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<document id="5160ED6277D7F4EFC0A77DEC867DA2FA" ID-DOI="10.11646/zootaxa.3846.1.1" ID-GBIF-Dataset="01a9d979-7b10-4c36-b1af-4acc2c0cb9a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286792" ID-ZooBank="97D4A04A-D75E-45CC-8A70-3EB3A4E94D9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460464243854" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2014" docId="113287AD9B5BFFD9FF13FC36FAB4FE7B" docLanguage="en" docName="zt03846p041.pdf" docOrigin="Zootaxa 3846 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvarium (Calvarium) australiense Zwick, 2014, n. sp." docType="treatment" docVersion="6" lastPageNumber="6" masterDocId="ED0BFFD59B59FFDCFF84FFF7FFB5FFF3" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer" masterLastPageNumber="41" masterPageNumber="1" pageNumber="3" updateTime="1698304595582" updateUser="plazi">
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<mods:title id="1646192253D0AD75C5E2219A116FAAC2">Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer</mods:title>
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<mods:name id="DB77842C85225ED8C9D29991F77DDF43" type="personal">
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<mods:namePart id="AA9307EE659932E65D5A26BED4BF8399">Zwick, Peter</mods:namePart>
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<mods:title id="8C8F018118E8CDEE2A10A8B7FE5B1D7D">Zootaxa</mods:title>
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<mods:part id="B6AA63AEF41B1B0A1396EC128CEA10F2">
<mods:date id="E6B38C8BDD8DC410C77E3E0C2FC4F839">2014</mods:date>
<mods:detail id="59D23F6478E3B97ADA6F636B1C61F874" type="volume">
<mods:number id="533F3C1B916A582FF635E346B6B94F75">3846</mods:number>
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<mods:classification id="668E58038FC487CEB64F1D9518C05A46">journal article</mods:classification>
<mods:identifier id="DB6072001C2BF81DFA2E2576F389754E" type="DOI">10.11646/zootaxa.3846.1.1</mods:identifier>
<mods:identifier id="E7F23781D2889CE26D0A3000836E79DA" type="GBIF-Dataset">01a9d979-7b10-4c36-b1af-4acc2c0cb9a7</mods:identifier>
<mods:identifier id="AA278536F831EFC832B9F7528EAEAB64" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F71238D162299C9B88AED6FFDDB94DC5" type="Zenodo-Dep">286792</mods:identifier>
<mods:identifier id="C3FF2FC265F4F0B9FA5B2CFB3B6D7152" type="ZooBank">97D4A04A-D75E-45CC-8A70-3EB3A4E94D9B</mods:identifier>
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<treatment id="113287AD9B5BFFD9FF13FC36FAB4FE7B" ID-DOI="http://doi.org/10.5281/zenodo.6126803" ID-GBIF-Taxon="127669611" ID-Zenodo-Dep="6126803" LSID="urn:lsid:plazi:treatment:113287AD9B5BFFD9FF13FC36FAB4FE7B" httpUri="http://treatment.plazi.org/id/113287AD9B5BFFD9FF13FC36FAB4FE7B" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="D18165309B5BFFDEFF13FC36FE32FC0F" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="992436BB9B5BFFDEFF13FC36FD2BFC28" blockId="2.[151,670,961,1021]" box="[151,670,961,987]" pageId="2" pageNumber="3">
<heading id="C26C81D79B5BFFDEFF13FC36FD2BFC28" bold="true" box="[151,670,961,987]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="ABEFEAA99B5BFFDEFF13FC36FD2BFC28" bold="true" box="[151,670,961,987]" pageId="2" pageNumber="3">
<emphasis id="ABEFEAA99B5BFFDEFF13FC36FDE1FC28" bold="true" box="[151,596,961,987]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="5E9B4D389B5BFFDEFF13FC36FDFBFC28" box="[151,590,961,987]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="australiense" status="sp. nov." subGenus="Calvarium">Calvarium (Calvarium) australiense</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="B0DC57D29B5BFFDEFDDFFC36FD2BFC28" box="[603,670,961,987]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="992436BB9B5BFFDEFF13FC13FE32FC0F" blockId="2.[151,670,961,1021]" box="[151,391,996,1021]" pageId="2" pageNumber="3">
(
<figureCitation id="01A02A3E9B5BFFDEFF1AFC13FE9DFC0E" box="[158,296,996,1021]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="2" pageNumber="3">Figures 1−5</figureCitation>
,
<figureCitation id="01A02A3E9B5BFFDEFEB1FC13FEC8FC0F" box="[309,381,996,1020]" captionStart="FIGURES 10 − 13" captionStartId="4.[151,264,1270,1293]" captionTargetBox="[236,1346,204,1245]" captionTargetId="figure@4.[232,1355,195,1247]" captionTargetPageId="4" captionText="FIGURES 10 − 13. Calvarium (Calvarium) australiense, n. sp., male: 10, T 8; 11, T 9; 12, penis; 13, tegmen and parameres. All to the same scale." httpUri="https://zenodo.org/record/286794/files/figure.png" pageId="2" pageNumber="3">10−13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D18165309B5BFFDEFF13FBDBFAEFFB94" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="992436BB9B5BFFDEFF13FBDBFAEFFB94" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">
<emphasis id="ABEFEAA99B5BFFDEFF13FBDBFE67FBB6" bold="true" box="[151,466,1068,1093]" pageId="2" pageNumber="3">
<typeStatus id="462088199B5BFFDEFF13FBDBFF67FBB6" box="[151,210,1068,1093]" pageId="2" pageNumber="3">Type</typeStatus>
material.
<typeStatus id="462088199B5BFFDEFED8FBDBFE78FBB6" box="[348,461,1068,1093]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
:
</emphasis>
♂ “Groote Eylandt N.B.Tindale“.
<emphasis id="ABEFEAA99B5BFFDEFC04FBDBFBB4FBB6" bold="true" box="[896,1025,1068,1093]" pageId="2" pageNumber="3">
<typeStatus id="462088199B5BFFDEFC04FBDBFC49FBB6" box="[896,1020,1068,1093]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="8F9DFD329B5BFFDEFB94FBDBFB86FBB7" box="[1040,1075,1068,1092]" pageId="2" pageNumber="3" type="generic">1♂</specimenCount>
with the same data;
<specimenCount id="8F9DFD329B5BFFDEFAC6FBDBFAD0FBB7" box="[1346,1381,1068,1092]" pageId="2" pageNumber="3" type="generic">1♂</specimenCount>
“N. Queensland Blackb's Coll.” (all
<collectionCode id="FF8AAE7E9B5BFFDEFE7BFBB8FDE1FB94" LSID="urn:lsid:biocol.org:col:34982" box="[511,596,1103,1127]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="2" pageNumber="3">SAMA</collectionCode>
; the specimens had been previously dissected and are in poor shape).
</paragraph>
</subSubSection>
<subSubSection id="D18165309B5BFFDEFF43FB86FE57F81E" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="992436BB9B5BFFDEFF43FB86FAE0FB5E" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">
<emphasis id="ABEFEAA99B5BFFDEFF43FB86FEF4FB79" bold="true" box="[199,321,1137,1162]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
A convex species distinguished by its terminal segments and genitalia, mainly the deeply excised U-shaped T8, the pincer-like T9, and by the large paramedian tegminal lobes ending in a long straight spine.
</paragraph>
<paragraph id="992436BB9B5BFFDEFF43FB41FB99FAE6" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">
<emphasis id="ABEFEAA99B5BFFDEFF43FB41FE93FB3C" bold="true" box="[199,294,1206,1231]" pageId="2" pageNumber="3">Habitus</emphasis>
(no intact specimen available). BL ~
<quantity id="5E639B5E9B5BFFDEFD56FB40FC98FB3D" box="[722,813,1207,1231]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="3" unit="mm" value="3.3">3.3 mm</quantity>
, BL/BW ~1.6. Strongly convex, shining, pilosity fine. Head and pronotum finely, elytra densely and much more coarsely punctate. Pronotum with strongly projecting front angles, front margin straight. Chestnut brown, appendages yellowish brown.
</paragraph>
<paragraph id="992436BB9B5BFFDEFF43FAE9FF59FA8F" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">Head strongly transverse, twice as wide across eyes as long from occipital foramen to front of frontoclypeus. Head capsule width correponds to about 60% of total body width. Front edge of frontoclypeus and oral area very narrow.</paragraph>
<paragraph id="992436BB9B5BFFDEFF43FA70FB53FA16" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">A deep antennal sulcus between the mouthparts and the eye is laterally delimited by a continuation of the curved anterolateral edge of the frontoclypeus which turns into a low ridge above the antennal insertion. The ridge curves towards the eye, then down along the front edge of the eye, leaving a small triangular gena.</paragraph>
<paragraph id="992436BB9B5BFFDEFF13FA19FD98F961" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">
There are two separate ridges beneath the eye. The upper subocular ridge (
<figureCitation id="01A02A3E9B5BFFDEFC67FA19FB93F9F4" box="[995,1062,1518,1543]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
, sor) begins below the triangular gena and runs backward a short distance below the eye. Caudally, it diverges from the eye and tapers away in the area covered by the pronotal lobes. In anterior view this ridge forms the side edge of the head capsule. Beneath it, on the underside of the head, begins the subgenal ridge (
<figureCitation id="01A02A3E9B5BFFDEFC8CF9A1FCFFF99C" box="[776,842,1622,1647]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
, sgr). It runs backward far ventrally from the eye and eventually meets the occipital ridge.
</paragraph>
<paragraph id="992436BB9B5BFFDEFF43F96BFBB8F925" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">The gular sutures are shorter than 1/3 of the distance between them. Sutures indistinct in front of the tentorial invaginations, reappearing a short distance before the rear edge of the mentum.</paragraph>
<paragraph id="992436BB9B5BFFDEFF43F915FADDF8EE" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">Antenna a little longer than head width. Scape an almost circular disc with sharp front edge. Resting position as described for the genus. Distal flagellar segments subcylindrical, about 2.5 times longer than apically wide.</paragraph>
<paragraph id="992436BB9B5BFFDEFF43F8D1FC77F839" blockId="2.[151,1437,1068,2029]" pageId="2" pageNumber="3">
Labrum narrow, sides parallel, rounded front margin with a very shallow median excision (
<figureCitation id="01A02A3E9B5BFFDEFB4FF8D1FAA5F8CC" box="[1227,1296,1830,1855]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
). Mandibles symmetrical, slender, with pointed tip and one large subterminal tooth (
<figureCitation id="01A02A3E9B5BFFDEFC3BF8BEFBB0F891" box="[959,1029,1865,1890]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
). Contour undulating between tooth and flat, bare, shining molar area. Terminal segment of maxillary palpus about twice as long as penultimate segment, tubular, indistinctly concave, apex blunt (
<figureCitation id="01A02A3E9B5BFFDEFD54F879FCA0F854" box="[720,789,1934,1959]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
). Last segment of labial palpus slender, concave, standing near the inner edge of the apically inflated penultimate segment (
<figureCitation id="01A02A3E9B5BFFDEFCEBF846FC01F839" box="[879,948,1969,1994]" captionStart="FIGURES 1 − 9" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[179,1406,213,1835]" captionTargetId="figure@3.[173,1417,195,1859]" captionTargetPageId="3" captionText="FIGURES 1 − 9. Calvarium spp., head morphology. 1 – 5, C. (C.) australiense, n. sp.; 6 – 9, C. (Calvariellum) hamifer, n. sp.: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2 – 5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture." httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="992436BB9B5BFFDEFF43F823FE57F81E" blockId="2.[151,1437,1068,2029]" box="[199,482,2004,2029]" pageId="2" pageNumber="3">Ventral side as for genus.</paragraph>
</subSubSection>
<caption id="CDE466339B5AFFDFFF13F8ACFBA9F81F" httpUri="https://zenodo.org/record/286793/files/figure.png" pageId="3" pageNumber="4" targetBox="[179,1406,213,1835]" targetPageId="3">
<paragraph id="992436BB9B5AFFDFFF13F8ACFBA9F81F" blockId="3.[151,1436,1883,2028]" pageId="3" pageNumber="4">
<emphasis id="ABEFEAA99B5AFFDFFF13F8ACFE89F883" bold="true" box="[151,316,1883,1906]" pageId="3" pageNumber="4">FIGURES 1−9.</emphasis>
<taxonomicName id="5E9B4D389B5AFFDFFEC7F8ACFE1AF882" box="[323,431,1883,1905]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="ABEFEAA99B5AFFDFFEC7F8ACFE1AF882" box="[323,431,1883,1905]" italics="true" pageId="3" pageNumber="4">Calvarium</emphasis>
</taxonomicName>
spp., head morphology. 1–5,
<taxonomicName id="5E9B4D389B5AFFDFFD59F8ABFC13F882" box="[733,934,1883,1905]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="australiense" status="sp. nov." subGenus="Calvarium">
<emphasis id="ABEFEAA99B5AFFDFFD59F8ABFC13F882" box="[733,934,1883,1905]" italics="true" pageId="3" pageNumber="4">C. (C.) australiense</emphasis>
</taxonomicName>
,
<emphasis id="ABEFEAA99B5AFFDFFC37F8ABFC58F882" bold="true" box="[947,1005,1884,1905]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="B0DC57D29B5AFFDFFC37F8ABFC58F882" box="[947,1005,1884,1905]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; 6–9,
<emphasis id="ABEFEAA99B5AFFDFFBA9F8ABFA8EF882" box="[1069,1339,1883,1905]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="5E9B4D389B5AFFDFFBA9F8ABFA8DF882" box="[1069,1336,1883,1905]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hamifer" status="sp. nov." subGenus="Calvariellum">C. (Calvariellum) hamifer</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="B0DC57D29B5AFFDFFAC6F8ABFAC9F882" box="[1346,1404,1884,1905]" pageId="3" pageNumber="4" rank="species">
<emphasis id="ABEFEAA99B5AFFDFFAC6F8ABFAC0F882" bold="true" box="[1346,1397,1884,1905]" pageId="3" pageNumber="4">n. sp</emphasis>
.
</taxonomicNameLabel>
: 1, 6, ventral views of mounted heads; 2, left mandible, dorsal view; 3, labrum; 4, maxillary palpus; 5, labium; 7, frontal view, slightly from ventral side, diagrammatic, right antenna removed; 8, maxillary palpus; 9, labial palpus. Figures 2–5 and 8, 9 to the same scale, respectively; Fig. 7 not to scale. Abbreviations: af, antennal foramen; as, accessory suture; asu, antennal sulcus; la, labrum; md, mandible; sgr, subgenal ridge; sor, subocular ridge; trs, transverse suture.
</paragraph>
</caption>
<caption id="CDE466339B5DFFD8FF13FB01FEFFFADF" httpUri="https://zenodo.org/record/286794/files/figure.png" pageId="4" pageNumber="5" targetBox="[236,1346,204,1245]" targetPageId="4">
<paragraph id="992436BB9B5DFFD8FF13FB01FEFFFADF" blockId="4.[151,1435,1270,1324]" pageId="4" pageNumber="5">
<emphasis id="ABEFEAA99B5DFFD8FF13FB01FEE0FAFF" bold="true" box="[151,341,1270,1293]" pageId="4" pageNumber="5">FIGURES 10−13.</emphasis>
<taxonomicName id="5E9B4D389B5DFFD8FEDFFB00FD78FAFE" box="[347,717,1271,1293]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="australiense" status="sp. nov." subGenus="Calvarium">
<emphasis id="ABEFEAA99B5DFFD8FEDFFB00FD78FAFE" box="[347,717,1271,1293]" italics="true" pageId="4" pageNumber="5">Calvarium (Calvarium) australiense</emphasis>
</taxonomicName>
,
<emphasis id="ABEFEAA99B5DFFD8FD5EFB00FCA6FAFF" bold="true" box="[730,787,1271,1292]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="B0DC57D29B5DFFD8FD5EFB00FCA6FAFF" box="[730,787,1271,1292]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male: 10, T8; 11, T9; 12, penis; 13, tegmen and parameres. All to the same scale.
</paragraph>
</caption>
<subSubSection id="D18165309B5DFFD9FF43FAA4FB1BFE96" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="992436BB9B5DFFD8FF43FAA4FB36FA47" blockId="4.[151,1437,1363,2036]" pageId="4" pageNumber="5">
<emphasis id="ABEFEAA99B5DFFD8FF43FAA4FEB1FA9F" bold="true" box="[199,260,1363,1388]" pageId="4" pageNumber="5">Male</emphasis>
. T8 and T9 (
<figureCitation id="01A02A3E9B5DFFD8FE12FAA4FDAEFA9F" box="[406,539,1363,1388]" captionStart="FIGURES 10 − 13" captionStartId="4.[151,264,1270,1293]" captionTargetBox="[236,1346,204,1245]" captionTargetId="figure@4.[232,1355,195,1247]" captionTargetPageId="4" captionText="FIGURES 10 − 13. Calvarium (Calvarium) australiense, n. sp., male: 10, T 8; 11, T 9; 12, penis; 13, tegmen and parameres. All to the same scale." httpUri="https://zenodo.org/record/286794/files/figure.png" pageId="4" pageNumber="5">Figs. 10, 11</figureCitation>
) with strong straight apodemes. T8 with deep U-shaped notch as wide as 1/3 of total width. Each lobe covered with sharp denticles or asperities. T9 with convergent apodemes ending in sharp tips separated by a narrow gap, resembling pliers or pincers. S8, S9 not found in preparations.
</paragraph>
<paragraph id="992436BB9B5DFFD8FF13FA48FB38F9B0" blockId="4.[151,1437,1363,2036]" pageId="4" pageNumber="5">
Penis (
<figureCitation id="01A02A3E9B5DFFD8FF60FA48FE83FA2B" box="[228,310,1471,1496]" captionStart="FIGURES 10 − 13" captionStartId="4.[151,264,1270,1293]" captionTargetBox="[236,1346,204,1245]" captionTargetId="figure@4.[232,1355,195,1247]" captionTargetPageId="4" captionText="FIGURES 10 − 13. Calvarium (Calvarium) australiense, n. sp., male: 10, T 8; 11, T 9; 12, penis; 13, tegmen and parameres. All to the same scale." httpUri="https://zenodo.org/record/286794/files/figure.png" pageId="4" pageNumber="5">Fig. 12</figureCitation>
) consisting of two narrowly separated flat long plates supporting a strongly sclerotized mushroomshaped cap. On its convex surface articulate the knee-shaped bases of the parameroids, details concealed by strong muscles extending between the cap and the front end of the plates. Parameroids flat, gently curved, with narrow shaft which widens caudally and ends in an obliquely truncate finely serrate axe-like apex.
</paragraph>
<paragraph id="992436BB9B5DFFD8FF13F9A7FA28F927" blockId="4.[151,1437,1363,2036]" pageId="4" pageNumber="5">
Tegmen with a soft, short base from which the parameres originate (
<figureCitation id="01A02A3E9B5DFFD8FC10F9B8FC53F99B" box="[916,998,1615,1640]" captionStart="FIGURES 10 − 13" captionStartId="4.[151,264,1270,1293]" captionTargetBox="[236,1346,204,1245]" captionTargetId="figure@4.[232,1355,195,1247]" captionTargetPageId="4" captionText="FIGURES 10 − 13. Calvarium (Calvarium) australiense, n. sp., male: 10, T 8; 11, T 9; 12, penis; 13, tegmen and parameres. All to the same scale." httpUri="https://zenodo.org/record/286794/files/figure.png" pageId="4" pageNumber="5">Fig. 13</figureCitation>
). Each paramere is deeply divided into a curved external spine and a flat medial lobe rising from a curved base, with an inward projecting hard condyluslike knob situated medially near base. Approximately parallel over most of length, then narrowed strongly, with a tiny denticle on outside. Terminal fifth forming a long spine in straight continuation of the sclerotized medial edge.
</paragraph>
<paragraph id="992436BB9B5DFFD8FF43F928FE17F90B" blockId="4.[151,1437,1363,2036]" box="[199,418,1759,1784]" pageId="4" pageNumber="5">
<emphasis id="ABEFEAA99B5DFFD8FF43F928FEABF90B" bold="true" box="[199,286,1759,1784]" pageId="4" pageNumber="5">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="992436BB9B5DFFD8FF43F8F4FC9DF87B" blockId="4.[151,1437,1363,2036]" pageId="4" pageNumber="5">
<emphasis id="ABEFEAA99B5DFFD8FF43F8F4FEBFF8EF" bold="true" box="[199,266,1795,1820]" pageId="4" pageNumber="5">Notes</emphasis>
. The antennal sulcus, the resting position of the antennae, the prosternal process, width and shortness of mesoventrite, and the male genitalia place the new species in the genus
<taxonomicName id="5E9B4D389B5DFFD8FC3AF8D0FB7DF8B3" authority="Ruta 2010" authorityName="Ruta" authorityYear="2010" box="[958,1224,1831,1856]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="ABEFEAA99B5DFFD8FC3AF8D0FB83F8CC" box="[958,1078,1831,1855]" italics="true" pageId="4" pageNumber="5">Calvarium</emphasis>
(Ruta 2010)
</taxonomicName>
. A subocular ridge in addition to the subgenal ridge was never described, but occurs also in
<taxonomicName id="5E9B4D389B5DFFD8FC50F8BBFBD1F890" box="[980,1124,1867,1891]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="superbum">
<emphasis id="ABEFEAA99B5DFFD8FC50F8BBFBD1F890" box="[980,1124,1867,1891]" italics="true" pageId="4" pageNumber="5">C. superbum</emphasis>
</taxonomicName>
(below) as well as in other species of
<taxonomicName id="5E9B4D389B5DFFD8FE89F898FE30F874" box="[269,389,1903,1927]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="ABEFEAA99B5DFFD8FE89F898FE30F874" box="[269,389,1903,1927]" italics="true" pageId="4" pageNumber="5">Calvarium</emphasis>
</taxonomicName>
(s. str.) (Ruta, pers. communication).
</paragraph>
<paragraph id="992436BB9B5DFFD9FF43F864FBB6FF27" blockId="4.[151,1437,1363,2036]" lastBlockId="5.[151,1437,151,392]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">The male genitalia are very similar to several other species. Caudal parameroids whose basal condyli rest on a hard, cap-like part of penis connected by muscles to the anterior penis edge occur in all species whose genitalia have been described. Axe-shaped parameroids occur in several species, a deeply notched or even completely divided and often spinulose T8 occurs in several species. T9 is usually much reduced. The tegmen is often divided into slender lateral spikes and a pair of variably shaped paramedian processes.</paragraph>
<paragraph id="992436BB9B5CFFD9FF43FF28FB1BFE96" blockId="5.[151,1437,151,392]" pageId="5" pageNumber="6">
The most similar species is
<taxonomicName id="5E9B4D389B5CFFD9FE7AFF17FC1AFF0B" authority="Yoshitomi &amp; Sato 2004" authorityName="Yoshitomi &amp; Sato" authorityYear="2004" box="[510,943,223,248]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="johorense">
<emphasis id="ABEFEAA99B5CFFD9FE7AFF17FD3EFF0B" box="[510,651,224,248]" italics="true" pageId="5" pageNumber="6">C. johorense</emphasis>
(Yoshitomi &amp; Sato 2004)
</taxonomicName>
from
<collectingCountry id="E18C762B9B5CFFD9FC77FF28FBEFFF0B" box="[1011,1114,223,248]" name="Malaysia" pageId="5" pageNumber="6">Malaysia</collectingCountry>
which shares the pincer-like T9. However, it is readily distinguished by the rounded overall contour of T8 and especially by the tegmen which has long lateral rods but short club-shaped paramedian lobes. The very large paramedian tegminal lobes ending in a long straight spine distinguish
<taxonomicName id="5E9B4D389B5CFFD9FD86FEBBFD19FE97" box="[514,684,332,356]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="australiense">
<emphasis id="ABEFEAA99B5CFFD9FD86FEBBFD19FE97" box="[514,684,332,356]" italics="true" pageId="5" pageNumber="6">C. australiense</emphasis>
</taxonomicName>
from the described genitalia of all congeners.
</paragraph>
</subSubSection>
<subSubSection id="D18165309B5CFFD9FF43FE98FAB4FE7B" box="[199,1281,367,392]" pageId="5" pageNumber="6" type="etymology">
<paragraph id="992436BB9B5CFFD9FF43FE98FAB4FE7B" blockId="5.[151,1437,151,392]" box="[199,1281,367,392]" pageId="5" pageNumber="6">
<emphasis id="ABEFEAA99B5CFFD9FF43FE98FEF3FE7B" bold="true" box="[199,326,367,392]" pageId="5" pageNumber="6">Etymology</emphasis>
. The name is a Latin adjective, nominative singular, neutrum, meaning “Australian”.
</paragraph>
</subSubSection>
</treatment>
</document>
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