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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3841.3.1" ID-GBIF-Dataset="d22b4cec-127a-4017-89f0-7f0a96b9fdc7" ID-ISSN="1175-5326" ID-Zenodo-Dep="228244" ID-ZooBank="29968B5C-8DA9-46B3-BEE9-4F569C12143C" checkinTime="1460461675414" checkinUser="plazi" docAuthor="Richard, Jasmine &amp; Clark, Paul F." docDate="2014" docId="03B4878D180DFFFAB6C6FC13FA94FA78" docLanguage="en" docName="zt03841p338.pdf" docOrigin="Zootaxa 3841 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caridina coulaborensis, sp. nov." docType="treatment" docVersion="12" lastPageNumber="326" masterDocId="FF8DFFF5181AFFE3B651FFB6FF80FC4E" masterDocTitle="Caridina simoni Bouvier, 1904 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) and the synonymy by Johnson, 1963" masterLastPageNumber="338" masterPageNumber="301" pageNumber="324" updateTime="1676484100243" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Caridina simoni Bouvier, 1904 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) and the synonymy by Johnson, 1963</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Richard, Jasmine</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Clark, Paul F.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-07-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3841</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>301</mods:start>
<mods:end>338</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3841.3.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">d22b4cec-127a-4017-89f0-7f0a96b9fdc7</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">228244</mods:identifier>
<mods:identifier type="ZooBank">29968B5C-8DA9-46B3-BEE9-4F569C12143C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5315520" ID-GBIF-Taxon="119512662" ID-Zenodo-Dep="5315520" LSID="urn:lsid:plazi:treatment:03B4878D180DFFFAB6C6FC13FA94FA78" httpUri="http://treatment.plazi.org/id/03B4878D180DFFFAB6C6FC13FA94FA78" lastPageId="25" lastPageNumber="326" pageId="23" pageNumber="324">
<subSubSection box="[151,534,933,959]" pageId="23" pageNumber="324" type="nomenclature">
<paragraph blockId="23.[151,534,933,992]" box="[151,534,933,959]" pageId="23" pageNumber="324">
<heading bold="true" box="[151,534,933,959]" fontSize="11" level="1" pageId="23" pageNumber="324" reason="1">
<emphasis bold="true" box="[151,534,933,959]" pageId="23" pageNumber="324">
<taxonomicName ID-CoL="8VK62" ID-ENA="2486195" authority="Richard &amp; Clark, 2014" box="[151,433,933,959]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="coulaborensis" status="sp. nov.">
<emphasis bold="true" box="[151,433,933,959]" italics="true" pageId="23" pageNumber="324">Caridina coulaborensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[441,534,933,959]" pageId="23" pageNumber="324" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,249,967,992]" pageId="23" pageNumber="324" type="description">
<paragraph blockId="23.[151,534,933,992]" box="[151,249,967,992]" pageId="23" pageNumber="324">
(
<figureCitation box="[159,240,967,992]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="23" pageNumber="324">
Fig.
<number box="[212,240,968,992]" pageId="23" pageNumber="324" value="13.0">13</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="324" type="materials_examined">
<paragraph blockId="23.[151,1437,1039,2037]" pageId="23" pageNumber="324">
<emphasis bold="true" box="[151,388,1039,1064]" pageId="23" pageNumber="324">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="4029644323" collectingDate="2012-02-06" collectionCode="NMB" collectorName="F. Sarasin &amp; J. Roux" country="New Caledonia" location="Coula-Boreare" municipality="Coula-Boreare" pageId="23" pageNumber="324" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<taxonomicName box="[398,650,1039,1064]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis box="[398,650,1039,1064]" italics="true" pageId="23" pageNumber="324">
<originalDetermination box="[398,650,1039,1064]" pageId="23" pageNumber="324">Caridina meridionalis</originalDetermination>
</emphasis>
</taxonomicName>
. Types:
<typeStatus box="[753,861,1039,1064]" pageId="23" pageNumber="324">Holotype</typeStatus>
. Cotype.
<emphasis bold="true" box="[972,1164,1039,1064]" pageId="23" pageNumber="324">
<collectingCountry box="[972,1160,1039,1064]" name="New Caledonia" pageId="23" pageNumber="324">New Caledonia</collectingCountry>
.
</emphasis>
<collectingMunicipality box="[1174,1342,1039,1064]" pageId="23" pageNumber="324">Coula-Boréaré</collectingMunicipality>
, coll.
<collectorName pageId="23" pageNumber="324">F. Sarasin &amp; J. Roux</collectorName>
,
<number box="[376,415,1076,1101]" pageId="23" pageNumber="324" value="6.2">6.2</number>
.
<number box="[418,446,1076,1101]" pageId="23" pageNumber="324" value="12.0">12</number>
(another label, Drs. F. Sarasin &amp; J. Roux, Coula Fluss bei Boréaré,
<number box="[1228,1243,1076,1101]" pageId="23" pageNumber="324" value="6.0">6</number>
.II.
<number box="[1273,1303,1076,1101]" pageId="23" pageNumber="324" value="12.0">12</number>
)
<collectionCode box="[1321,1385,1077,1100]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="23" pageNumber="324">NMB</collectionCode>
reg.
<number box="[151,166,1112,1136]" pageId="23" pageNumber="324" value="4.0">4</number>
.VIII.e,
<specimenCount box="[254,291,1112,1136]" pageId="23" pageNumber="324" type="male">
<number box="[254,266,1112,1136]" pageId="23" pageNumber="324" value="1.0">1</number>
♂
</specimenCount>
;
</materialsCitation>
<materialsCitation collectingDate="2012-02-06" collectionCode="NMB" collectorName="F. Sarasin &amp; J. Roux" location="Coula-Boreare" municipality="Coula-Boreare" pageId="23" pageNumber="324" specimenCount="26" specimenCount-female="15" specimenCount-male="11" typeStatus="paratype">
<typeStatus box="[305,418,1112,1136]" pageId="23" pageNumber="324">Paratypes</typeStatus>
. Cotypes.
<collectingMunicipality box="[535,703,1111,1136]" pageId="23" pageNumber="324">Coula-Boréaré</collectingMunicipality>
, coll.
<collectorName box="[772,1010,1111,1136]" pageId="23" pageNumber="324">F. Sarasin &amp; J. Roux</collectorName>
,
<date box="[1022,1092,1111,1136]" pageId="23" pageNumber="324" value="12-02-06">
<collectingDate box="[1022,1092,1111,1136]" pageId="23" pageNumber="324" value="12-02-06">
<number box="[1022,1061,1111,1136]" pageId="23" pageNumber="324" value="6.2">6.2</number>
.
<number box="[1064,1092,1111,1136]" pageId="23" pageNumber="324" value="12.0">12</number>
</collectingDate>
</date>
(another label, Drs. F. Sarasin &amp; J. Roux, Coula Fluss bei Boréaré,
<number box="[566,581,1148,1173]" pageId="23" pageNumber="324" value="6.0">6</number>
.II.
<number box="[611,641,1148,1173]" pageId="23" pageNumber="324" value="12.0">12</number>
)
<collectionCode box="[658,722,1149,1172]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="23" pageNumber="324">NMB</collectionCode>
reg.
<number box="[779,794,1148,1172]" pageId="23" pageNumber="324" value="4.0">4</number>
.VIII.e,
<specimenCount box="[882,932,1148,1172]" pageId="23" pageNumber="324" type="male">
<number box="[882,910,1148,1172]" pageId="23" pageNumber="324" value="11.0">11</number>
♂
</specimenCount>
,
<specimenCount box="[944,979,1148,1172]" pageId="23" pageNumber="324" type="female">
<number box="[944,962,1148,1172]" pageId="23" pageNumber="324" value="4.0">4</number>
♀
</specimenCount>
ovig.,
<specimenCount box="[1058,1108,1148,1172]" pageId="23" pageNumber="324" type="female">
<number box="[1058,1090,1148,1172]" pageId="23" pageNumber="324" value="11.0">11</number>
♀
</specimenCount>
;
</materialsCitation>
<materialsCitation collectingDate="2011-08-20" collectionCode="NMB" collectorName="F. Sarasin &amp; J. Roux" location="Tiouaka Tal" municipality="Tiouaka Tal" pageId="23" pageNumber="324" specimenCount="12" specimenCount-female="7" specimenCount-male="5">
<collectingMunicipality box="[1121,1258,1148,1173]" pageId="23" pageNumber="324">Tiouaka Tal</collectingMunicipality>
, coll.
<collectorName pageId="23" pageNumber="324">F. Sarasin &amp; J. Roux</collectorName>
,
<number box="[277,326,1184,1208]" pageId="23" pageNumber="324" value="20.8">20.8</number>
.
<number box="[333,360,1184,1208]" pageId="23" pageNumber="324" value="11.0">11</number>
(another label, Drs. F. Sarasin &amp; J. Roux, Mune Tiouaka
<number box="[1000,1028,1184,1208]" pageId="23" pageNumber="324" value="20.0">20</number>
avec C. sp.)
<collectionCode box="[1172,1236,1184,1207]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="23" pageNumber="324">NMB</collectionCode>
reg.
<number box="[1292,1307,1184,1208]" pageId="23" pageNumber="324" value="4.0">4</number>
.VIII.c,
<specimenCount box="[1394,1431,1183,1208]" pageId="23" pageNumber="324" type="male">
<number box="[1394,1409,1183,1208]" pageId="23" pageNumber="324" value="5.0">5</number>
♂
</specimenCount>
,
<specimenCount box="[151,186,1220,1244]" pageId="23" pageNumber="324" type="female">
<number box="[151,165,1220,1244]" pageId="23" pageNumber="324" value="1.0">1</number>
♀
</specimenCount>
ovig.,
<specimenCount box="[266,302,1220,1245]" pageId="23" pageNumber="324" type="female">
<number box="[266,284,1220,1245]" pageId="23" pageNumber="324" value="6.0">6</number>
♀
</specimenCount>
,
<number box="[316,330,1220,1244]" pageId="23" pageNumber="324" value="3.0">3</number>
damaged specimens;
</materialsCitation>
<materialsCitation collectingDate="1911-08-09" collectionCode="NMB" collectorName="F Sarasin &amp; J. Roux" location="Kone" municipality="Kone" pageId="23" pageNumber="324" specimenCount="15" specimenCount-female="7" specimenCount-male="8">
<collectingMunicipality box="[581,645,1220,1244]" pageId="23" pageNumber="324">Koné</collectingMunicipality>
, coll.
<collectorName box="[714,948,1220,1245]" pageId="23" pageNumber="324">F Sarasin &amp; J. Roux</collectorName>
,
<date box="[961,1058,1220,1244]" pageId="23" pageNumber="324" value="1911-08-09">
<collectingDate box="[961,1058,1220,1244]" pageId="23" pageNumber="324" value="1911-08-09">
<number box="[961,998,1220,1244]" pageId="23" pageNumber="324" value="9.8">9.8</number>
.
<number box="[1001,1058,1220,1244]" pageId="23" pageNumber="324" value="1911.0">1911</number>
</collectingDate>
</date>
<collectionCode box="[1065,1129,1221,1244]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="23" pageNumber="324">NMB</collectionCode>
reg.
<date box="[1181,1256,1220,1245]" pageId="23" pageNumber="324">
<number box="[1181,1196,1220,1245]" pageId="23" pageNumber="324" value="4.0">4</number>
.VIII.
</date>
d
<specimenCount box="[1279,1316,1220,1244]" pageId="23" pageNumber="324" type="male">
<number box="[1279,1293,1220,1244]" pageId="23" pageNumber="324" value="8.0">8</number>
♂
</specimenCount>
,
<specimenCount box="[1329,1364,1220,1244]" pageId="23" pageNumber="324" type="female">
<number box="[1329,1347,1220,1244]" pageId="23" pageNumber="324" value="4.0">4</number>
♀
</specimenCount>
ovig.,
<specimenCount box="[151,187,1256,1280]" pageId="23" pageNumber="324" type="female">
<number box="[151,168,1256,1280]" pageId="23" pageNumber="324" value="3.0">3</number>
♀
</specimenCount>
;
</materialsCitation>
<materialsCitation box="[206,889,1255,1280]" collectingDate="2012-01-16" collectionCode="NMB" location="Lao Foa" pageId="23" pageNumber="324" specimenCount="28" specimenCount-female="12" specimenCount-male="16">
<location LSID="urn:lsid:plazi:treatment:03B4878D180DFFFAB6C6FC13FA94FA78:8EC26040180DFFF4B69FFB5EFEB2F8B1" box="[206,306,1256,1279]" name="Lao Foa" pageId="23" pageNumber="324">Lao Foa</location>
,
<number box="[323,376,1255,1280]" pageId="23" pageNumber="324" value="16.1">16.1</number>
.
<number box="[379,409,1255,1280]" pageId="23" pageNumber="324" value="12.0">12</number>
,
<collectionCode box="[426,490,1256,1279]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="23" pageNumber="324">NMB</collectionCode>
reg.
<number box="[557,572,1255,1280]" pageId="23" pageNumber="324" value="4.0">4</number>
.VIII.f
<specimenCount box="[654,706,1255,1280]" pageId="23" pageNumber="324" type="male">
<number box="[654,684,1255,1280]" pageId="23" pageNumber="324" value="16.0">16</number>
♂
</specimenCount>
,
<specimenCount box="[723,758,1256,1279]" pageId="23" pageNumber="324" type="female">
<number box="[723,741,1256,1279]" pageId="23" pageNumber="324" value="7.0">7</number>
♀
</specimenCount>
ovig.,
<specimenCount box="[846,883,1255,1280]" pageId="23" pageNumber="324" type="female">
<number box="[846,864,1255,1280]" pageId="23" pageNumber="324" value="5.0">5</number>
♀
</specimenCount>
;
</materialsCitation>
<materialsCitation box="[901,1380,1255,1280]" collectingDate="2012-12-01" collectionCode="NMB" location="Coinde" pageId="23" pageNumber="324" specimenCount="3" specimenCount-male="3">
<location LSID="urn:lsid:plazi:treatment:03B4878D180DFFFAB6C6FC13FA94FA78:8EC26040180DFFF4B5D4FB51FC56F94E" box="[901,982,1255,1280]" name="Coinde" pageId="23" pageNumber="324">Coindé</location>
<number box="[994,1047,1256,1280]" pageId="23" pageNumber="324" value="12.1">12.1</number>
.
<number box="[1050,1080,1256,1280]" pageId="23" pageNumber="324" value="12.0">12</number>
,
<collectionCode box="[1097,1161,1256,1279]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="23" pageNumber="324">NMB</collectionCode>
reg.
<number box="[1228,1243,1256,1280]" pageId="23" pageNumber="324" value="4.0">4</number>
.VIII.g,
<specimenCount box="[1337,1375,1256,1280]" pageId="23" pageNumber="324" type="male">
<number box="[1337,1351,1256,1280]" pageId="23" pageNumber="324" value="3.0">3</number>
♂
</specimenCount>
;
</materialsCitation>
<materialsCitation collectingDate="2011-03" collectionCode="NMB" collectorName="F. Sarasin &amp; J. Roux" location="Noumea &amp; Paita" pageId="23" pageNumber="324" specimenCount="1" specimenCount-female="1">
ZW
<location LSID="urn:lsid:plazi:treatment:03B4878D180DFFFAB6C6FC13FA94FA78:8EC26040180DFFF4B6C6FABAFED4F96A" box="[151,340,1292,1316]" name="Noumea &amp; Paita" pageId="23" pageNumber="324">Nouméa &amp; Païta</location>
,
<date box="[351,415,1292,1316]" pageId="23" pageNumber="324" value="2011-03">
<collectingDate box="[351,415,1292,1316]" pageId="23" pageNumber="324" value="2011-03">
III.
<number box="[388,415,1292,1316]" pageId="23" pageNumber="324" value="11.0">11</number>
</collectingDate>
</date>
, coll.
<collectorName box="[483,717,1292,1317]" pageId="23" pageNumber="324">F. Sarasin &amp; J. Roux</collectorName>
,
<collectionCode box="[728,792,1293,1316]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="23" pageNumber="324">NMB</collectionCode>
reg.
<number box="[849,864,1292,1317]" pageId="23" pageNumber="324" value="4.0">4</number>
.VIII.h,
<specimenCount box="[953,988,1292,1316]" pageId="23" pageNumber="324" type="female">
<number box="[953,967,1292,1316]" pageId="23" pageNumber="324" value="1.0">1</number>
♀
</specimenCount>
ovig.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="326" pageId="23" pageNumber="324" type="description">
<paragraph blockId="23.[151,1437,1039,2037]" box="[199,1346,1327,1352]" pageId="23" pageNumber="324">
<emphasis bold="true" box="[199,345,1327,1352]" pageId="23" pageNumber="324">Description.</emphasis>
From male
<typeStatus box="[480,578,1327,1352]" pageId="23" pageNumber="324">holotype</typeStatus>
and paraptypes. Adult size
<quantity box="[889,1014,1327,1352]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.5" metricValueMin="2.3" pageId="23" pageNumber="324" unit="mm" value="29.0" valueMax="35.0" valueMin="23.0">23–35 mm</quantity>
. Carapace length 3.0–
<quantity box="[1262,1346,1327,1352]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="324" unit="mm" value="4.5">4.5 mm</quantity>
</paragraph>
<paragraph blockId="23.[151,1437,1039,2037]" pageId="23" pageNumber="324">
<emphasis box="[199,294,1365,1388]" italics="true" pageId="23" pageNumber="324">Rostrum</emphasis>
(
<figureCitation box="[311,395,1364,1389]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="23" pageNumber="324">
Fig.
<number box="[365,395,1364,1388]" pageId="23" pageNumber="324" value="13.0">13</number>
</figureCitation>
a): Length variable reaching antennal scale or fractionally longer or as shorter as middle of third antennular peduncle segment. Rostrum
<quantity box="[675,814,1399,1424]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="23" pageNumber="324" unit="mm" value="3.5" valueMax="4.5" valueMin="2.5">2.5–4.5 mm</quantity>
<number box="[825,860,1399,1424]" pageId="23" pageNumber="324" value="0.8">0.8</number>
–1.0×long as carapace
<number box="[1129,1197,1400,1424]" pageId="23" pageNumber="324" value="16.5" valueMax="22.0" valueMin="11.0">11–22</number>
teeth proximally on dorsal margin leaving
<number box="[405,502,1436,1461]" pageId="23" pageNumber="324" value="0.375" valueMax="0.45" valueMin="0.3">0.3–0.45</number>
of the dorsal margin unarmed distally.
<number box="[949,991,1436,1460]" pageId="23" pageNumber="324" value="1.5" valueMax="2.0" valueMin="1.0">1–2</number>
post orbital teeth. Tip bifid.
<number box="[1318,1374,1436,1460]" pageId="23" pageNumber="324" value="12.0" valueMax="17.0" valueMin="7.0">7–17</number>
teeth from the proximal end to the tip or with a short distal unarmed end on the ventral margin. Formula (
<number box="[1299,1343,1472,1496]" pageId="23" pageNumber="324" value="1.5" valueMax="2.0" valueMin="1.0">1–2</number>
)
<number box="[1359,1427,1472,1496]" pageId="23" pageNumber="324" value="16.5" valueMax="22.0" valueMin="11.0">11–22</number>
/
<number box="[151,210,1508,1532]" pageId="23" pageNumber="324" value="12.0" valueMax="17.0" valueMin="7.0">7–17</number>
.
</paragraph>
<paragraph blockId="23.[151,1437,1039,2037]" pageId="23" pageNumber="324">
<emphasis box="[199,449,1543,1568]" italics="true" pageId="23" pageNumber="324">Antennular peduncle</emphasis>
(
<figureCitation box="[480,579,1543,1568]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="23" pageNumber="324">
Fig.
<number box="[549,579,1544,1568]" pageId="23" pageNumber="324" value="13.0">13</number>
</figureCitation>
a):
<number box="[630,729,1543,1568]" pageId="23" pageNumber="324" value="0.675" valueMax="0.75" valueMin="0.6">0.6–0.75</number>
×carapace. Stylocerite
<number box="[1015,1100,1543,1568]" pageId="23" pageNumber="324" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7</number>
×length of basal segment. Anterolateral teeth of basal segment
<number box="[562,675,1580,1605]" pageId="23" pageNumber="324" value="0.2" valueMax="0.25" valueMin="0.15">0.15–0.25</number>
×second segment.
<number box="[882,938,1580,1604]" pageId="23" pageNumber="324" value="10.5" valueMax="12.0" valueMin="9.0">9–12</number>
segments bearing aesthetascs.
</paragraph>
<paragraph blockId="23.[151,1437,1039,2037]" pageId="23" pageNumber="324">
<emphasis box="[199,381,1615,1640]" italics="true" pageId="23" pageNumber="324">First pereiopod</emphasis>
(
<figureCitation box="[404,493,1615,1640]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="23" pageNumber="324">
Fig.
<number box="[464,493,1615,1640]" pageId="23" pageNumber="324" value="13.0">13</number>
</figureCitation>
b): Dactylus
<number box="[653,735,1615,1640]" pageId="23" pageNumber="324" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4</number>
×palm of propodus. Chela
<number box="[1068,1151,1615,1640]" pageId="23" pageNumber="324" value="2.0999999999999996" valueMax="2.3" valueMin="1.9">1.9–2.3</number>
×long as broad. Carpus
<number box="[151,235,1652,1677]" pageId="23" pageNumber="324" value="2.1" valueMax="2.4" valueMin="1.8">1.8–2.4</number>
×long as broad, with the anterior excavation.
</paragraph>
<paragraph blockId="23.[151,1437,1039,2037]" pageId="23" pageNumber="324">
<emphasis box="[199,404,1687,1712]" italics="true" pageId="23" pageNumber="324">Second pereiopod</emphasis>
(
<figureCitation box="[425,513,1687,1712]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="23" pageNumber="324">
Fig.
<number box="[483,513,1688,1712]" pageId="23" pageNumber="324" value="13.0">13</number>
</figureCitation>
c): Dactylus
<number box="[665,748,1687,1712]" pageId="23" pageNumber="324" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4</number>
×long as palm of propodus. Chela
<number box="[1168,1203,1687,1712]" pageId="23" pageNumber="324" value="2.8">2.8</number>
–3.0×long as broad. Carpus
<number box="[237,322,1724,1749]" pageId="23" pageNumber="324" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5</number>
×long as broad.
</paragraph>
<paragraph blockId="23.[151,1437,1039,2037]" pageId="23" pageNumber="324">
<emphasis box="[199,381,1759,1784]" italics="true" pageId="23" pageNumber="324">Third pereiopod</emphasis>
(
<figureCitation box="[400,483,1759,1784]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="23" pageNumber="324">
Fig.
<number box="[455,483,1759,1784]" pageId="23" pageNumber="324" value="13.0">13</number>
</figureCitation>
d, e): Dactylus 2.0–
<number box="[712,746,1759,1784]" pageId="23" pageNumber="324" value="2.3">2.3</number>
×long as broad.
<number box="[934,989,1759,1784]" pageId="23" pageNumber="324" value="8.5" valueMax="11.0" valueMin="6.0">6–11</number>
spines on dactylus (including terminal spines). Propodus 6.0–7.0×long as dactylus and
<number box="[723,791,1796,1821]" pageId="23" pageNumber="324" value="13.0" valueMax="14.0" valueMin="12.0">12–14</number>
×long as broad with
<number box="[1044,1100,1796,1820]" pageId="23" pageNumber="324" value="11.5" valueMax="14.0" valueMin="9.0">9–14</number>
spines arranged along inner margin. Carpus
<number box="[339,452,1831,1856]" pageId="23" pageNumber="324" value="0.6000000000000001" valueMax="0.65" valueMin="0.55">0.55–0.65</number>
×long as propodus, with
<number box="[747,761,1832,1856]" pageId="23" pageNumber="324" value="1.0">1</number>
large spine and
<number box="[963,1003,1831,1856]" pageId="23" pageNumber="324" value="3.5" valueMax="5.0" valueMin="2.0">2–5</number>
small spines on inner margin. Merus
<number box="[151,232,1868,1893]" pageId="23" pageNumber="324" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8</number>
×carpus length. Merus with
<number box="[550,564,1868,1892]" pageId="23" pageNumber="324" value="3.0">3</number>
large spines along posterior margin.
</paragraph>
<paragraph blockId="23.[151,1437,1039,2037]" pageId="23" pageNumber="324">
<emphasis box="[199,373,1903,1928]" italics="true" pageId="23" pageNumber="324">Fifth pereiopod</emphasis>
(
<figureCitation box="[387,466,1903,1928]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="23" pageNumber="324">
Fig.
<number box="[439,466,1903,1928]" pageId="23" pageNumber="324" value="13.0">13</number>
</figureCitation>
f, g): Dactylus
<number box="[634,668,1903,1928]" pageId="23" pageNumber="324" value="3.7">3.7</number>
–5.0×long as broad with
<number box="[947,1017,1903,1928]" pageId="23" pageNumber="324" value="52.5" valueMax="60.0" valueMin="45.0">45–60</number>
spines arranged in comb-like fashion on inner margin. Propodus
<number box="[462,530,1940,1965]" pageId="23" pageNumber="324" value="15.0" valueMax="16.0" valueMin="14.0">14–16</number>
×long as broad and
<number box="[760,845,1940,1965]" pageId="23" pageNumber="324" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5</number>
×long as dactylus with
<number box="[1110,1180,1940,1965]" pageId="23" pageNumber="324" value="12.5" valueMax="15.0" valueMin="10.0">10–15</number>
spines arranged along posterior margin. Carpus
<number box="[443,541,1976,2001]" pageId="23" pageNumber="324" value="0.525" valueMax="0.6" valueMin="0.45">0.45–0.6</number>
×propodus length and with one large spine and
<number box="[1090,1132,1976,2001]" pageId="23" pageNumber="324" value="4.5" valueMax="6.0" valueMin="3.0">3–6</number>
minute spines along inner margin. Merus
<number box="[322,403,2012,2037]" pageId="23" pageNumber="324" value="1.7" valueMax="1.9" valueMin="1.5">1.5–1.9</number>
×carpus length, with
<number box="[643,657,2012,2036]" pageId="23" pageNumber="324" value="3.0">3</number>
large spines along posterior margin.
</paragraph>
<caption httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="24" pageNumber="325" targetBox="[182,1408,261,1918]" targetPageId="24">
<paragraph blockId="24.[151,1436,1919,2033]" pageId="24" pageNumber="325">
<emphasis bold="true" box="[151,289,1919,1941]" pageId="24" pageNumber="325">FIGURE 13.</emphasis>
<taxonomicName box="[296,537,1919,1941]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="coulaborensis" status="sp. nov.">
<emphasis box="[296,537,1919,1941]" italics="true" pageId="24" pageNumber="325">
<taxonomicName box="[296,388,1919,1941]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="325" phylum="Arthropoda" rank="genus">Caridina</taxonomicName>
coulaborensis
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[544,626,1919,1940]" pageId="24" pageNumber="325" rank="species">
<emphasis bold="true" box="[544,626,1919,1940]" pageId="24" pageNumber="325">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype NMB 4.VIII.e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ovig.: i) Egg; j) Posterior margin of telson; NMB 4.VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson.
</paragraph>
</caption>
<paragraph blockId="25.[151,1437,151,1590]" box="[199,716,151,176]" pageId="25" pageNumber="326">
<emphasis box="[199,327,151,176]" italics="true" pageId="25" pageNumber="326">Setobranch</emphasis>
: With two setae on all pereiopods.
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" box="[199,1239,188,213]" pageId="25" pageNumber="326">
<emphasis box="[199,412,188,213]" italics="true" pageId="25" pageNumber="326">First male pleopod</emphasis>
(
<figureCitation box="[427,509,188,213]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="25" pageNumber="326">
Fig.
<number box="[480,509,188,213]" pageId="25" pageNumber="326" value="13.0">13</number>
</figureCitation>
h): Endopod
<number box="[654,767,188,213]" pageId="25" pageNumber="326" value="0.475" valueMax="0.5" valueMin="0.45">0.45–0.50</number>
×exopod length; appendix interna present.
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" box="[199,846,223,248]" pageId="25" pageNumber="326">
<emphasis box="[199,432,223,248]" italics="true" pageId="25" pageNumber="326">First female pleopod</emphasis>
: Endopod
<number box="[555,654,223,248]" pageId="25" pageNumber="326" value="0.525" valueMax="0.6" valueMin="0.45">0.45–0.6</number>
×long as exopod.
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" box="[199,879,260,285]" pageId="25" pageNumber="326">
<emphasis box="[199,255,261,284]" italics="true" pageId="25" pageNumber="326">Eggs</emphasis>
(
<figureCitation box="[270,353,260,285]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="25" pageNumber="326">
Fig.
<number box="[323,353,260,285]" pageId="25" pageNumber="326" value="13.0">
<date box="[323,353,260,285]" pageId="25" pageNumber="326" value="2013-01">13</date>
</number>
</figureCitation>
i): ca.
<specimenCount box="[421,507,260,285]" pageId="25" pageNumber="326" type="egg">
<number box="[421,449,260,285]" pageId="25" pageNumber="326" value="95.0">95</number>
eggs
</specimenCount>
of
<number box="[545,644,260,285]" pageId="25" pageNumber="326" value="0.625" valueMax="0.65" valueMin="0.6">0.6–0.65</number>
×
<quantity box="[658,822,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="25" pageNumber="326" unit="mm" value="0.39" valueMax="0.43" valueMin="0.35">0.35–0.43 mm</quantity>
size.
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" box="[199,1139,295,320]" pageId="25" pageNumber="326">
<emphasis box="[199,438,295,320]" italics="true" pageId="25" pageNumber="326">Second male pleopod</emphasis>
: appendix masculina
<number box="[683,715,295,320]" pageId="25" pageNumber="326" value="1.7">1.7</number>
–2.0×appendix interna,
<number box="[984,1020,295,320]" pageId="25" pageNumber="326" value="0.4">0.4</number>
×endopod.
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" box="[199,739,331,359]" pageId="25" pageNumber="326">
<number box="[199,213,334,359]" pageId="25" pageNumber="326" value="6.0">
<emphasis box="[199,213,334,359]" italics="true" pageId="25" pageNumber="326">6</emphasis>
</number>
<emphasis box="[213,225,331,345]" italics="true" pageId="25" pageNumber="326">th</emphasis>
<emphasis box="[232,431,334,359]" italics="true" pageId="25" pageNumber="326">abdominal somite</emphasis>
:
<number box="[446,531,334,359]" pageId="25" pageNumber="326" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7</number>
×long as carapace.
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" pageId="25" pageNumber="326">
<emphasis box="[199,269,373,398]" italics="true" pageId="25" pageNumber="326">Telson</emphasis>
(
<figureCitation box="[289,372,373,398]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="25" pageNumber="326">
Fig.
<number box="[344,372,373,398]" pageId="25" pageNumber="326" value="13.0">13</number>
</figureCitation>
j–l): 1.0–
<number box="[479,511,373,398]" pageId="25" pageNumber="326" value="1.1">1.1</number>
×long as
<number box="[621,635,373,398]" pageId="25" pageNumber="326" value="6.0">6</number>
th abdominal somite.
<number box="[871,913,374,398]" pageId="25" pageNumber="326" value="5.5" valueMax="7.0" valueMin="4.0">4–7</number>
pairs of dorsal spines (including subterminal spines). Posterior margin exhibits variations. It is mostly rounded without a median process rarely triangular with a median process. In specimens from Tiouaka (
<collectionCode box="[656,721,446,470]" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="25" pageNumber="326">NMB</collectionCode>
<date box="[721,784,446,470]" pageId="25" pageNumber="326">
<number box="[721,735,446,470]" pageId="25" pageNumber="326" value="4.0">4</number>
VIII
</date>
c), the telson tip is much broader in the females (
<figureCitation box="[1333,1412,445,470]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="25" pageNumber="326">
Fig.
<number box="[1385,1412,445,470]" pageId="25" pageNumber="326" value="13.0">13</number>
</figureCitation>
k) and comparatively narrower in the males (
<figureCitation box="[637,722,481,506]" captionStart="FIGURE 13" captionStartId="24.[151,250,1919,1941]" captionTargetBox="[182,1408,261,1918]" captionTargetId="figure@24.[178,1408,193,1932]" captionTargetPageId="24" captionText="FIGURE 13. Caridina coulaborensis sp. nov. Holotype NMB 4. VIII. e, ♂: a) Anterior region of the cepahalothorax; b) First pereiopod; c) Second pereiopod; d) Third pereiopod; e) Dactylus of third pereiopod; f) Fifth pereiopod; g) Dactylus of fifth pereiopod; h) Endopod of first male pleopod; Paratypes ♀ ovig.: i) Egg; j) Posterior margin of telson; NMB 4. VIIIc ♀: k) Posterior margin of telson; Holotype ♂: l) Posterior margin of telson." httpUri="https://zenodo.org/record/228256/files/figure.png" pageId="25" pageNumber="326">
Fig.
<number box="[692,722,481,506]" pageId="25" pageNumber="326" value="13.0">13</number>
</figureCitation>
l).
<number box="[753,767,482,506]" pageId="25" pageNumber="326" value="1.0">1</number>
pair of lateral spines and
<number box="[1069,1111,482,506]" pageId="25" pageNumber="326" value="2.5" valueMax="3.0" valueMin="2.0">2–3</number>
pairs of intermediate spines present. The intermediate spines are either slightly shorter, equal to or fractionally longer than the lateral spines.
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" box="[199,567,553,578]" pageId="25" pageNumber="326">
<emphasis box="[199,285,553,578]" italics="true" pageId="25" pageNumber="326">Uropod</emphasis>
:
<number box="[300,356,554,578]" pageId="25" pageNumber="326" value="11.5" valueMax="14.0" valueMin="9.0">9–14</number>
diaeresis spinules.
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" box="[199,576,589,614]" pageId="25" pageNumber="326">
<emphasis box="[199,341,589,614]" italics="true" pageId="25" pageNumber="326">Anal carina:</emphasis>
Armed with a spine.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="326" type="etymology">
<paragraph blockId="25.[151,1437,151,1590]" pageId="25" pageNumber="326">
<emphasis bold="true" box="[199,334,625,650]" pageId="25" pageNumber="326">Etymology.</emphasis>
<taxonomicName box="[344,500,625,650]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="coulaborensis">
<emphasis box="[344,500,625,650]" italics="true" pageId="25" pageNumber="326">coulaborensis</emphasis>
</taxonomicName>
: named for the
<typeStatus box="[691,739,626,650]" pageId="25" pageNumber="326">type</typeStatus>
locality in
<collectingCountry box="[876,1052,625,650]" name="New Caledonia" pageId="25" pageNumber="326">New Caledonia</collectingCountry>
being an abitory combination of Coula-Boréaré.
</paragraph>
</subSubSection>
<subSubSection box="[199,540,697,722]" pageId="25" pageNumber="326" type="distribution">
<paragraph blockId="25.[151,1437,151,1590]" box="[199,540,697,722]" pageId="25" pageNumber="326">
<emphasis bold="true" box="[199,352,697,722]" pageId="25" pageNumber="326">Distribution.</emphasis>
<collectingCountry box="[359,536,697,722]" name="New Caledonia" pageId="25" pageNumber="326">New Caledonia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,724,733,758]" pageId="25" pageNumber="326" type="materials_examined">
<paragraph blockId="25.[151,1437,151,1590]" box="[199,724,733,758]" pageId="25" pageNumber="326">
<emphasis bold="true" box="[199,357,733,758]" pageId="25" pageNumber="326">
<typeStatus box="[199,258,733,758]" pageId="25" pageNumber="326">Type</typeStatus>
locality.
</emphasis>
Coula-Boréaré,
<collectingCountry box="[543,720,733,758]" name="New Caledonia" pageId="25" pageNumber="326">New Caledonia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="326" type="discussion">
<paragraph blockId="25.[151,1437,151,1590]" pageId="25" pageNumber="326">
<emphasis bold="true" box="[199,315,769,794]" pageId="25" pageNumber="326">Remarks.</emphasis>
The specimens described by Roux as
<taxonomicName box="[746,920,769,794]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis box="[746,920,769,794]" italics="true" pageId="25" pageNumber="326">C. meridionalis</emphasis>
</taxonomicName>
from Coula-Boréaré, Tiouaka Tal, Koné, Lao Foa, Coindé, SW Nouméa &amp; Païta,
<collectingCountry box="[559,733,805,830]" name="New Caledonia" pageId="25" pageNumber="326">New Caledonia</collectingCountry>
were considered to be an undescribed species, named here as
<taxonomicName box="[151,415,842,867]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="coulaborensis" status="sp. nov.">
<emphasis box="[151,415,842,867]" italics="true" pageId="25" pageNumber="326">Caridina coulaborensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[422,515,841,867]" pageId="25" pageNumber="326" rank="species">
<emphasis bold="true" box="[422,515,841,867]" pageId="25" pageNumber="326">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" pageId="25" pageNumber="326">
<taxonomicName box="[199,470,878,903]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="coulaborensis" status="sp. nov.">
<emphasis box="[199,470,878,903]" italics="true" pageId="25" pageNumber="326">Caridina coulaborensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[484,581,878,903]" pageId="25" pageNumber="326" rank="species">
<emphasis bold="true" box="[484,581,878,903]" pageId="25" pageNumber="326">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<emphasis box="[750,966,878,903]" italics="true" pageId="25" pageNumber="326">
C. n.
<taxonomicName box="[825,966,878,903]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="meridionalis">meridionalis</taxonomicName>
</emphasis>
in having a rostrum with
<number box="[1298,1367,879,903]" pageId="25" pageNumber="326" value="16.5" valueMax="22.0" valueMin="11.0">11–22</number>
teeth proximally on the dorsal margin leaving
<number box="[609,707,915,940]" pageId="25" pageNumber="326" value="0.375" valueMax="0.45" valueMin="0.3">0.3–0.45</number>
unarmed distally (vs.
<number box="[958,1028,915,939]" pageId="25" pageNumber="326" value="23.0" valueMax="27.0" valueMin="19.0">19–27</number>
teeth on the dorsal margin to the tip in
<emphasis box="[183,393,950,975]" italics="true" pageId="25" pageNumber="326">
C. n.
<taxonomicName box="[251,393,950,975]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="meridionalis">meridionalis</taxonomicName>
</emphasis>
), tip of the rostrum bifid (vs. mostly tip pointed, rarely bifid in
<taxonomicName box="[1153,1331,950,975]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis box="[1153,1331,950,975]" italics="true" pageId="25" pageNumber="326">C. meridionalis</emphasis>
</taxonomicName>
), ventral margin of rostrum with
<number box="[421,477,987,1011]" pageId="25" pageNumber="326" value="12.0" valueMax="17.0" valueMin="7.0">7–17</number>
teeth (vs.
<number box="[595,651,987,1011]" pageId="25" pageNumber="326" value="7.0" valueMax="10.0" valueMin="4.0">4–10</number>
teeth in
<emphasis box="[750,953,987,1012]" italics="true" pageId="25" pageNumber="326">
C. n.
<taxonomicName box="[812,953,987,1012]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="meridionalis">meridionalis</taxonomicName>
</emphasis>
) and posterior margin of the telson mostly rounded rarely triangular with
<number box="[523,565,1023,1047]" pageId="25" pageNumber="326" value="2.5" valueMax="3.0" valueMin="2.0">2–3</number>
pairs of intermediate spines that are either slightly shorter, equal to or fractionally longer than the lateral spines (vs. triangular posterior margin converging to a median pointed process with
<number box="[208,222,1095,1119]" pageId="25" pageNumber="326" value="2.0">2</number>
pairs of intermediate spines that are slightly shorter than the lateral spines in
<emphasis box="[1089,1291,1094,1119]" italics="true" pageId="25" pageNumber="326">
C. n.
<taxonomicName box="[1150,1291,1094,1119]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="meridionalis">meridionalis</taxonomicName>
</emphasis>
).
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" pageId="25" pageNumber="326">
The proximal placement of teeth on the dorsal margin of the rostrum with a distal unarmed end in the new species is similar to that described for
<taxonomicName box="[603,799,1166,1191]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[603,799,1166,1191]" italics="true" pageId="25" pageNumber="326">Caridina nilotica</emphasis>
</taxonomicName>
(see
<bibRefCitation author="Richard &amp; Clark" box="[865,1128,1166,1191]" firstAuthor="Richard" pageId="25" pageNumber="326" refString="Richard, J. &amp; Clark P. F. (2005) Caridina nilotica (P. Roux, 1833) [Crustacea: Decapoda: Caridea: Atyidae] from East Africa with a description of four new species. Proceedings of the Biological Society of Washington, 118, 706 - 730." type="journal article" year="2005">
Richard &amp; Clark
<number box="[1068,1128,1166,1191]" pageId="25" pageNumber="326" value="2005.0">2005</number>
</bibRefCitation>
), but it differs in having a rounded or triangular posterior margin of the telson with long spines (vs. the posterior margin of the telson being almost flat and not protruding with
<number box="[557,599,1239,1263]" pageId="25" pageNumber="326" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
pairs of short stout spines present in
<taxonomicName box="[1026,1145,1238,1263]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[1026,1145,1238,1263]" italics="true" pageId="25" pageNumber="326">C. nilotica</emphasis>
</taxonomicName>
).
<taxonomicName box="[1170,1436,1238,1263]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="coulaborensis" status="sp. nov.">
<emphasis box="[1170,1436,1238,1263]" italics="true" pageId="25" pageNumber="326">Caridina coulaborensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[151,247,1275,1301]" pageId="25" pageNumber="326" rank="species">
<emphasis bold="true" box="[151,247,1275,1301]" pageId="25" pageNumber="326">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName box="[400,595,1275,1300]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[400,595,1275,1300]" italics="true" pageId="25" pageNumber="326">C. brachydactyla</emphasis>
</taxonomicName>
in having a rounded posterior margin of telson with or without a median process (vs. triangular posterior margin always ending in a median process in
<taxonomicName box="[1042,1236,1312,1337]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1042,1236,1312,1337]" italics="true" pageId="25" pageNumber="326">C. brachydactyla</emphasis>
</taxonomicName>
) and ca.
<number box="[1344,1372,1312,1337]" pageId="25" pageNumber="326" value="95.0">95</number>
large eggs of
<number box="[239,338,1348,1373]" pageId="25" pageNumber="326" value="0.625" valueMax="0.65" valueMin="0.6">0.6–0.65</number>
×
<quantity box="[353,516,1348,1373]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="25" pageNumber="326" unit="mm" value="0.39" valueMax="0.43" valueMin="0.35">0.35–0.43 mm</quantity>
size (vs. ca.
<number box="[661,703,1348,1373]" pageId="25" pageNumber="326" value="850.0">850</number>
small eggs
<number box="[835,947,1348,1373]" pageId="25" pageNumber="326" value="0.38" valueMax="0.41" valueMin="0.35">0.35–0.41</number>
×
<quantity box="[962,1112,1348,1373]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="25" pageNumber="326" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.2–0.23 mm</quantity>
in
<taxonomicName box="[1148,1340,1348,1373]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1148,1340,1348,1373]" italics="true" pageId="25" pageNumber="326">C. brachydactyla</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="25.[151,1437,151,1590]" pageId="25" pageNumber="326">
<taxonomicName box="[199,463,1385,1410]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="coulaborensis" status="sp. nov.">
<emphasis box="[199,463,1385,1410]" italics="true" pageId="25" pageNumber="326">Caridina coulaborensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[471,565,1384,1410]" pageId="25" pageNumber="326" rank="species">
<emphasis bold="true" box="[471,565,1384,1410]" pageId="25" pageNumber="326">sp. nov.</emphasis>
</taxonomicNameLabel>
differs distinctly from
<taxonomicName box="[826,934,1386,1409]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[826,934,1386,1409]" italics="true" pageId="25" pageNumber="326">C. simoni</emphasis>
</taxonomicName>
in possessing
<number box="[1098,1166,1385,1409]" pageId="25" pageNumber="326" value="16.5" valueMax="22.0" valueMin="11.0">11–22</number>
teeth proximally on the dorsal margin of rostrum leaving
<number box="[526,624,1421,1446]" pageId="25" pageNumber="326" value="0.375" valueMax="0.45" valueMin="0.3">0.3–0.45</number>
of the dorsal margin unarmed distally (vs.
<number box="[1107,1177,1421,1446]" pageId="25" pageNumber="326" value="20.0" valueMax="25.0" valueMin="15.0">15–25</number>
proximal teeth leaving distally
<number box="[244,342,1457,1482]" pageId="25" pageNumber="326" value="0.325" valueMax="0.4" valueMin="0.25">0.25–0.4</number>
unarmed or interrupted by
<number box="[667,710,1458,1482]" pageId="25" pageNumber="326" value="2.5" valueMax="4.0" valueMin="1.0">1–4</number>
teeth in
<taxonomicName box="[818,929,1458,1481]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[818,929,1458,1481]" italics="true" pageId="25" pageNumber="326">C. simoni</emphasis>
</taxonomicName>
), tip of rostrum bifid (vs. tip pointed in
<taxonomicName class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="simoni">
<emphasis italics="true" pageId="25" pageNumber="326">C. simoni</emphasis>
</taxonomicName>
), ventral margin of rostrum with
<number box="[611,667,1494,1518]" pageId="25" pageNumber="326" value="12.0" valueMax="17.0" valueMin="7.0">7–17</number>
teeth proximally to the tip or with a short distally unarmed margin (vs. with
<number box="[263,319,1529,1554]" pageId="25" pageNumber="326" value="9.5" valueMax="14.0" valueMin="5.0">5–14</number>
teeth proximally leaving the distal ventral margin always unarmed in
<taxonomicName box="[1145,1256,1530,1553]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[1145,1256,1530,1553]" italics="true" pageId="25" pageNumber="326">C. simoni</emphasis>
</taxonomicName>
) and in having comparatively smaller eggs of
<number box="[496,595,1565,1590]" pageId="25" pageNumber="326" value="0.625" valueMax="0.65" valueMin="0.6">0.6–0.65</number>
×
<quantity box="[609,772,1565,1590]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="25" pageNumber="326" unit="mm" value="0.39" valueMax="0.43" valueMin="0.35">0.35–0.43 mm</quantity>
size (vs.
<number box="[878,927,1565,1590]" pageId="25" pageNumber="326" value="0.65">0.65</number>
–1.0×
<quantity box="[990,1141,1565,1590]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="25" pageNumber="326" unit="mm" value="0.525" valueMax="0.6" valueMin="0.45">0.45–0.6 mm</quantity>
in
<taxonomicName box="[1176,1284,1566,1589]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[1176,1284,1566,1589]" italics="true" pageId="25" pageNumber="326">C. simoni</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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