167 lines
16 KiB
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167 lines
16 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.386.6913" ID-GBIF-Dataset="3a2df501-1763-4f11-a2ea-f9abaac99c00" ID-PMC="PMC3970064" ID-Pensoft-Pub="1313-2970-386-29" ID-PubMed="24693214" ID-ZBK="357FE9805295436EB40CFDD307D00D48" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-386-29" ModsDocOrigin="ZooKeys 386" ModsDocTitle="The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species" checkinTime="1451246199718" checkinUser="pensoft" docAuthor="Kerr, Peter H." docDate="2014" docId="FC8A0EE6B77FF9B5F8090DDA0715C992" docLanguage="en" docName="ZooKeys 386: 29-83" docOrigin="ZooKeys 386" docSource="http://dx.doi.org/10.3897/zookeys.386.6913" docTitle="Megophthalmidia perignea Kerr, 2014, sp. n." docType="treatment" docUuid="B80ABEC9-7043-4E7C-8273-EF6C2176B7C8" docUuidSource="ZooBank" docVersion="4" lastPageNumber="59" masterDocId="FFE4CC07FFFAC50AFFB0FFF4FFD6FFF4" masterDocTitle="The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species" masterLastPageNumber="83" masterPageNumber="29" pageNumber="55" updateTime="1668157894205" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kerr, Peter H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>386</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>83</mods:end>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.386.6913</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.386.6913</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-386-29</mods:identifier>
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<mods:identifier type="ZBK">357FE9805295436EB40CFDD307D00D48</mods:identifier>
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<mods:identifier type="ZooBank">357FE9805295436EB40CFDD307D00D48</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152051500" LSID="urn:lsid:zoobank.org:act:B80ABEC9-7043-4E7C-8273-EF6C2176B7C8" httpUri="http://treatment.plazi.org/id/FC8A0EE6B77FF9B5F8090DDA0715C992" lastPageId="30" lastPageNumber="59" pageId="26" pageNumber="55">
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<subSubSection pageId="26" pageNumber="55" type="nomenclature">
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<paragraph pageId="26" pageNumber="55">
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<taxonomicName LSID="http://zoobank.org/B80ABEC9-7043-4E7C-8273-EF6C2176B7C8" class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="26" pageNumber="55" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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<taxonomicNameLabel pageId="26" pageNumber="55">sp. n.</taxonomicNameLabel>
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Figs 62-71
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="27" lastPageNumber="56" pageId="26" pageNumber="55" type="type material">
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<paragraph pageId="26" pageNumber="55">Type material.</paragraph>
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<paragraph pageId="26" pageNumber="55">
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Holotype: ♂, "USA: ARIZONA: Mohave Co; 11km, ESE Kingman; Malaise nr Hualapai, Mt. Recreational Park; 5-15.vi.2012, ME Irwin; 1740m;
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="35.111332">35°06.68'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-113.902336">113°54.14'W</geoCoordinate>
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CSCA13L225" / "HOLOTYPE 13N210,
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="26" pageNumber="55" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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♂, Kerr, 2014" [red label]. Deposited in CSCA, mounted on gray point, entire specimen in excellent condition (Fig. 62), missing part of left wing.
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</paragraph>
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<paragraph lastPageId="27" lastPageNumber="56" pageId="26" pageNumber="55">
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Paratypes (all bearing a blue paratype label): 4 ♂♂, 3 ♀♀, same locality as holotype / [CSCA; specimen numbers 13M587 (dissected ♂, Figs 63-71), 13M569 (♂), 13M659 (♂, in alcohol), 13N390 (♂), 13N391 (♂),13N392 (♀), 13N393
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<pageBreakToken pageId="27" pageNumber="56" start="start">(</pageBreakToken>
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♀), 13N394 (♀, Fig. 62)]; ♂, "USA: ARIZONA, Gila Co; 21km S Globe; Malaise in Oak-Juniper, hillside thicket; 15-20.v.2013, ME Irwin; 1480m;
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="33.235332">33°14.12'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-110.782">110°46.92'W</geoCoordinate>
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CSCA13L218" [CSCA; specimen # 13M544]; ♂, ♀, "USA: ARIZONA, Gila Co; 21km S Globe; Malaise in Oak-Juniper, hillside thicket; 20-25.v.2013, ME Irwin; 1480m;
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="33.235332">33°14.12'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-110.782">110°46.92'W</geoCoordinate>
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CSCA13L222" [UAIC; specimen numbers 13M556 (♂), 13N389 (♀)].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="56" type="diagnosis">
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<paragraph pageId="27" pageNumber="56">Diagnosis.</paragraph>
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<paragraph pageId="27" pageNumber="56">
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Like its putative sister taxon,
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia ignea" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="ignea">Megophthalmidia ignea</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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sp. n. is superficially similar to
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia browni" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="browni">Megophthalmidia browni</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia mckibbeni" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="mckibbeni">Megophthalmidia mckibbeni</taxonomicName>
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sp. n. because of its brown thorax and contrasting cream-colored tibia, but may be separated from these taxa by its shortened and bare apical epandrial processes (Figs 63, 64).
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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is similar to
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia ignea" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="ignea">Megophthalmidia ignea</taxonomicName>
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in nearly all aspects except in relatively subtle features of the male genitalia. At its base, the short aedeagal tine of
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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has approximately the diameter as the long aedeagal tine. Also, in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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, the apical epandrial processes are shorter, wider, and turned inward more than in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia ignea" order="Diptera" pageId="27" pageNumber="56" phylum="Arthropoda" rank="species" species="ignea">Megophthalmidia ignea</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="30" lastPageNumber="59" pageId="27" pageNumber="56" type="description">
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<paragraph pageId="27" pageNumber="56">Description.</paragraph>
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<paragraph pageId="27" pageNumber="56">Male. Body length: 2.6-2.9, 2.8 [2.9] mm (n=6). Wing length: 2.5-2.9, 2.7 [2.5] mm (n=7).</paragraph>
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<paragraph pageId="28" pageNumber="57">
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<pageBreakToken pageId="28" pageNumber="57" start="start">Coloration</pageBreakToken>
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(Fig. 62). Head dark brown; antennal scape, pedicel, and flagellomeres brown; face dark brown, clypeus and labrum brown to dark brown; palps and labellum cream-colored to pale yellow (palpomeres 1-3 usually slightly darker than others, palpomere 2 with light patch where sensilla present). Thorax brown to dark brown throughout, except at the anterolateral margin of the dorsum and dorsal apronotal area, where it may be narrowly cream-colored or pale yellow; scutum setae golden brown to dark brown. Coxae clearly lighter in color than thorax, cream-colored to pale yellow; femora cream-colored throughout, dark brown at apical margin; tibiae and tarsi cream-colored to pale yellow, with densely-arranged dark brown setae; hind tibial comb yellowish, preceded by 0-3 (usually 3) dark brown setae. Wing hyaline without markings, wing veins brown; haltere stem and knob white to cream-colored. Abdominal segments concolorous brown. Terminalia light brown to brown.
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</paragraph>
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<caption pageId="28" pageNumber="57">
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<paragraph pageId="28" pageNumber="57">
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Figure 62.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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sp. n., habitus [holotype male above, # 13N210; female below, # 13N394]. Scale bar = 1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="29" pageNumber="58">
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<pageBreakToken pageId="29" pageNumber="58" start="start">Head</pageBreakToken>
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. Ocelli slightly raised, median ocellus in line with anterior margin of lateral ocelli, median ocellus approx. 0.3
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<normalizedToken originalValue="×–0.5×">x-0.5x</normalizedToken>
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size of lateral ocelli; lateral ocellus located approx. 1.8
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<normalizedToken originalValue="×">x</normalizedToken>
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diameter of ocellus from eye margin, separated from median ocellus by approx. 2.0
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<normalizedToken originalValue="×">x</normalizedToken>
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its own diameter. Eyes with microsetae, which are approximately as long as width of facet. Frons microtrichose, without setae, flattened. Antennal length 1.1-1.3, 1.2 [1.3] mm (n=6). Face clearly longer than wide, setose; clypeus and labrum microtrichose, without setae. Palpus with four palpomeres; palpomere 1 oblong, without setae; other palpomeres with brown setae; palpomere 2 bearing small pocket of sensilla; palpomere 1 length longer than or subequal in length to palpomere 2; palpomere 3 approx. same length as combined length of palpomeres 1 and 2; palpomere 4 subequal or slightly longer in length to combined length of palpomeres 2 and 3.
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</paragraph>
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<paragraph pageId="29" pageNumber="58">
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Thorax. Dorsum with evenly-distributed, short, appressed setae, bearing longer setae only along lateral and posterior margins. Antepronotum, proepisternum, and laterotergite bearing setae; remaining lateral thoracic sclerites bare. Costal wing vein extends beyond R5, approx. three-fifths distance between R5 and M1; R1 approximately the same length as r-m or slightly longer; cubital fork below or proximad of r-m base (as in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia occidentalis" order="Diptera" pageId="29" pageNumber="58" phylum="Arthropoda" rank="species" species="occidentalis">Megophthalmidia occidentalis</taxonomicName>
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, Fig. 52); R1, M1, M2, CuA1, and CuA2 with setae on upper surface (lacking setae on M1 + M2). Wing veins A1 and CuP absent.
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</paragraph>
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<paragraph pageId="30" pageNumber="59">
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<pageBreakToken pageId="30" pageNumber="59" start="start">Male</pageBreakToken>
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genitalia (Figs 63-71). Posterior margin of epandrium broadly emarginate at center (Fig. 65). Posterior processes of epandrium relatively short, approx. 2
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide, separated at base by approx. 1
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<normalizedToken originalValue="×">x</normalizedToken>
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width of process, turned inward (i.e., broad in lateral view (Fig. 63)); bare (Fig. 64). Gonocoxites as in Figs 66-68. Aedeagal fork bifurcated into tines; shorter tine slightly thickened (nearly same width as paired tine at base), both tines pointed outward apically (Fig. 70).
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</paragraph>
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<caption pageId="30" pageNumber="59">
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<paragraph pageId="30" pageNumber="59">
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Figures 63-65.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="30" pageNumber="59" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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sp. n., male epandrium [paratype, # 13M587] 63 Lateral view 64 Posterior view 65 Dorsal view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="30" pageNumber="59">
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<paragraph pageId="30" pageNumber="59">
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Figures 66-68.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="30" pageNumber="59" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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sp. n., male hypandrium [paratype, # 12J587] 66 Lateral view 67 Dorsal view 68 Ventral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="30" pageNumber="59">
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<paragraph pageId="30" pageNumber="59">
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Figures 69-71.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia perignea" order="Diptera" pageId="30" pageNumber="59" phylum="Arthropoda" rank="species" species="perignea">Megophthalmidia perignea</taxonomicName>
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sp. n., male aedeagus [paratype, # 13M587] 69 Lateral view 70 Dorsal view 71 Posterior view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="30" pageNumber="59">Female. Body length: 2.6-3.3, 2.8 mm (n=4). Antennal length: 0.8-1.0, 0.9 mm (n=4). Wing length: 2.5-3.1, 2.8 mm (n=4).</paragraph>
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<paragraph pageId="30" pageNumber="59">Coloration (Fig. 62). Same as male; cerci light brown to brown.</paragraph>
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<paragraph pageId="30" pageNumber="59">Head and thorax. Same as male, except antenna length shorter.</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="59" type="etymology">
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<paragraph pageId="30" pageNumber="59">Etymology.</paragraph>
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<paragraph pageId="30" pageNumber="59">
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The species epithet
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<normalizedToken originalValue="“perignea”">"perignea"</normalizedToken>
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is an adjective, derived for the Latin word for "near fiery" in reference to the distribution of this species, relative to that of its sister taxon,
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia ignea" order="Diptera" pageId="30" pageNumber="59" phylum="Arthropoda" rank="species" species="ignea">Megophthalmidia ignea</taxonomicName>
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sp. n.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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