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<document id="D7E1F8D0D51F10B96031CEE464F548E6" ID-DOI="10.11646/zootaxa.1575.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5096833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626202096956" checkinUser="felipe" docAuthor="Chaverri, Luis Guillermo &amp; Borkent, Art" docDate="2007" docId="03A187C6FF90FF914EF8FAD6FB2BFC9A" docLanguage="en" docName="zt01575p034.pdf" docOrigin="Zootaxa 1575 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1575.1.1" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Dixella woodi Chaverri and Borkent 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="24" masterDocId="FF98FFBEFF84FF864E6FFFA4FFEFFF90" masterDocTitle="The Meniscus midges of Costa Rica (Diptera: Dixidae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="21" updateTime="1699003511584" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="77424CFAA3BC9CC84C6E54498E495A64" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0FA24C0A1323853DF68827D3660D34B3">
<mods:title id="95D61843D018663A000BCD032FAE8DF1">The Meniscus midges of Costa Rica (Diptera: Dixidae)</mods:title>
</mods:titleInfo>
<mods:name id="7A6C0797E6270FE5FEA19D7FB3C0C976" type="personal">
<mods:role id="20D21372EC1A903E77B2F22903C15800">
<mods:roleTerm id="54116DDC335BABC7AC44AE699CD53402">Author</mods:roleTerm>
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<mods:namePart id="4B921530631A0B9AFA98099EB2C5D606">Chaverri, Luis Guillermo</mods:namePart>
<mods:affiliation id="CA37C4021B858F5D7A7F0A3B6B3462A7">Instituto Nacional de Biodiversidad, 400 Mts Oeste del Cementerio, A. P. 22 - 3100 Santo Domingo de Heredia, Costa Rica. E-mail: lchaverr @ inbio. ac. cr Research Associate, Royal British Columbia Museum, American Museum of Natural History, and Instituto Nacional de Biodiversidad, 691 - 8 th Ave. SE, Salmon Arm, British Columbia, V 1 E 2 C 2, Canada. E-mail: aborkent @ sunlite. ca</mods:affiliation>
</mods:name>
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<mods:role id="C83A47FA56092CD92F5A295F1F9102CF">
<mods:roleTerm id="C8A2E7008C7C3A27B23BA24E85C176D5">Author</mods:roleTerm>
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<mods:namePart id="76D8E9965E840DA1C61945882830CBDA">Borkent, Art</mods:namePart>
</mods:name>
<mods:typeOfResource id="CC95D34292E0C35442DBE2AB1A60BF2B">text</mods:typeOfResource>
<mods:relatedItem id="4416B9C587BFCA99B0F6C6A57E316EF3" type="host">
<mods:titleInfo id="40931128816E28970B2EA5B2191B788E">
<mods:title id="E853E0F4F0F8DA2037470529F67274F5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D3F1C065EA3F5D7807ED5CF1EDF292FA">
<mods:date id="1D952D909A792CE3F52582C8D30B934B">2007</mods:date>
<mods:detail id="2A3091C711187B063BFE53C266A43662" type="pubDate">
<mods:number id="FE058BB0168AD267C176FA8DE25740DD">2007-09-05</mods:number>
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<mods:detail id="0CFCE5DC1D6051ACE79CA37A195D96F5" type="volume">
<mods:number id="3EA4F6E093B31E5C7F7EE955B89693F5">1575</mods:number>
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<mods:detail id="4CD00628FBF150422579C8752487AEF1" type="issue">
<mods:number id="BB30B7123D5307B5A0D36FB2D11EB231">1</mods:number>
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<mods:url id="8397761C6C6CE4AFA310173391F600B7">https://biotaxa.org/Zootaxa/article/view/zootaxa.1575.1.1</mods:url>
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<mods:classification id="826B1AAFF8CF0CB809C7A00B259BD8FE">journal article</mods:classification>
<mods:identifier id="B7E1953C1ACBAAC8334A10604C91459F" type="DOI">10.11646/zootaxa.1575.1.1</mods:identifier>
<mods:identifier id="D2B04EF2E1BE293A498CD8469CF1013B" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="94CC51120643C1EBBA9BF77A2D2E9708" type="Zenodo-Dep">5096833</mods:identifier>
<mods:identifier id="595BF8897626A06AF69973AE8C2E3F82" type="ZooBank">6D448CFC-0B62-46C0-92BF-24629AE3FCE0</mods:identifier>
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<treatment id="03A187C6FF90FF914EF8FAD6FB2BFC9A" LSID="urn:lsid:plazi:treatment:03A187C6FF90FF914EF8FAD6FB2BFC9A" httpUri="http://treatment.plazi.org/id/03A187C6FF90FF914EF8FAD6FB2BFC9A" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
<subSubSection id="C312655BFF90FF924EF8FAD6FD4BFA1C" box="[151,676,1394,1420]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BB736D0FF90FF924EF8FAD6FD4BFA1C" blockId="20.[151,676,1394,1455]" box="[151,676,1394,1420]" pageId="20" pageNumber="21">
<heading id="D0FF81BCFF90FF924EF8FAD6FD4BFA1C" bold="true" box="[151,676,1394,1420]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B97CEAC2FF90FF924EF8FAD6FD4BFA1C" bold="true" box="[151,676,1394,1420]" pageId="20" pageNumber="21">
<taxonomicName id="4C084D53FF90FF924EF8FAD6FDB5FA1C" ID-CoL="3726N" authority="Chaverri and Borkent" authorityName="Chaverri and Borkent" authorityYear="2007" box="[151,602,1394,1420]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="woodi" status="sp. nov.">
<emphasis id="B97CEAC2FF90FF924EF8FAD6FED1FA1C" bold="true" box="[151,318,1394,1420]" italics="true" pageId="20" pageNumber="21">Dixella woodi</emphasis>
Chaverri and Borkent
</taxonomicName>
<taxonomicNameLabel id="A24F57B9FF90FF924C0DFAD6FD4BFA1C" box="[610,676,1394,1420]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C312655BFF90FF924EF8FA31FD9AFA3F" box="[151,629,1429,1455]" pageId="20" pageNumber="21" type="description">
<paragraph id="8BB736D0FF90FF924EF8FA31FD9AFA3F" blockId="20.[151,676,1394,1455]" box="[151,629,1429,1455]" pageId="20" pageNumber="21">
(
<figureCitation id="13332A55FF90FF924EF0FA31FECAFA3F" box="[159,293,1429,1455]" captionStart="FIGURE 1" captionStartId="4.[151,255,1898,1922]" captionTargetBox="[168,1416,300,1854]" captionTargetId="figure-91@4.[151,1436,277,1872]" captionTargetPageId="4" captionText="FIGURE 1. Head capsules of male Dixella, anterior view. (A) D. maculata; (B) D. hernandezi; (C) D. shannoni; (D) D. hansoni; (E) D. fernandezae; (F) D. jironi; (G) D. venezuelensis; (H) D. woodi; (I) D. lobata; (J) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096835" httpUri="https://zenodo.org/record/5096835/files/figure.png" pageId="20" pageNumber="21">Figures 1H</figureCitation>
,
<figureCitation id="13332A55FF90FF924F5DFA31FEB8FA3F" box="[306,343,1429,1455]" captionStart="FIGURE 3" captionStartId="7.[151,255,2001,2025]" captionTargetBox="[223,1380,216,1927]" captionTargetId="figure-42@7.[151,1436,144,1977]" captionTargetPageId="7" captionText="FIGURE 3. Thoraces of male Dixella, dorsal view. (A) D. venezuelensis; (B) D. woodi; (C) D. lobata; (D) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096839" httpUri="https://zenodo.org/record/5096839/files/figure.png" pageId="20" pageNumber="21">3B</figureCitation>
,
<figureCitation id="13332A55FF90FF924F0CFA31FE68FA3F" box="[355,391,1429,1455]" captionStart="FIGURE 5" captionStartId="9.[151,255,1696,1720]" captionTargetBox="[196,1384,318,1669]" captionTargetId="figure-141@9.[151,1436,268,1714]" captionTargetPageId="9" captionText="FIGURE 5. Thoraces of male Dixella, lateral view. (A) D. venezuelensis; (B) D. woodi; (C) D. lobata; (D) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096843" httpUri="https://zenodo.org/record/5096843/files/figure.png" pageId="20" pageNumber="21">5B</figureCitation>
,
<figureCitation id="13332A55FF90FF924FFBFA31FE55FA3F" box="[404,442,1429,1455]" captionStart="FIGURE 6" captionStartId="12.[151,255,1607,1631]" captionTargetBox="[229,1364,197,1574]" captionTargetId="figure-139@12.[160,1427,128,1623]" captionTargetPageId="12" captionText="FIGURE 6. Wings of male Dixella. (A) D. maculata; (B) D. hernandezi; (C) D. shannoni; (D) D. hansoni; (E) D. fernandezae; (F) D. jironi; (G) D. venezuelensis; (H) D. woodi; (I) D. lobata; (J) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096845" httpUri="https://zenodo.org/record/5096845/files/figure.png" pageId="20" pageNumber="21">6H</figureCitation>
,
<figureCitation id="13332A55FF90FF924FA9FA31FE02FA3F" box="[454,493,1429,1455]" captionStart="FIGURE 9" captionStartId="16.[151,255,1514,1538]" captionTargetBox="[204,1380,315,1432]" captionTargetId="figure-202@16.[152,1436,264,1488]" captionTargetPageId="16" captionText="FIGURE 9. Claws of male Dixella. (A) D. woodi; (B) D. lobata; (C) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096851" httpUri="https://zenodo.org/record/5096851/files/figure.png" pageId="20" pageNumber="21">9A</figureCitation>
,
<figureCitation id="13332A55FF90FF924F96FA31FDC3FA3F" box="[505,556,1429,1455]" captionStart="FIGURE 13" captionStartId="22.[151,255,1815,1839]" captionTargetBox="[190,1380,245,1750]" captionTargetId="figure-101@22.[159,1427,194,1791]" captionTargetPageId="22" captionText="FIGURE 13. Genitalia of male Dixella, dorsal view. (A) D. venezuelensis; (B) D. woodi." figureDoi="http://doi.org/10.5281/zenodo.5096859" httpUri="https://zenodo.org/record/5096859/files/figure.png" pageId="20" pageNumber="21">13B</figureCitation>
,
<figureCitation id="13332A55FF90FF924C57FA31FD83FA3F" box="[568,620,1429,1455]" captionStart="FIGURE 19" captionStartId="31.[151,255,1961,1985]" captionTargetBox="[234,1362,204,1945]" captionTargetId="figure-39@31.[224,1377,194,1955]" captionTargetPageId="31" captionText="FIGURE 19. Distributions of Dixella species in Costa Rica: (A) D. woodi—O, D. lobata—#; (B) D. venezuelensis—O, D. suzukii—#." figureDoi="http://doi.org/10.5281/zenodo.5096872" httpUri="https://zenodo.org/record/5096872/files/figure.png" pageId="20" pageNumber="21">19A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C312655BFF90FF924EF8FA41FD2EF9DF" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="8BB736D0FF90FF924EF8FA41FD2EF9DF" blockId="20.[151,1436,1509,2021]" pageId="20" pageNumber="21">
<emphasis id="B97CEAC2FF90FF924EF8FA41FEF8FA6F" bold="true" box="[151,279,1509,1535]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
<emphasis id="B97CEAC2FF90FF924F4BFA42FE8FFA6F" box="[292,352,1510,1535]" italics="true" pageId="20" pageNumber="21">Male</emphasis>
: only species of
<taxonomicName id="4C084D53FF90FF924C4BFA41FD6EFA6F" authorityName="Chaverri &amp; Borkent" authorityYear="2007" box="[548,641,1509,1535]" class="Insecta" family="Dixidae" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Dixidae</taxonomicName>
in Central America with elongate, dark acrostichal and dorsocentral setae (
<figureCitation id="13332A55FF90FF924E8EF9A9FEA7F9B7" box="[225,328,1549,1575]" captionStart="FIGURE 3" captionStartId="7.[151,255,2001,2025]" captionTargetBox="[223,1380,216,1927]" captionTargetId="figure-42@7.[151,1436,144,1977]" captionTargetPageId="7" captionText="FIGURE 3. Thoraces of male Dixella, dorsal view. (A) D. venezuelensis; (B) D. woodi; (C) D. lobata; (D) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096839" httpUri="https://zenodo.org/record/5096839/files/figure.png" pageId="20" pageNumber="21">Figs. 3B</figureCitation>
;
<figureCitation id="13332A55FF90FF924F3AF9A9FE95F9B7" box="[341,378,1549,1575]" captionStart="FIGURE 5" captionStartId="9.[151,255,1696,1720]" captionTargetBox="[196,1384,318,1669]" captionTargetId="figure-141@9.[151,1436,268,1714]" captionTargetPageId="9" captionText="FIGURE 5. Thoraces of male Dixella, lateral view. (A) D. venezuelensis; (B) D. woodi; (C) D. lobata; (D) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096843" httpUri="https://zenodo.org/record/5096843/files/figure.png" pageId="20" pageNumber="21">5B</figureCitation>
). Also, only species in this region with setae on the katepisternum (
<figureCitation id="13332A55FF90FF924AF6F9A9FB1DF9B7" box="[1177,1266,1549,1575]" captionStart="FIGURE 5" captionStartId="9.[151,255,1696,1720]" captionTargetBox="[196,1384,318,1669]" captionTargetId="figure-141@9.[151,1436,268,1714]" captionTargetPageId="9" captionText="FIGURE 5. Thoraces of male Dixella, lateral view. (A) D. venezuelensis; (B) D. woodi; (C) D. lobata; (D) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096843" httpUri="https://zenodo.org/record/5096843/files/figure.png" pageId="20" pageNumber="21">Fig. 5B</figureCitation>
) (but see Taxonomic Discussion below).
<emphasis id="B97CEAC2FF90FF924F8AF992FDD2F9DF" box="[485,573,1590,1615]" italics="true" pageId="20" pageNumber="21">Female</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C312655BFF90FF934EA9F9F9FF71FE62" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="8BB736D0FF90FF934EA9F9F9FACCFE32" blockId="20.[151,1436,1509,2021]" lastBlockId="21.[151,1436,152,498]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="B97CEAC2FF90FF924EA9F9F9FEB1F9E7" bold="true" box="[198,350,1629,1655]" pageId="20" pageNumber="21">Description.</emphasis>
<emphasis id="B97CEAC2FF90FF924F1CF9FAFE40F9E7" box="[371,431,1630,1655]" italics="true" pageId="20" pageNumber="21">Male</emphasis>
.
<emphasis id="B97CEAC2FF90FF924FAEF9F9FDEBF9E7" bold="true" box="[449,516,1629,1655]" pageId="20" pageNumber="21">Head</emphasis>
(
<figureCitation id="13332A55FF90FF924C79F9F9FD99F9E7" box="[534,630,1629,1655]" captionStart="FIGURE 1" captionStartId="4.[151,255,1898,1922]" captionTargetBox="[168,1416,300,1854]" captionTargetId="figure-91@4.[151,1436,277,1872]" captionTargetPageId="4" captionText="FIGURE 1. Head capsules of male Dixella, anterior view. (A) D. maculata; (B) D. hernandezi; (C) D. shannoni; (D) D. hansoni; (E) D. fernandezae; (F) D. jironi; (G) D. venezuelensis; (H) D. woodi; (I) D. lobata; (J) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096835" httpUri="https://zenodo.org/record/5096835/files/figure.png" pageId="20" pageNumber="21">Fig. 1H</figureCitation>
): brown, densely spiculate. Clypeus brown, 1.22 longer than basal width. Antennal scape brown, pedicel dark brown, flagellomeres light brown.
<emphasis id="B97CEAC2FF90FF924A59F921FB7CF90F" bold="true" box="[1078,1171,1669,1695]" pageId="20" pageNumber="21">Thorax</emphasis>
(
<figureCitation id="13332A55FF90FF924ACCF921FB10F90F" box="[1187,1279,1669,1695]" captionStart="FIGURE 3" captionStartId="7.[151,255,2001,2025]" captionTargetBox="[223,1380,216,1927]" captionTargetId="figure-42@7.[151,1436,144,1977]" captionTargetPageId="7" captionText="FIGURE 3. Thoraces of male Dixella, dorsal view. (A) D. venezuelensis; (B) D. woodi; (C) D. lobata; (D) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096839" httpUri="https://zenodo.org/record/5096839/files/figure.png" pageId="20" pageNumber="21">Fig. 3B</figureCitation>
,
<figureCitation id="13332A55FF90FF924B63F921FADEF90F" box="[1292,1329,1669,1695]" captionStart="FIGURE 5" captionStartId="9.[151,255,1696,1720]" captionTargetBox="[196,1384,318,1669]" captionTargetId="figure-141@9.[151,1436,268,1714]" captionTargetPageId="9" captionText="FIGURE 5. Thoraces of male Dixella, lateral view. (A) D. venezuelensis; (B) D. woodi; (C) D. lobata; (D) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096843" httpUri="https://zenodo.org/record/5096843/files/figure.png" pageId="20" pageNumber="21">5B</figureCitation>
): scutum with brown medial vittae extending from anterior margin to slightly beyond middle of scutum, divided longitudinally by yellow strip that continues posteriorly in prescutellar area as brown strip; lateral vitta extending from supraalar area to prescutal suture; acrostichal, dorsocentral setae elongate, dark; area near prescutal suture, antealar area with scattered setae; scutellum with setae along anterior margin. Scutellum, mediotergite light brown. Pleura with katepisternum, posterior anepisternum, anepimeron light brown, rest of pleura yellowish; scattered setae on upper katepisternum.
<emphasis id="B97CEAC2FF90FF924C8EF8D1FCCAF81F" bold="true" box="[737,805,1909,1935]" pageId="20" pageNumber="21">Wing</emphasis>
(
<figureCitation id="13332A55FF90FF924D58F8D1FC78F81F" box="[823,919,1909,1935]" captionStart="FIGURE 6" captionStartId="12.[151,255,1607,1631]" captionTargetBox="[229,1364,197,1574]" captionTargetId="figure-139@12.[160,1427,128,1623]" captionTargetPageId="12" captionText="FIGURE 6. Wings of male Dixella. (A) D. maculata; (B) D. hernandezi; (C) D. shannoni; (D) D. hansoni; (E) D. fernandezae; (F) D. jironi; (G) D. venezuelensis; (H) D. woodi; (I) D. lobata; (J) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096845" httpUri="https://zenodo.org/record/5096845/files/figure.png" pageId="20" pageNumber="21">Fig. 6H</figureCitation>
;
<tableCitation id="C68A036BFF90FF924DCAF8D1FC11F81F" box="[933,1022,1909,1935]" captionStart="TABLE 1" captionStartId="5.[151,241,151,175]" captionText="TABLE 1. Length of wings of male Dixella in Costa Rica (in mm)" pageId="20" pageNumber="21">Table 1</tableCitation>
): without dark spot over r-m; R2+3 originating at or distal to r-m; stem of R2+3 0.30 length of R3.
<emphasis id="B97CEAC2FF90FF924D3BF804FC4AF82A" bold="true" box="[852,933,1952,1978]" pageId="20" pageNumber="21">Halter</emphasis>
: pale, 0.40 length of fore femur.
<emphasis id="B97CEAC2FF90FF924B4AF804FA8FF82A" bold="true" box="[1317,1376,1952,1978]" pageId="20" pageNumber="21">Legs</emphasis>
: yellowish; claws as figured (
<figureCitation id="13332A55FF90FF924FA0F86FFDDFF875" box="[463,560,1995,2021]" captionStart="FIGURE 9" captionStartId="16.[151,255,1514,1538]" captionTargetBox="[204,1380,315,1432]" captionTargetId="figure-202@16.[152,1436,264,1488]" captionTargetPageId="16" captionText="FIGURE 9. Claws of male Dixella. (A) D. woodi; (B) D. lobata; (C) D. suzukii." figureDoi="http://doi.org/10.5281/zenodo.5096851" httpUri="https://zenodo.org/record/5096851/files/figure.png" pageId="20" pageNumber="21">Fig. 9A</figureCitation>
).
<emphasis id="B97CEAC2FF90FF924C27F86FFD53F875" bold="true" box="[584,700,1995,2021]" pageId="20" pageNumber="21">Genitalia</emphasis>
(
<figureCitation id="13332A55FF90FF924CA0F86FFCD1F875" box="[719,830,1995,2021]" captionStart="FIGURE 13" captionStartId="22.[151,255,1815,1839]" captionTargetBox="[190,1380,245,1750]" captionTargetId="figure-101@22.[159,1427,194,1791]" captionTargetPageId="22" captionText="FIGURE 13. Genitalia of male Dixella, dorsal view. (A) D. venezuelensis; (B) D. woodi." figureDoi="http://doi.org/10.5281/zenodo.5096859" httpUri="https://zenodo.org/record/5096859/files/figure.png" pageId="20" pageNumber="21">Fig. 13B</figureCitation>
): tergite 9 with anterior, posterior margins nearly straight, with numerous scattered setae. Sternite 9 with posterior margin lacking narrow excavation, with elongate scattered lateral, sublateral setae, bare medially. Gonocoxite wider than long; basal lobe not visible; apical lobe as long, as wide as gonostylus, truncate apically with anteromedial margin slightly projecting, with elongate basal seta, one elongate, two short subapical setae, a brown basal projection as a beak and a strong subbasal seta. Gonostylus with dense setae, somewhat swollen subapically, apex sharply tapered. Parameres, aedeagus not identified with certainty; anterior internal structures as figured. Tergite 10 broad, tapering gradully posteriorly, with minute and elongate spicules, posterior margin with two submedial setae.
</paragraph>
<paragraph id="8BB736D0FF91FF934EA9FE14FDB2FE5A" blockId="21.[151,1436,152,498]" box="[198,605,432,458]" pageId="21" pageNumber="22">
<emphasis id="B97CEAC2FF91FF934EA9FE14FE3BFE59" box="[198,468,432,457]" italics="true" pageId="21" pageNumber="22">Female and immatures</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BB736D0FF91FF934EF8FE7CFF71FE62" blockId="21.[151,1436,152,498]" box="[151,158,472,498]" pageId="21" pageNumber="22">.</paragraph>
</subSubSection>
<caption id="DF776658FF91FF934EF8F868FBAFF874" ID-DOI="http://doi.org/10.5281/zenodo.5096857" ID-Zenodo-Dep="5096857" box="[151,1088,1996,2020]" httpUri="https://zenodo.org/record/5096857/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,255,1996,2020]" targetBox="[214,1361,502,1969]" targetPageId="21">
<paragraph id="8BB736D0FF91FF934EF8F868FBAFF874" blockId="21.[151,1088,1996,2020]" box="[151,1088,1996,2020]" pageId="21" pageNumber="22">
<emphasis id="B97CEAC2FF91FF934EF8F868FEC9F874" bold="true" box="[151,294,1996,2020]" pageId="21" pageNumber="22">FIGURE 12.</emphasis>
Genitalia of male
<taxonomicName id="4C084D53FF91FF934F9BF869FDADF874" box="[500,578,1997,2020]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAC2FF91FF934F9BF869FDADF874" box="[500,578,1997,2020]" italics="true" pageId="21" pageNumber="22">Dixella</emphasis>
</taxonomicName>
, dorsal view. (A)
<taxonomicName id="4C084D53FF91FF934D6DF869FC4AF874" authorityName="Chaverri and Borkent" authorityYear="2007" box="[770,933,1997,2020]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fernandezae">
<emphasis id="B97CEAC2FF91FF934D6DF869FC4AF874" box="[770,933,1997,2020]" italics="true" pageId="21" pageNumber="22">D. fernandezae</emphasis>
</taxonomicName>
; (B)
<taxonomicName id="4C084D53FF91FF934DB2F869FBD3F874" authorityName="Chaverri and Borkent" authorityYear="2007" box="[989,1084,1997,2020]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="jironi">D. jironi</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF776658FF92FF904EF8F8B3FBBAF8BF" ID-DOI="http://doi.org/10.5281/zenodo.5096859" ID-Zenodo-Dep="5096859" box="[151,1109,1815,1839]" httpUri="https://zenodo.org/record/5096859/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,255,1815,1839]" targetBox="[190,1380,245,1750]" targetPageId="22">
<paragraph id="8BB736D0FF92FF904EF8F8B3FBBAF8BF" blockId="22.[151,1109,1815,1839]" box="[151,1109,1815,1839]" pageId="22" pageNumber="23">
<emphasis id="B97CEAC2FF92FF904EF8F8B3FEC9F8BF" bold="true" box="[151,294,1815,1839]" pageId="22" pageNumber="23">FIGURE 13.</emphasis>
Genitalia of male
<taxonomicName id="4C084D53FF92FF904F9BF8BCFDADF8BF" box="[500,578,1816,1839]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEAC2FF92FF904F9BF8BCFDADF8BF" box="[500,578,1816,1839]" italics="true" pageId="22" pageNumber="23">Dixella</emphasis>
</taxonomicName>
, dorsal view. (A)
<taxonomicName id="4C084D53FF92FF904D6DF8BCFC5CF8BF" box="[770,947,1816,1839]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B97CEAC2FF92FF904D6DF8BCFC5CF8BF" box="[770,947,1816,1839]" italics="true" pageId="22" pageNumber="23">D. venezuelensis</emphasis>
</taxonomicName>
; (B)
<taxonomicName id="4C084D53FF92FF904D83F8BCFBA2F8BF" authorityName="Chaverri and Borkent" authorityYear="2007" box="[1004,1101,1816,1839]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="woodi">
<emphasis id="B97CEAC2FF92FF904D83F8BCFBA2F8BF" box="[1004,1101,1816,1839]" italics="true" pageId="22" pageNumber="23">D. woodi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C312655BFF92FF914EA9F8C6FB2BFC9A" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BB736D0FF92FF904EA9F8C6FB23F864" blockId="22.[151,1436,1890,2036]" pageId="22" pageNumber="23">
<emphasis id="B97CEAC2FF92FF904EA9F8C6FDCDF8EC" bold="true" box="[198,546,1890,1916]" pageId="22" pageNumber="23">Distribution and bionomics.</emphasis>
This species is recorded from two localities in
<collectingCountry id="F31F7640FF92FF904A20F8C6FB3DF8EC" box="[1103,1234,1890,1916]" name="Costa Rica" pageId="22" pageNumber="23">Costa Rica</collectingCountry>
, at altitudes from
<quantity id="4CF09B35FF92FF904EF8F82EFEC1F834" box="[151,302,1930,1956]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="13.6" metricValueMin="3.4" pageId="22" pageNumber="23" unit="m" value="850.0" valueMax="1360.0" valueMin="340.0">340–1360 m</quantity>
(
<figureCitation id="13332A55FF92FF904F2EF82EFE41F834" box="[321,430,1930,1956]" captionStart="FIGURE 19" captionStartId="31.[151,255,1961,1985]" captionTargetBox="[234,1362,204,1945]" captionTargetId="figure-39@31.[224,1377,194,1955]" captionTargetPageId="31" captionText="FIGURE 19. Distributions of Dixella species in Costa Rica: (A) D. woodi—O, D. lobata—#; (B) D. venezuelensis—O, D. suzukii—#." figureDoi="http://doi.org/10.5281/zenodo.5096872" httpUri="https://zenodo.org/record/5096872/files/figure.png" pageId="22" pageNumber="23">Fig. 19A</figureCitation>
) during the dry season (February). Habitats include Premontane Moist Forest and Tropical Lower Montane Forest. The
<typeStatus id="54B38872FF92FF904C07F816FD22F85C" box="[616,717,1970,1996]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
was swept with an aerial net along the margin of a narrow stream where pools had formed and were collected together with adults of
<taxonomicName id="4C084D53FF92FF904A6DF87FFB2AF864" box="[1026,1221,2011,2036]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="B97CEAC2FF92FF904A6DF87FFB2AF864" box="[1026,1221,2011,2036]" italics="true" pageId="22" pageNumber="23">D. venezuelensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB736D0FF93FF914EA9FF33FA73FE5A" blockId="23.[151,1436,151,778]" pageId="23" pageNumber="24">
<emphasis id="B97CEAC2FF93FF914EA9FF33FE31FF21" bold="true" box="[198,478,151,177]" pageId="23" pageNumber="24">Taxonomic discussion.</emphasis>
A female collected at
<collectingCountry id="F31F7640FF93FF914C85FF3CFC81FF22" box="[746,878,152,178]" name="Costa Rica" pageId="23" pageNumber="24">Costa Rica</collectingCountry>
,
<collectingRegion id="49CCF832FF93FF914D14FF3CFC10FF22" box="[891,1023,152,178]" country="Costa Rica" name="Puntarenas" pageId="23" pageNumber="24">Puntarenas</collectingRegion>
,
<collectingRegion id="49CCF832FF93FF914A65FF3CFB61FF22" box="[1034,1166,152,178]" country="Costa Rica" name="Puntarenas" pageId="23" pageNumber="24">Puntarenas</collectingRegion>
, Monteverde, 2 Km E.
<collectingCountry id="F31F7640FF93FF914EF8FF64FECDFF4A" box="[151,290,192,218]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="23" pageNumber="24">Santa Elena</collectingCountry>
(
<quantity id="4CF09B35FF93FF914F5BFF64FE7EFF4A" box="[308,401,192,218]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="23" pageNumber="24" unit="m" value="1540.0">1540 m</quantity>
,
<date id="FFB61010FF93FF914FF0FF64FDF2FF4A" box="[415,541,192,218]" pageId="23" pageNumber="24" value="2000-06-02">2-VI-2000</date>
, J.M. Cumming, CD 5084; INBC) was initially thought to be the female of
<taxonomicName id="4C084D53FF93FF914EF8FF4CFEEDFE91" authorityName="Chaverri and Borkent" authorityYear="2007" box="[151,258,232,258]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="woodi">
<emphasis id="B97CEAC2FF93FF914EF8FF4CFF43FE91" box="[151,172,232,257]" italics="true" pageId="23" pageNumber="24">D</emphasis>
.
<emphasis id="B97CEAC2FF93FF914ED4FF4CFEEDFE91" box="[187,258,232,257]" italics="true" pageId="23" pageNumber="24">woodi</emphasis>
</taxonomicName>
because it had a thorax with a similarly distinctive setation (including on the katepisternum). However, the clypeus of this female was short, suggesting the presence of a second Costa Rican species with setae on the katepisternum (otherwise unique in this area). A second female we examined, from
<collectingCountry id="F31F7640FF93FF914AA7FE9CFAD0FEC2" box="[1224,1343,312,338]" name="Colombia" pageId="23" pageNumber="24">Colombia</collectingCountry>
,
<collectingRegion id="49CCF832FF93FF914B25FE9CFF12FEEA" country="Colombia" name="Cundinamarca" pageId="23" pageNumber="24">Cundinamarca</collectingRegion>
, Finca Bella Vista, near Sasaima (
<date id="FFB61010FF93FF914CFEFEC4FCF3FEEA" box="[657,796,352,378]" pageId="23" pageNumber="24" value="1965-04-19">19-IV-1965</date>
, P.R. Craig; CAS), had katepisternal setae and a moderately elongate clypeus and may represent the female of
<taxonomicName id="4C084D53FF93FF914D2EFE2CFC42FE31" authorityName="Chaverri and Borkent" authorityYear="2007" box="[833,941,392,417]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="woodi">
<emphasis id="B97CEAC2FF93FF914D2EFE2CFC42FE31" box="[833,941,392,417]" italics="true" pageId="23" pageNumber="24">D. woodi</emphasis>
</taxonomicName>
. However, the distance between the
<typeStatus id="54B38872FF93FF914B06FE2CFA74FE32" box="[1385,1435,392,418]" pageId="23" pageNumber="24">type</typeStatus>
locality and the site in
<collectingCountry id="F31F7640FF93FF914FF0FE14FDFCFE5A" box="[415,531,432,458]" name="Colombia" pageId="23" pageNumber="24">Colombia</collectingCountry>
makes us suspect that this female represents yet another undescribed species.
</paragraph>
<paragraph id="8BB736D0FF93FF914EA9FE73FB19FD02" blockId="23.[151,1436,151,778]" pageId="23" pageNumber="24">
<emphasis id="B97CEAC2FF93FF914EA9FE73FEF7FE61" bold="true" box="[198,280,471,497]" pageId="23" pageNumber="24">Types.</emphasis>
<materialsCitation id="3B603C8DFF93FF914F4DFE7CFBFDFDD2" accessionNumber="LN 202650" collectingDate="2004-02-17" collectionCode="INBC" collectorName="D. M. Wood" country="Costa Rica" county="Escazu" elevation="1360" location="Km" municipality="San Antonio" pageId="23" pageNumber="24" specimenCount="2" stateProvince="San Jose" typeStatus="holotype">
<typeStatus id="54B38872FF93FF914F4DFE7CFE7DFE62" box="[290,402,472,498]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
, male adult on microscope slide, labeled “
<typeStatus id="54B38872FF93FF914DF0FE7CFBD2FE62" box="[927,1085,472,498]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C084D53FF93FF914A26FE7CFF14FD8A" authority="Chaverri and Borkent" authorityName="Chaverri and Borkent" authorityYear="2007" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="woodi">
<emphasis id="B97CEAC2FF93FF914A26FE7CFB1CFE61" box="[1097,1267,472,497]" italics="true" pageId="23" pageNumber="24">Dixella woodi</emphasis>
Chaverri and Borkent
</taxonomicName>
”, “
<collectingCountry id="F31F7640FF93FF914F72FDA4FE4AFD8A" box="[285,421,512,538]" name="Costa Rica" pageId="23" pageNumber="24">Costa Rica</collectingCountry>
, Prov.
<collectingRegion id="49CCF832FF93FF914F97FDA4FD8CFD8A" box="[504,611,512,538]" country="Costa Rica" name="San Jose" pageId="23" pageNumber="24">San José</collectingRegion>
,
<collectingCounty id="62D64E5CFF93FF914C1EFDA4FD28FD8A" box="[625,711,512,538]" pageId="23" pageNumber="24">Escazú</collectingCounty>
,
<collectingMunicipality id="6BD3ACAAFF93FF914CBBFDA4FC82FD8A" box="[724,877,512,538]" pageId="23" pageNumber="24">San Antonio</collectingMunicipality>
, 4
<location id="8ED7600BFF93FF914DFDFDA4FC51FD8A" LSID="urn:lsid:plazi:treatment:03A187C6FF90FF914EF8FAD6FB2BFC9A:8ED7600BFF93FF914DFDFDA4FC51FD8A" box="[914,958,512,538]" country="Costa Rica" county="Escazu" municipality="San Antonio" name="Km" pageId="23" pageNumber="24" stateProvince="San Jose">Km</location>
E de
<location id="8ED7600BFF93FF914A67FDA4FB6FFD8A" LSID="urn:lsid:plazi:treatment:03A187C6FF90FF914EF8FAD6FB2BFC9A:8ED7600BFF93FF914A67FDA4FB6FFD8A" box="[1032,1152,512,538]" country="Costa Rica" county="Escazu" municipality="San Antonio" name="Palmichal" pageId="23" pageNumber="24" stateProvince="San Jose">Palmichal</location>
,
<location id="8ED7600BFF93FF914AE2FDA4FEE3FDD2" LSID="urn:lsid:plazi:treatment:03A187C6FF90FF914EF8FAD6FB2BFC9A:8ED7600BFF93FF914AE2FDA4FEE3FDD2" country="Costa Rica" county="Escazu" municipality="San Antonio" name="Albergue Nacientes de Palmichal" pageId="23" pageNumber="24" stateProvince="San Jose">Albergue Nacientes de Palmichal</location>
,
<quantity id="4CF09B35FF93FF914F74FD8CFE97FDD2" box="[283,376,552,578]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="23" pageNumber="24" unit="m" value="1360.0">
<elevation id="0025D1E3FF93FF914F74FD8CFE97FDD2" box="[283,376,552,578]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="23" pageNumber="24" unit="m" value="1360.0">1360 m</elevation>
</quantity>
,
<date id="FFB61010FF93FF914FE8FD8CFDE6FDD2" box="[391,521,552,578]" pageId="23" pageNumber="24" value="2004-02-17">
<collectingDate id="EFF2E9F8FF93FF914FE8FD8CFDE6FDD2" box="[391,521,552,578]" pageId="23" pageNumber="24" value="2004-02-17">17-II-2004</collectingDate>
</date>
, LN 202650 516850,
<collectorName id="26FD5306FF93FF914D7DFD8CFC4EFDD2" box="[786,929,552,578]" pageId="23" pageNumber="24">D.M. Wood</collectorName>
” (
<collectionCode id="ED19AE15FF93FF914DD2FD8CFBE6FDD2" box="[957,1033,552,578]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="23" pageNumber="24">INBC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B603C8DFF93FF914A46FD8CFB1EFD02" accessionNumber="LN 172094" collectingDate="2003-02-15" collectingDateMax="2003-02-17" collectingDateMin="2003-02-15" collectionCode="CNCI" collectorName="A. Borkent" country="Costa Rica" county="Tarrazu" elevation="340" location="Reserva Riosparaiso" municipality="San Carlos" pageId="23" pageNumber="24" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
<typeStatus id="54B38872FF93FF914A46FD8CFB7BFDD2" box="[1065,1172,552,578]" pageId="23" pageNumber="24" type="paratype">Paratype</typeStatus>
: male adult on microscope slide,
<collectingCountry id="F31F7640FF93FF914F45FDF4FE41FDFA" box="[298,430,592,618]" name="Costa Rica" pageId="23" pageNumber="24">Costa Rica</collectingCountry>
, Prov.
<collectingRegion id="49CCF832FF93FF914C6EFDF4FD84FDFA" box="[513,619,592,618]" country="Costa Rica" name="San Jose" pageId="23" pageNumber="24">San José</collectingRegion>
,
<collectingCounty id="62D64E5CFF93FF914C17FDF4FD3BFDFA" box="[632,724,592,618]" pageId="23" pageNumber="24">Tarrazú</collectingCounty>
,
<collectingMunicipality id="6BD3ACAAFF93FF914C8EFDF4FC8AFDFA" box="[737,869,592,618]" pageId="23" pageNumber="24">San Carlos</collectingMunicipality>
,
<location id="8ED7600BFF93FF914D1CFDF4FB88FDFA" LSID="urn:lsid:plazi:treatment:03A187C6FF90FF914EF8FAD6FB2BFC9A:8ED7600BFF93FF914D1CFDF4FB88FDFA" box="[883,1127,592,618]" country="Costa Rica" county="Tarrazu" municipality="San Carlos" name="Reserva Riosparaiso" pageId="23" pageNumber="24" stateProvince="San Jose">Reserva Riosparaíso</location>
, orilla del
<location id="8ED7600BFF93FF914A87FDF4FA9EFDFA" LSID="urn:lsid:plazi:treatment:03A187C6FF90FF914EF8FAD6FB2BFC9A:8ED7600BFF93FF914A87FDF4FA9EFDFA" box="[1256,1393,592,618]" country="Costa Rica" county="Tarrazu" municipality="San Carlos" name="Rio Blanco" pageId="23" pageNumber="24" stateProvince="San Jose">Río Blanco</location>
,
<locationDeviation id="9A052FF2FF93FF914B10FDF4FE9EFD02" location="Quepos" pageId="23" pageNumber="24">
<quantity id="4CF09B35FF93FF914B10FDF4FF53FD02" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="23" pageNumber="24" unit="km" value="16.0">16 km</quantity>
NE de Quepos
</locationDeviation>
,
<quantity id="4CF09B35FF93FF914F12FDDCFE26FD02" box="[381,457,632,658]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="23" pageNumber="24" unit="m" value="340.0">
<elevation id="0025D1E3FF93FF914F12FDDCFE26FD02" box="[381,457,632,658]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="23" pageNumber="24" unit="m" value="340.0">340 m</elevation>
</quantity>
,
<date id="FFB61010FF93FF914FB9FDDCFD91FD02" box="[470,638,632,658]" pageId="23" pageNumber="24" value="2003-02-15" valueMax="2003-02-17" valueMin="2003-02-15">
<collectingDate id="EFF2E9F8FF93FF914FB9FDDCFD91FD02" box="[470,638,632,658]" pageId="23" pageNumber="24" value="2003-02-15" valueMax="2003-02-17" valueMin="2003-02-15">15-17-II-2003</collectingDate>
</date>
, LN 172094 522815,
<collectorName id="26FD5306FF93FF914DE9FDDCFBE4FD02" box="[902,1035,632,658]" pageId="23" pageNumber="24">A. Borkent</collectorName>
, CD 5400&quot; (
<collectionCode id="ED19AE15FF93FF914AF2FDDCFB05FD02" box="[1181,1258,632,658]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="23" pageNumber="24">CNCI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB736D0FF93FF914EA9FD3BFB2BFC9A" blockId="23.[151,1436,151,778]" pageId="23" pageNumber="24">
<emphasis id="B97CEAC2FF93FF914EA9FD3BFDDAFD29" bold="true" box="[198,565,671,697]" pageId="23" pageNumber="24">Derivation of specific epithet.</emphasis>
This species is named for D. Monty Wood in recognition of his important and continuing research on Neotropical
<taxonomicName id="4C084D53FF93FF914C19FD6CFD21FD72" baseAuthorityName="McAlpine" baseAuthorityYear="1981" box="[630,718,712,738]" kingdom="Plantae" order="Diptera" pageId="23" pageNumber="24" rank="order">Diptera</taxonomicName>
and for his invaluable teaching, advice and contributions to developing an understanding of
<taxonomicName id="4C084D53FF93FF914C7FFD54FD87FC9A" baseAuthorityName="McAlpine" baseAuthorityYear="1981" box="[528,616,752,778]" kingdom="Plantae" order="Diptera" pageId="23" pageNumber="24" rank="order">Diptera</taxonomicName>
at the Instituto Nacional de Biodiversidad (INBio).
</paragraph>
</subSubSection>
</treatment>
</document>
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