCitation id="13582A61FFCFFFF0FE8E41AEBE0D3985" box="[294,383,933,959]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="49" pageNumber="50">Fig. 1P</figureCitation>
|
|||
|
|
). For the female, the head shows a cephalic trituberculate carina; with almost aligned, conical tubercles; middle tubercle slightly more developed than lateral tubercles; frons with almost effaced punctures; pronotal process trituberculate, with posterior concavity; rounded pronotal tubercles; with middle tubercle slightly more developed than lateral tubercles; posterior pronotal midline distinctly impressed (
|
|||
|
|
<figureCitation id="13582A61FFCFFFF0FEEB463BBEDC3E70" box="[323,430,1072,1098]" captionStart="Fig" captionStartId="48.[189,232,1877,1903]" captionTargetBox="[228,1363,265,1833]" captionTargetId="figure-40@48.[189,1399,265,1833]" captionTargetPageId="48" captionText="Fig. 14. A–D. Phanaeus zapotecus Edmonds, 2006. A. ♂ (TAMU). B. ♀ (TAMU). C. Minor ♂ (VMC). D. Tergite VIII (TAMU). E. Phanaeus dionysius Kohlmann, Arriaga-Jiménez & Rös, 2018 syn. nov., tergite VIII (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836794" httpUri="https://zenodo.org/record/4836794/files/figure.png" pageId="49" pageNumber="50">Fig. 14B</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDC36E4FFCFFFF0FF15467ABE763F02" blockId="49.[188,1400,1136,1336]" pageId="49" pageNumber="50">
|
|||
|
|
<bibRefCitation id="EFF24B15FFCFFFF0FF15467ABEF13EB1" author="Edmonds W. D." box="[189,387,1136,1163]" pageId="49" pageNumber="50" pagination="31 - 37" refId="ref37655" refString="Edmonds W. D. 2006. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico. Zootaxa 1171 (3): 31 - 37. https: // doi. org / 10.11646 / zootaxa. 1171.1.3" type="journal article" year="2006">Edmonds (2006)</bibRefCitation>
|
|||
|
|
commented that the
|
|||
|
|
<typeStatus id="54D88846FFCFFFF0FD29467ABD9A3EB1" box="[641,744,1137,1163]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="9D65FD6DFFCFFFF0FC8C467ABCAA3EB1" box="[804,984,1137,1163]" pageId="49" pageNumber="50" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
|
|||
|
|
(
|
|||
|
|
<specimenCount id="9D65FD6DFFCFFFF0FC45467ABB693EB1" box="[1005,1051,1137,1163]" pageId="49" pageNumber="50" type="male">1 ♂</specimenCount>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D65FD6DFFCFFFF0FB81467ABB193EB1" box="[1065,1131,1137,1163]" pageId="49" pageNumber="50" type="female">2 ♀♀</specimenCount>
|
|||
|
|
) of
|
|||
|
|
<taxonomicName id="4C634D67FFCFFFF0FB0A467ABA473EB0" authorityName="Edmonds" authorityYear="2006" box="[1186,1333,1137,1162]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FB0A467ABA473EB0" box="[1186,1333,1137,1162]" italics="true" pageId="49" pageNumber="50">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were housed at IEXA. However, the type material was not found by us at IEXA in a recent search. Therefore, the
|
|||
|
|
<typeStatus id="54D88846FFCFFFF0FF4F46BDBE3C3EEA" box="[231,334,1206,1232]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
|
|||
|
|
and these
|
|||
|
|
<typeStatus id="54D88846FFCFFFF0FE6046BCBD4A3EEB" box="[456,568,1207,1233]" pageId="49" pageNumber="50" type="paratype">paratypes</typeStatus>
|
|||
|
|
are considered temporarily lost. It is possible that the former collection manager of IEXA, Professor Miguel Angel Morón Ríos, knew the location of the
|
|||
|
|
<typeStatus id="54D88846FFCFFFF0FBC046D2BBBD3EC9" box="[1128,1231,1241,1267]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
|
|||
|
|
and
|
|||
|
|
<typeStatus id="54D88846FFCFFFF0FAAD46D2BA073EC9" box="[1285,1397,1241,1267]" pageId="49" pageNumber="50" type="paratype">paratypes</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C634D67FFCFFFF0FF7646F7BE033F2F" authorityName="Edmonds" authorityYear="2006" box="[222,369,1276,1301]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="zapotecus">
|
|||
|
|
<emphasis id="B917EAF6FFCFFFF0FF7646F7BE033F2F" box="[222,369,1276,1301]" italics="true" pageId="49" pageNumber="50">P. zapotecus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but, unfortunately, we were not able to confirm this assumption because of his recent death.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|