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<document id="A8505C9E1A30AB0B25BDDA844E7ECF99" ID-CLB-Dataset="22879" ID-DOI="10.11646/zootaxa.4763.4.8" ID-GBIF-Dataset="83ffcc85-572b-4157-9cee-953c6bd08319" ID-ISSN="1175-5334" ID-Zenodo-Dep="3762198" ID-ZooBank="urn:lsid:zoobank.org:pub:2B670CBB-B791-477A-8519-311DC96741A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587572388770" checkinUser="carolina" docAuthor="Hendrich, Lars, Surbakti, Suriani &amp; Balke, Michael" docDate="2020" docId="804D87E1805EFFB7FF6AB4F4FCB2589B" docLanguage="en" docName="zootaxa.4763.4.8.pdf.imf" docOrigin="Zootaxa 4763 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limbodessus skalei Hendrich &amp; Surbakti &amp; Balke 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="588" masterDocId="7C74FF99805FFFB6FFFDB640FF9F5F66" masterDocTitle="Limbodessus skalei sp. nov., a new diving beetle species from the west Papuan Archipelago (Coleoptera: Dytiscidae, Bidessini)" masterLastPageNumber="592" masterPageNumber="587" pageNumber="588" updateTime="1698835551230" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="23F9D522D466794D463402267861C704" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5E4CCCB2F947449888539193921F2311">
<mods:title id="E4FA4593991A27F4BB25459CABA86A78">Limbodessus skalei sp. nov., a new diving beetle species from the west Papuan Archipelago (Coleoptera: Dytiscidae, Bidessini)</mods:title>
</mods:titleInfo>
<mods:name id="ADE4E2BBEC37015FA2DCEC3C66F76D93" type="personal">
<mods:role id="BFF1A03A1BB3C6A6327DE372B26A4D41">
<mods:roleTerm id="F7C7CC1702040689AB648D0D3BF0C101">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15A5CFBE672F61814754F6A871607BB4">Hendrich, Lars</mods:namePart>
<mods:affiliation id="0A938A47477F0FDC2F1BE94661713CE0">SNSB-Zoologische Staatssammlung München. Germany</mods:affiliation>
<mods:nameIdentifier id="2CA0F42CE6E5DCFE04F8D037F838525D" type="email">hendrich@snsb.de</mods:nameIdentifier>
</mods:name>
<mods:name id="92F38541CE6659F437436B44E11A0CC8" type="personal">
<mods:role id="5BD65DFF544A6BBAA50F0C50CF90EF25">
<mods:roleTerm id="080E8576FAF6C3D7DBC034DD0F403085">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="681DA571C593442459D0C42A032E7CA4">Surbakti, Suriani</mods:namePart>
<mods:affiliation id="D1488EAB9CF5C8F7D03B8AD7AB1CBED0">Universitas Cenderawasih, Department of Biology, Waena, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name id="71797713025F5704514C054D5D5F288B" type="personal">
<mods:role id="A259548C99293491B8CE324FA2A793B4">
<mods:roleTerm id="0D3DEE9F834807F75EC8F7688898D63B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4ABA4208284DAF7253544932CF309B93">Balke, Michael</mods:namePart>
<mods:affiliation id="6A02991DC16A28C1F735599E0C42DE7C">SNSB-Zoologische Staatssammlung München. Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="75614C70F06F891EC23E62B13FA32BCE">text</mods:typeOfResource>
<mods:relatedItem id="9A6E773BBAE8B8886DAF406A59B86D5A" type="host">
<mods:titleInfo id="15D50B597DCB5D638609F0C1E5881A17">
<mods:title id="63CF98F8D15727F59D1C3CF45E864F25">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7F97CE40E0147742F23C4393B9B28362">
<mods:date id="2D8F2B8A672323386BDA56F750CB3D95">2020</mods:date>
<mods:detail id="64D7A8F97135124C2B04A821ADA37377" type="pubDate">
<mods:number id="EE791289E0ED22EAD67C8227AADDFE10">2020-04-15</mods:number>
</mods:detail>
<mods:detail id="398AAFDC0A82DB55F85E6EADFEA4DE33" type="volume">
<mods:number id="34CF6564EFD644DF6EA378168B18E21A">4763</mods:number>
</mods:detail>
<mods:detail id="B7C7DCB62D635195FBD8E254BF919B60" type="issue">
<mods:number id="A6376A1149BC2DCE7AE91FA6ECE57AE0">4</mods:number>
</mods:detail>
<mods:extent id="82825580E46CE92615E17A96AE5C5639" unit="page">
<mods:start id="947FE6093479F4EA24DDEAFB6FF790F3">587</mods:start>
<mods:end id="23930DDC61EC0FED07B7CFF0AA3DA192">592</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8430DE8089025A556215D4EDA2B4B117">journal article</mods:classification>
<mods:identifier id="E218FC51A0F575DD9EDF1C725FA0A1A1" type="CLB-Dataset">22879</mods:identifier>
<mods:identifier id="C1F520EC138AC97F8554E70304DCDF49" type="DOI">10.11646/zootaxa.4763.4.8</mods:identifier>
<mods:identifier id="2E275C4E8DCC39ABF7345156FEBCFC69" type="GBIF-Dataset">83ffcc85-572b-4157-9cee-953c6bd08319</mods:identifier>
<mods:identifier id="63DB2CC47595E860A4C370F1375E40D6" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="9ED2B5F17ADDDAD9A08E0AA611E401D1" type="Zenodo-Dep">3762198</mods:identifier>
<mods:identifier id="535F578CEEF6B4A042405A9B0E61CEA7" type="ZooBank">urn:lsid:zoobank.org:pub:2B670CBB-B791-477A-8519-311DC96741A6</mods:identifier>
</mods:mods>
<treatment id="804D87E1805EFFB7FF6AB4F4FCB2589B" ID-DOI="http://doi.org/10.5281/zenodo.3804774" ID-GBIF-Taxon="163732329" ID-Zenodo-Dep="3804774" LSID="urn:lsid:plazi:treatment:804D87E1805EFFB7FF6AB4F4FCB2589B" httpUri="http://treatment.plazi.org/id/804D87E1805EFFB7FF6AB4F4FCB2589B" lastPageNumber="588" pageId="1" pageNumber="588">
<subSubSection id="40FE657C805EFFB7FF6AB4F4FE795DA9" box="[151,486,692,719]" pageId="1" pageNumber="588" type="nomenclature">
<paragraph id="085B36F7805EFFB7FF6AB4F4FE795DA9" blockId="1.[151,486,692,719]" box="[151,486,692,719]" pageId="1" pageNumber="588">
<heading id="5313819B805EFFB7FF6AB4F4FE795DA9" bold="true" box="[151,486,692,719]" fontSize="10" level="2" pageId="1" pageNumber="588" reason="2">
<emphasis id="3A90EAE5805EFFB7FF6AB4F4FE795DA9" bold="true" box="[151,486,692,719]" pageId="1" pageNumber="588">
<taxonomicName id="CFE44D74805EFFB7FF6AB4F4FE1E5DA9" ID-CoL="8LZYL" authority="Hendrich &amp; Surbakti &amp; Balke, 2020" authorityName="Hendrich &amp; Surbakti &amp; Balke" authorityYear="2020" box="[151,385,692,719]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="skalei" status="sp. nov.">
<emphasis id="3A90EAE5805EFFB7FF6AB4F4FE1E5DA9" bold="true" box="[151,385,692,719]" italics="true" pageId="1" pageNumber="588">Limbodessus skalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21A3579E805EFFB7FE75B4F5FE795DA9" box="[392,486,693,719]" pageId="1" pageNumber="588" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="40FE657C805EFFB7FF6AB4B7FC815C50" pageId="1" pageNumber="588" type="distribution">
<paragraph id="085B36F7805EFFB7FF6AB4B7FC815C50" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF6AB4B7FEA95C77" bold="true" box="[151,310,758,785]" pageId="1" pageNumber="588">
<typeStatus id="D75F8855805EFFB7FF6AB4B7FF4D5C77" box="[151,210,759,785]" pageId="1" pageNumber="588">Type</typeStatus>
locality.
</emphasis>
Riverbank of Werabiai River approx.
<quantity id="CF1C9B12805EFFB7FD14B4B7FCBC5C77" box="[745,803,759,785]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="588" unit="km" value="8.0">8 km</quantity>
from river mouth following the water course [
<geoCoordinate id="6DD05030805EFFB7FACFB4B6FA0C5C77" box="[1330,1427,758,785]" degrees="0.2585" direction="south" orientation="latitude" pageId="1" pageNumber="588" precision="5" value="-0.2585">0.2585S</geoCoordinate>
°
<geoCoordinate id="6DD05030805EFFB7FF6AB55AFE865C53" box="[151,281,794,821]" degrees="130.9856" direction="east" orientation="longitude" pageId="1" pageNumber="588" precision="5" value="130.9856">130.9856E</geoCoordinate>
°], Waigeo, Raja Ampat Province,
<collectingRegion id="CA20F815805EFFB7FD6EB55CFC865C50" box="[659,793,796,822]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="588">West Papua</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="40FE657C805EFFB7FF3AB57FFDEB5CA3" pageId="1" pageNumber="588" type="materials_examined">
<paragraph id="085B36F7805EFFB7FF3AB57FFDEB5CA3" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB57FFEE45C3F" bold="true" box="[199,379,830,857]" pageId="1" pageNumber="588">Type material:</emphasis>
<materialsCitation id="B88C3CAA805EFFB7FE7BB57FFE165CC7" ID-GBIF-Occurrence="2605167501" collectingDate="2004-11-01" collectionCode="MZB" country="Indonesia" determinerName="Balke" elevation="197" latitude="-0.258519" location="W-Papua, Raja Ampat Prov" longitude="130.985685" pageId="1" pageNumber="588" specimenCount="1" typeStatus="holotype">
<emphasis id="3A90EAE5805EFFB7FE7BB57FFE6C5C3F" bold="true" box="[390,499,831,857]" pageId="1" pageNumber="588">
<typeStatus id="D75F8855805EFFB7FE7BB57FFE6C5C3F" box="[390,499,831,857]" pageId="1" pageNumber="588" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="1EE2FD7E805EFFB7FDF8B57FFDDE5C3F" box="[517,577,831,857]" pageId="1" pageNumber="588" type="male">Male</specimenCount>
, “
<collectingCountry id="70F37667805EFFB7FDA7B57FFD535C3F" box="[602,716,831,857]" name="Indonesia" pageId="1" pageNumber="588">Indonesia</collectingCountry>
,
<location id="0D3B602C805EFFB7FD26B500FB805C3C" LSID="urn:lsid:plazi:treatment:804D87E1805EFFB7FF6AB4F4FCB2589B:0D3B602C805EFFB7FD26B500FB805C3C" box="[731,1055,831,858]" country="Indonesia" name="W-Papua, Raja Ampat Prov" pageId="1" pageNumber="588">W-Papua, Raja Ampat Prov</location>
., Waigeo, Werabiai River,
<quantity id="CF1C9B12805EFFB7FAA8B57FFA0A5C3F" box="[1365,1429,831,857]" metricMagnitude="2" metricUnit="m" metricValue="1.97" pageId="1" pageNumber="588" unit="m" value="197.0">
<elevation id="83C9D1C4805EFFB7FAA8B57FFA0A5C3F" box="[1365,1429,831,857]" metricMagnitude="2" metricUnit="m" metricValue="1.97" pageId="1" pageNumber="588" unit="m" value="197.0">197m</elevation>
</quantity>
,
<geoCoordinate id="6DD05030805EFFB7FF6AB522FE2E5C1B" box="[151,433,866,893]" pageId="1" pageNumber="588">-0.258519° 130.985685°</geoCoordinate>
,
<date id="7C5A1037805EFFB7FE43B523FDAD5C1B" box="[446,562,867,893]" pageId="1" pageNumber="588" value="2004-11-01">
<collectingDate id="6C1EE9DF805EFFB7FE43B523FDAD5C1B" box="[446,562,867,893]" pageId="1" pageNumber="588" value="2004-11-01">11.1.2004</collectingDate>
</date>
”, “
<typeStatus id="D75F8855805EFFB7FDAEB523FD755C1B" box="[595,746,867,893]" pageId="1" pageNumber="588" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="CFE44D74805EFFB7FD0EB523FC515C1B" authority="Hendrich &amp; Surbakti &amp; Balke, 2020" authorityName="Hendrich &amp; Surbakti &amp; Balke" authorityYear="2020" box="[755,974,867,893]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="skalei" status="sp. nov.">
<emphasis id="3A90EAE5805EFFB7FD0EB523FC515C1B" box="[755,974,867,893]" italics="true" pageId="1" pageNumber="588">Limbodessus skalei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21A3579E805EFFB7FC2AB524FB935C18" box="[983,1036,868,894]" pageId="1" pageNumber="588" rank="species">sp.n.</taxonomicNameLabel>
Hendrich &amp;
<determinerName id="FD87DA33805EFFB7FB58B523FB795C1B" box="[1189,1254,867,893]" pageId="1" pageNumber="588">Balke</determinerName>
det. 2019” [red printed label] (
<collectionCode id="6EF5AE32805EFFB7FEC2B5C7FE1E5CC7" box="[319,385,903,929]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="588" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
<emphasis id="3A90EAE5805EFFB7FE68B5C7FD925CC7" bold="true" box="[405,525,903,929]" pageId="1" pageNumber="588">
<typeStatus id="D75F8855805EFFB7FE68B5C7FD925CC7" box="[405,525,903,929]" pageId="1" pageNumber="588" type="paratype">Paratypes</typeStatus>
</emphasis>
: One male and one female with same data as
<typeStatus id="D75F8855805EFFB7FBE6B5C7FBE25CC7" box="[1051,1149,903,929]" pageId="1" pageNumber="588" type="holotype">holotype</typeStatus>
(
<collectionCode id="6EF5AE32805EFFB7FB70B5C7FB535CC7" box="[1165,1228,903,929]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="1" pageNumber="588" type="Museum">ZSM</collectionCode>
). Each
<typeStatus id="D75F8855805EFFB7FADCB5C8FA1E5CC4" box="[1313,1409,904,930]" pageId="1" pageNumber="588" type="paratype">paratype</typeStatus>
is provided with a red printed
<typeStatus id="D75F8855805EFFB7FE32B5ECFDB05CA0" box="[463,559,940,966]" pageId="1" pageNumber="588" type="paratype">paratype</typeStatus>
label.
</paragraph>
</subSubSection>
<subSubSection id="40FE657C805EFFB7FF3AB58FFB2C59DC" pageId="1" pageNumber="588" type="description">
<paragraph id="085B36F7805EFFB7FF3AB58FFE5B5B57" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB58FFEE15C8F" bold="true" box="[199,382,975,1001]" pageId="1" pageNumber="588">Measurements.</emphasis>
<typeStatus id="D75F8855805EFFB7FE7EB58FFE6F5C8F" box="[387,496,975,1001]" pageId="1" pageNumber="588" type="holotype">Holotype</typeStatus>
: TL =
<quantity id="CF1C9B12805EFFB7FDCBB58EFD045C8C" box="[566,667,974,1002]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="1" pageNumber="588" unit="mm" value="2.05">2.05 mm</quantity>
, TL-H =
<quantity id="CF1C9B12805EFFB7FCFCB58EFCF95C8C" box="[769,870,974,1002]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="588" unit="mm" value="1.85">1.85 mm</quantity>
, MW =
<quantity id="CF1C9B12805EFFB7FC42B58EFBBB5C8C" box="[959,1060,974,1002]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="588" unit="mm" value="0.95">0.95 mm</quantity>
, TL/MW = 2.16.
<typeStatus id="D75F8855805EFFB7FB19B590FACB5C8C" box="[1252,1364,976,1002]" pageId="1" pageNumber="588" type="paratype">Paratypes</typeStatus>
, male: TL =
<quantity id="CF1C9B12805EFFB7FF2AB5B2FEA15B68" box="[215,318,1010,1038]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="1" pageNumber="588" unit="mm" value="2.05">2.05 mm</quantity>
, TL-H =
<quantity id="CF1C9B12805EFFB7FE55B5B2FD915B68" box="[424,526,1010,1038]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="1" pageNumber="588" unit="mm" value="1.85">1.85 mm</quantity>
, MW =
<quantity id="CF1C9B12805EFFB7FD91B5B2FD4D5B68" box="[620,722,1010,1038]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="588" unit="mm" value="0.95">0.95 mm</quantity>
, TL/MW = 2.16; female: TL = 2.0 mm, TL-H =
<quantity id="CF1C9B12805EFFB7FB0AB5B2FAC25B68" box="[1271,1373,1010,1038]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="588" unit="mm" value="1.75">1.75 mm</quantity>
, MW = 1.0 mm, TL/MW = 2.00.
</paragraph>
<paragraph id="085B36F7805EFFB7FF3AB27AFD665B1F" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB27AFEC65B33" bold="true" box="[199,345,1082,1109]" pageId="1" pageNumber="588">Description.</emphasis>
A small yellow
<taxonomicName id="CFE44D74805EFFB7FDEDB27BFD3E5B33" authorityName="Guignot" authorityYear="1939" box="[528,673,1083,1109]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="3A90EAE5805EFFB7FDEDB27BFD3E5B33" box="[528,673,1083,1109]" italics="true" pageId="1" pageNumber="588">Limbodessus</emphasis>
</taxonomicName>
, with dark markings on elytra and slight habitus disruption between pronotum and elytron. Dorsoventrally rather flattened.
</paragraph>
<paragraph id="085B36F7805EFFB7FF3AB2C2FA9F5B83" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB2C2FED75BFB" bold="true" box="[199,328,1154,1181]" pageId="1" pageNumber="588">Coloration</emphasis>
: Antennae, palpi and head yellowish, pronotum completely testaceous, basally darkened between pronotal striae. Elytron testaceous with a wide basal and subbasal dark blackish-brown band (
<figureCitation id="90DF2A72805EFFB7FB46B2E7FB615BA7" box="[1211,1278,1191,1217]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1969]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure@2.[163,1425,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. The Limbodessus species known from New Guinea and adjacent islands. Dorsal habitus of L. skalei sp. nov., holotype, male (a); L. alexanderi (b), L. baliem (c) and L. compactus (d). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3762200" httpUri="https://zenodo.org/record/3762200/files/figure.png" pageId="1" pageNumber="588">Fig. 1</figureCitation>
). Ventral side including prosternal process and metacoxal plates somewhat darkened, legs and epipleura yellowish.
</paragraph>
<paragraph id="085B36F7805EFFB7FF3AB2AEFE5C594C" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB2AEFE7A5A6F" bold="true" box="[199,485,1262,1289]" pageId="1" pageNumber="588">Sculpture and structure</emphasis>
:
<emphasis id="3A90EAE5805EFFB7FE09B2AFFDAE5A6F" box="[500,561,1263,1289]" italics="true" pageId="1" pageNumber="588">Head</emphasis>
: Punctation fine, somewhat sparse, posteriorly slightly coarser and somewhat denser. Cervical line present (faint or partly dissolved into punctures in female).
<emphasis id="3A90EAE5805EFFB7FBCDB354FB005A4B" box="[1072,1183,1300,1325]" italics="true" pageId="1" pageNumber="588">Pronotum</emphasis>
: Broadest at posterior corners. Punctation coarse, dense, almost evenly distributed, microreticulation absent. Sides of pronotum margined and almost evenly rounded. Angle between pronotum and elytra rather distinct, pronotal and elytral plicae present. Striae almost straight, ¾ length of pronotum and strongly impressed.
<emphasis id="3A90EAE5805EFFB7FC4FB33FFC675AFF" box="[946,1016,1407,1433]" italics="true" pageId="1" pageNumber="588">Elytra</emphasis>
: Striae almost straight, 1/6 length of elytra. Punctation coarse, dense, almost evenly distributed, microreticulation not visible, shiny.
<emphasis id="3A90EAE5805EFFB7FB2DB3E3FAC95ADB" box="[1232,1366,1443,1469]" italics="true" pageId="1" pageNumber="588">Ventral side</emphasis>
: Base of elytral epipleuron without raised transverse carina delineating a basal pit. Metacoxae and metaventrite covered with numerous larger punctures, surface shiny, without microreticulation. Sexes dimorphic considering shape of antennomeres (see below).
</paragraph>
<paragraph id="085B36F7805EFFB7FF3AB073FCAB59F3" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB073FE9B592B" bold="true" box="[199,260,1587,1613]" pageId="1" pageNumber="588">Male</emphasis>
: Antenna filiform. Dorsal surface with coarse punctures but otherwise with shiny surface (
<figureCitation id="90DF2A72805EFFB7FB12B073FAB0592B" box="[1263,1327,1587,1613]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1969]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure@2.[163,1425,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. The Limbodessus species known from New Guinea and adjacent islands. Dorsal habitus of L. skalei sp. nov., holotype, male (a); L. alexanderi (b), L. baliem (c) and L. compactus (d). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3762200" httpUri="https://zenodo.org/record/3762200/files/figure.png" pageId="1" pageNumber="588">Fig. 1</figureCitation>
). Median lobe of aedeagus as in
<figureCitation id="90DF2A72805EFFB7FE61B017FD935917" box="[412,524,1623,1649]" captionStart="FIGURE 2" captionStartId="3.[151,250,1056,1081]" captionTargetBox="[151,1436,179,1028]" captionTargetId="figure@3.[151,1436,179,1028]" captionTargetPageId="3" captionText="FIGURE 2. Limbodessus skalei sp. nov., median lobe of aedeagus in ventral (a), lateral view (b), paramere, lateral view (c). Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762202" httpUri="https://zenodo.org/record/3762202/files/figure.png" pageId="1" pageNumber="588">Fig. 2a, b</figureCitation>
. Shape of penis fairly uniform, apex straight and pointed, with tiny setae on tip. Paramere with hook or bent finger like apical part (
<figureCitation id="90DF2A72805EFFB7FD28B03BFCB859F3" box="[725,807,1659,1685]" captionStart="FIGURE 2" captionStartId="3.[151,250,1056,1081]" captionTargetBox="[151,1436,179,1028]" captionTargetId="figure@3.[151,1436,179,1028]" captionTargetPageId="3" captionText="FIGURE 2. Limbodessus skalei sp. nov., median lobe of aedeagus in ventral (a), lateral view (b), paramere, lateral view (c). Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762202" httpUri="https://zenodo.org/record/3762202/files/figure.png" pageId="1" pageNumber="588">Fig. 2c</figureCitation>
).
</paragraph>
<paragraph id="085B36F7805EFFB7FF3AB0DFFB2C59DC" blockId="1.[151,1437,758,2046]" box="[199,1203,1695,1722]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB0DFFE8159DF" bold="true" box="[199,286,1695,1721]" pageId="1" pageNumber="588">Female</emphasis>
: Antenna more or less moniliform and body slightly broader (see measurements).
</paragraph>
</subSubSection>
<subSubSection id="40FE657C805EFFB7FF3AB083FD455864" pageId="1" pageNumber="588" type="etymology">
<paragraph id="085B36F7805EFFB7FF3AB083FD455864" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB083FED159BB" bold="true" box="[199,334,1731,1757]" pageId="1" pageNumber="588">Etymology.</emphasis>
The species is named after our friend and engaged coleopterist André Skale (Gera,
<collectingCountry id="70F37667805EFFB7FB0FB083FAC059BB" box="[1266,1375,1731,1757]" name="Germany" pageId="1" pageNumber="588">Germany</collectingCountry>
). The specific epithet is a substantive in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="40FE657C805EFFB7FF3AB14AFE735808" pageId="1" pageNumber="588" type="diagnosis">
<paragraph id="085B36F7805EFFB7FF3AB14AFE735808" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB14AFE4F5843" bold="true" box="[199,464,1802,1829]" pageId="1" pageNumber="588">Differential diagnosis.</emphasis>
The new species can be easily separated from all other epigean
<taxonomicName id="CFE44D74805EFFB7FB46B14BFAD35843" authorityName="Guignot" authorityYear="1939" box="[1211,1356,1803,1829]" class="Insecta" family="Dytiscidae" genus="Limbodessus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="3A90EAE5805EFFB7FB46B14BFAD35843" box="[1211,1356,1803,1829]" italics="true" pageId="1" pageNumber="588">Limbodessus</emphasis>
</taxonomicName>
by the combination of its colorful dorsal surface (
<figureCitation id="90DF2A72805EFFB7FD86B16FFD22582F" box="[635,701,1839,1865]" captionStart="FIGURE 1" captionStartId="2.[151,250,1944,1969]" captionTargetBox="[163,1425,181,1919]" captionTargetId="figure@2.[163,1425,181,1919]" captionTargetPageId="2" captionText="FIGURE 1. The Limbodessus species known from New Guinea and adjacent islands. Dorsal habitus of L. skalei sp. nov., holotype, male (a); L. alexanderi (b), L. baliem (c) and L. compactus (d). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3762200" httpUri="https://zenodo.org/record/3762200/files/figure.png" pageId="1" pageNumber="588">Fig. 1</figureCitation>
), smaller size and form of the median lobe (
<figureCitation id="90DF2A72805EFFB7FB4AB16FFB64582F" box="[1207,1275,1839,1865]" captionStart="FIGURE 2" captionStartId="3.[151,250,1056,1081]" captionTargetBox="[151,1436,179,1028]" captionTargetId="figure@3.[151,1436,179,1028]" captionTargetPageId="3" captionText="FIGURE 2. Limbodessus skalei sp. nov., median lobe of aedeagus in ventral (a), lateral view (b), paramere, lateral view (c). Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3762202" httpUri="https://zenodo.org/record/3762202/files/figure.png" pageId="1" pageNumber="588">Fig. 2</figureCitation>
) (see
<bibRefCitation id="6C754B06805EFFB7FAC2B170FE8E580B" author="Watts, C. H. S. &amp; Leijs, R." pageId="1" pageNumber="588" pagination="1 - 13" refId="ref2591" refString="Watts, C. H. S. &amp; Leijs, R. (2005) Review of the epigean species of Australian Limbodessus Guignot (Insecta: Coleoptera: Dytiscidae). Transactions of the Royal Society of South Australia, 129 (1), 1 - 13." type="journal article" year="2005">Watts &amp; Leijs 2005</bibRefCitation>
,
<bibRefCitation id="6C754B06805EFFB7FEE1B113FE7F5808" author="Balke, M. &amp; Ruthensteiner, B. &amp; Warikar, E. &amp; Neven, K. &amp; Hendrich, L." box="[284,480,1875,1902]" pageId="1" pageNumber="588" pagination="7096" refId="ref2376" refString="Balke, M., Ruthensteiner, B., Warikar, E., Neven, K. &amp; Hendrich, L. (2016) Two new species of Limbodessus diving beetles from New Guinea-short verbal descriptions flanked by online content (digital photography, μCT scans, drawings and DNA sequence data). Biodiversity Data Journal, 3, e 7096. https: // doi. org / 10.3897 / BDJ. 3. e 7096" type="journal article" year="2016">
Balke
<emphasis id="3A90EAE5805EFFB7FE99B114FE09580B" box="[356,406,1875,1901]" italics="true" pageId="1" pageNumber="588">et al</emphasis>
. 2016
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="40FE657C805EFFB7FF3AB136FF2C58D0" pageId="1" pageNumber="588" type="distribution">
<paragraph id="085B36F7805EFFB7FF3AB136FF2C58D0" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB136FEFE58F7" bold="true" box="[199,353,1910,1937]" pageId="1" pageNumber="588">Distribution.</emphasis>
Known only from the
<typeStatus id="D75F8855805EFFB7FD99B138FD0B58F4" box="[612,660,1912,1938]" pageId="1" pageNumber="588">type</typeStatus>
locality on Waigeo, around eight kilometers upstream Werabiai Riv- er.
</paragraph>
</subSubSection>
<subSubSection id="40FE657C805EFFB7FF3AB1FFFCB2589B" pageId="1" pageNumber="588" type="biology_ecology">
<paragraph id="085B36F7805EFFB7FF3AB1FFFCB2589B" blockId="1.[151,1437,758,2046]" pageId="1" pageNumber="588">
<emphasis id="3A90EAE5805EFFB7FF3AB1FFFEB358BF" bold="true" box="[199,300,1983,2009]" pageId="1" pageNumber="588">Ecology.</emphasis>
The three specimens were collected at the edge of the Werabiai River (
<figureCitation id="90DF2A72805EFFB7FB93B1FFFB2A58BF" box="[1134,1205,1983,2009]" captionStart="FIGURE 3" captionStartId="4.[151,260,1738,1765]" captionTargetBox="[282,1304,179,1711]" captionTargetId="figure@4.[282,1304,179,1711]" captionTargetPageId="4" captionText="FIGURE 3. Werabiai River and river banks approx. 8 km from river mouth, Waigeo island (Photo Dmitry Telnov, Natural History Museum, London, 20.II.2012)." figureDoi="http://doi.org/10.5281/zenodo.3762204" httpUri="https://zenodo.org/record/3762204/files/figure.png" pageId="1" pageNumber="588">Fig. 3</figureCitation>
), among stones and pebbles, of small, exposed puddles without any vegetation.
</paragraph>
</subSubSection>
</treatment>
</document>
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