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<document id="B8EB378B915FE79366F54D0F8B0EE7D4" ID-CLB-Dataset="21863" ID-DOI="10.5852/ejt.2016.241" ID-GBIF-Dataset="f694842b-3af3-4198-b01e-13d0b5466b5d" ID-ISSN="2118-9773" ID-Zenodo-Dep="3850299" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590443252253" checkinUser="valdenar" docAuthor="Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2016" docId="C4027A09140F5556D35BFB4EFB00B02F" docLanguage="en" docName="ejt-2016-241.pdf.imf" docOrigin="European Journal of Taxonomy 241" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Orphnus (Phornus) ferrierei Frolov &amp; Akhmetova 2016, sp. nov." docType="treatment" docUuid="BB79E7BA-8F60-4A97-BCFF-40017A1FBCD0" docUuidSource="ZooBank" docVersion="10" lastPageNumber="13" masterDocId="383B02711405555AD167FFB9FFF7B60D" masterDocTitle="Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="11" updateTime="1698841278433" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D9D5E7285BD9B70AF335EAD9375EC263" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="96B3C7E9B186EB036C78EF8B789BBBE4">
<mods:title id="1D0F8C1238EAC476895CC4E31D0617D5">Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae)</mods:title>
</mods:titleInfo>
<mods:name id="5E0C76176CF70EB5302C819808C63CB0" type="personal">
<mods:role id="F7BAB56938EC90B7588ACAEE3EBB183B">
<mods:roleTerm id="489BBBC670E01D6C8895B3B699FEE720">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46C24B7AEA644C8091729F6697266C14">Frolov, Andrey V.</mods:namePart>
<mods:nameIdentifier id="D2FCCD6149CAFD0B0270A226B16AFF70" type="LSID">FB31931B-44B9-4C22-963B-1023CEA88A21</mods:nameIdentifier>
<mods:affiliation id="07DCD3A7D1F0A414C51F50F5653C83F9">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia. Federal University of Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900 Cuiabá, MT, Brazil.</mods:affiliation>
<mods:nameIdentifier id="73B18C88A5AA10E878137742DEF73A3D" type="email">frolov@scarabaeoidea.com</mods:nameIdentifier>
</mods:name>
<mods:name id="43AA4E93F31EC19E6932B1AAB9B49FCC" type="personal">
<mods:role id="25BE563372BD1A6483C27FAAF7710419">
<mods:roleTerm id="0E1642787C97D24B2193F87A890B40EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C04DD76B842B0C971FBF15F51BB109A">Akhmetova, Lilia A.</mods:namePart>
<mods:nameIdentifier id="91E0429E149A4725148F0E2A8F24D06C" type="LSID">15207F4B-F31F-4F06-8C9F-0190B1EA74B0</mods:nameIdentifier>
<mods:affiliation id="324183F045FEA96C98B0E6B9F41019FA">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia. Federal University of Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900 Cuiabá, MT, Brazil.</mods:affiliation>
<mods:nameIdentifier id="5C6484F882D74ABECE75269D91758252" type="email">akhmetova@scarabaeoidea.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F0D3E0D6979C090C387B4E4CDE5EFA34">text</mods:typeOfResource>
<mods:relatedItem id="5E8956E2B48E248D799F8783CCCCBECF" type="host">
<mods:titleInfo id="382567A188DE57824CC0D0A8DEB486A2">
<mods:title id="57F3820A580C83206FF09D6F9BA69A8B">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="E8D55F6EB79D6EA7EFCAECFF9CC5D725">
<mods:date id="A15BF2728C7CE6056AC8C054A21D26E8">2016</mods:date>
<mods:detail id="1D8A9BEEB0A91D8937D5CDB3C97CE1CD" type="pubDate">
<mods:number id="8B262EA06309A082D66F48590C63A34D">2016-11-07</mods:number>
</mods:detail>
<mods:detail id="CB83A90A447138DCB21495C29FA79409" type="volume">
<mods:number id="C52972EB2E512E050DE8D425ACDDD878">241</mods:number>
</mods:detail>
<mods:extent id="DED65A23A085F1B6E0A1281B22B3D258" unit="page">
<mods:start id="84A380F4DD37D166844BE17906C326D1">1</mods:start>
<mods:end id="2763EDF90AEA8B8A6069A9D5C882077B">20</mods:end>
</mods:extent>
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<mods:classification id="5D6E7439DDFBB7E0207E1944926210BF">journal article</mods:classification>
<mods:identifier id="AB7F38345C45CA689C0940497B4A2C35" type="CLB-Dataset">21863</mods:identifier>
<mods:identifier id="035C2CD346B018D45075F006C7E84A08" type="DOI">10.5852/ejt.2016.241</mods:identifier>
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<mods:identifier id="68D4215740030B5398D67BAABDCA952D" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="2FDB0FBD304871F5698FA1F41250EF5A" type="Zenodo-Dep">3850299</mods:identifier>
<mods:identifier id="EF98A8D9EF550A67A61E9FB367BCF7CE" type="ZooBank">FEBC79B8-0F1B-4D15-937D-7D35C45D1408</mods:identifier>
</mods:mods>
<treatment id="C4027A09140F5556D35BFB4EFB00B02F" ID-DOI="http://doi.org/10.5281/zenodo.3854780" ID-GBIF-Taxon="164283133" ID-Zenodo-Dep="3854780" LSID="urn:lsid:zoobank.org:act:BB79E7BA-8F60-4A97-BCFF-40017A1FBCD0" httpUri="http://treatment.plazi.org/id/C4027A09140F5556D35BFB4EFB00B02F" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="04B19894140F5550D35BFB4EFC0FB31F" box="[572,1016,1271,1298]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="4C14CB1F140F5550D35BFB4EFC0FB31F" blockId="10.[358,1229,1271,1368]" box="[572,1016,1271,1298]" pageId="10" pageNumber="11">
<heading id="175C7C73140F5550D35BFB4EFC0FB31F" box="[572,1016,1271,1298]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="8BABB09C140F5550D35BFB4EFC62B31C" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[572,917,1271,1298]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ferrierei" status="sp. nov." subGenus="Phornus">
<emphasis id="7EDF170D140F5550D35BFB4EFC62B31C" bold="true" box="[572,917,1271,1298]" pageId="10" pageNumber="11">
<emphasis id="7EDF170D140F5550D35BFB4EFD5FB31F" bold="true" box="[572,680,1271,1298]" italics="true" pageId="10" pageNumber="11">Orphnus</emphasis>
(
<emphasis id="7EDF170D140F5550D3DEFB4EFCD6B31F" bold="true" box="[697,801,1271,1298]" italics="true" pageId="10" pageNumber="11">Phornus</emphasis>
)
<emphasis id="7EDF170D140F5550D255FB4EFC62B31C" bold="true" box="[818,917,1271,1297]" italics="true" pageId="10" pageNumber="11">ferrierei</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA76140F5550D2FAFB41FC0FB31F" box="[925,1016,1272,1298]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="04B19894140F5550D001FAA3FB3AB339" box="[358,1229,1306,1332]" pageId="10" pageNumber="11" type="description">
<paragraph id="4C14CB1F140F5550D001FAA3FB3AB339" blockId="10.[358,1229,1271,1368]" box="[358,1229,1306,1332]" pageId="10" pageNumber="11">
<uri id="383AC71D140F5550D001FAA3FB3AB339" box="[358,1229,1306,1332]" pageId="10" pageNumber="11">
urn:lsid:zoobank.org:act:
<uuid id="380DF1CA140F5550D3E3FAA3FB3AB339" box="[644,1229,1306,1332]" pageId="10" pageNumber="11">BB79E7BA-8F60-4A97-BCFF-40017A1FBCD0</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="04B19894140F5550D3A9FA84FC91B35A" box="[718,870,1341,1368]" pageId="10" pageNumber="11" type="description">
<paragraph id="4C14CB1F140F5550D3A9FA84FC91B35A" blockId="10.[358,1229,1271,1368]" box="[718,870,1341,1368]" pageId="10" pageNumber="11">
<figureCitation id="D490D79A140F5550D3A9FA84FCDAB355" box="[718,813,1341,1368]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="10" pageNumber="11">Figs 1O</figureCitation>
,
<figureCitation id="D490D79A140F5550D25DFA87FCBCB355" box="[826,843,1342,1368]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="10" pageNumber="11">4</figureCitation>
,
<figureCitation id="D490D79A140F5550D230FA84FC91B35A" box="[855,870,1341,1367]" captionStart="Fig" captionStartId="16.[189,232,1966,1992]" captionTargetBox="[194,1393,1011,1923]" captionTargetId="figure@16.[189,1398,1009,1925]" captionTargetPageId="16" captionText="Fig. 6. Orphnus (Phornus) spp. Locality map. Orphnus parastrangulatus sp. nov. is not mapped, because there are no more or less precise locality data available for this species." figureDoi="http://doi.org/10.5281/zenodo.3850323" httpUri="https://zenodo.org/record/3850323/files/figure.png" pageId="10" pageNumber="11">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="04B19894140F5550D1DAFAC6FDCCB03F" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="4C14CB1F140F5550D1DAFAC6FEC1B394" blockId="10.[189,310,1407,1433]" box="[189,310,1407,1433]" pageId="10" pageNumber="11">
<heading id="175C7C73140F5550D1DAFAC6FEC1B394" bold="true" box="[189,310,1407,1433]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7EDF170D140F5550D1DAFAC6FEC1B394" bold="true" box="[189,310,1407,1433]" pageId="10" pageNumber="11">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140F5550D1DAFA17FDCCB03F" blockId="10.[189,1399,1453,1586]" pageId="10" pageNumber="11">
<taxonomicName id="8BABB09C140F5550D1DAFA17FE7BB3C5" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[189,396,1454,1480]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ferrierei" status="sp. nov.">
<emphasis id="7EDF170D140F5550D1DAFA17FE7BB3C5" box="[189,396,1454,1480]" italics="true" pageId="10" pageNumber="11">Orphnus ferrierei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA76140F5550D0F4FA17FE19B3C5" box="[403,494,1454,1480]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="8BABB09C140F5550D3A0FA17FCBDB3C5" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[711,842,1454,1480]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="valeriae" status="sp. nov.">
<emphasis id="7EDF170D140F5550D3A0FA17FCBDB3C5" box="[711,842,1454,1480]" italics="true" pageId="10" pageNumber="11">O. valeriae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA76140F5550D235FA17FC58B3C5" box="[850,943,1454,1480]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, but can be separated from it in having pronotum smaller and more sparsely punctate with rounded punctures, distinct medial longitudinal stria basally on pronotum, frontoclypeal process tubercle-shaped, and less sclerotized parameres with angulate apices (in lateral view).
</paragraph>
</subSubSection>
<subSubSection id="04B19894140F5550D1DAF9E0FBB8B0AC" box="[189,1103,1625,1698]" pageId="10" pageNumber="11" type="etymology">
<paragraph id="4C14CB1F140F5550D1DAF9E0FEB3B07E" blockId="10.[189,324,1625,1651]" box="[189,324,1625,1651]" pageId="10" pageNumber="11">
<heading id="175C7C73140F5550D1DAF9E0FEB3B07E" bold="true" box="[189,324,1625,1651]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7EDF170D140F5550D1DAF9E0FEB3B07E" bold="true" box="[189,324,1625,1651]" pageId="10" pageNumber="11">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140F5550D1DAF93EFBB8B0AC" blockId="10.[189,1103,1671,1698]" box="[189,1103,1671,1698]" pageId="10" pageNumber="11">
This species is named after Pierre-Alfred Ferrière, collector of the
<typeStatus id="931075BD140F5550D2ADF931FC0AB0AF" box="[970,1021,1672,1698]" pageId="10" pageNumber="11">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="04B19894140F5550D1DAF971FAD8B1D2" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="4C14CB1F140F5550D1DAF971FE5DB0EF" blockId="10.[189,426,1736,1762]" box="[189,426,1736,1762]" pageId="10" pageNumber="11">
<heading id="175C7C73140F5550D1DAF971FE5DB0EF" bold="true" box="[189,426,1736,1762]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7EDF170D140F5550D1DAF971FE5DB0EF" bold="true" box="[189,426,1736,1762]" pageId="10" pageNumber="11">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="FCC3C142140F5550D18DF94EFD52B176" ID-GBIF-Occurrence="3761958346" collectionCode="MNHN" country="Central African Republic" determinerName="R. Paulian" location="Central African Republic" municipality="South-Western" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="4C14CB1F140F5550D18DF94EFEABB11C" blockId="10.[189,1399,1783,1915]" box="[234,348,1783,1809]" pageId="10" pageNumber="11">
<emphasis id="7EDF170D140F5550D18DF94EFEABB11C" bold="true" box="[234,348,1783,1809]" pageId="10" pageNumber="11">
<typeStatus id="931075BD140F5550D18DF94EFEABB11C" box="[234,348,1783,1809]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4C14CB1F140F5550D1DAF8A3FD52B176" blockId="10.[189,1399,1783,1915]" pageId="10" pageNumber="11">
<collectingCountry id="34BC8B8F140F5550D1DAF8A3FD82B139" box="[189,629,1818,1844]" name="Central African Republic" pageId="10" pageNumber="11">CENTRAL AFRICAN REPUBLIC</collectingCountry>
:
<specimenCount id="5AAD0096140F5550D3EFF8A2FD68B138" box="[648,671,1819,1845]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
, “MUSEUM PARIS CONGO FRANC. HAUTE-SANGA [
<collectingMunicipality id="AC705165140F5550D1A4F884FE83B15A" box="[195,372,1853,1879]" pageId="10" pageNumber="11">South-Western</collectingMunicipality>
<location id="49749DC4140F5550D0E7F884FD44B155" LSID="urn:lsid:plazi:treatment:C4027A09140F5556D35BFB4EFB00B02F:49749DC4140F5550D0E7F884FD44B155" box="[384,691,1853,1880]" country="Central African Republic" municipality="South-Western" name="Central African Republic" pageId="10" pageNumber="11">Central African Republic</location>
] P. A. FERRIÈRE 106-97 / Brachyorphnus ferrierei
<taxonomicNameLabel id="65ECAA76140F5550D458F887FA81B155" box="[1343,1398,1854,1880]" pageId="10" pageNumber="11" rank="species">n.sp.</taxonomicNameLabel>
<determinerName id="B9C827DB140F5550D1DAF8D8FEC0B176" box="[189,311,1889,1915]" pageId="10" pageNumber="11">R. Paulian</determinerName>
det. /
<typeStatus id="931075BD140F5550D0E6F8D8FDE8B176" box="[385,543,1889,1915]" pageId="10" pageNumber="11" type="holotype">HOLOTYPE</typeStatus>
” (
<collectionCode id="2ABA53DA140F5550D35FF8D8FD6EB176" box="[568,665,1889,1915]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="FCC3C142140F5550D18DF81BFAD8B1D2" ID-GBIF-Occurrence="3761958364" collectionCode="MNHN" country="Central African Republic" location="CENTRAL AFRICAN REPUBLIC" pageId="10" pageNumber="11" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<paragraph id="4C14CB1F140F5550D18DF81BFE90B1B1" blockId="10.[189,1327,1954,2016]" box="[234,359,1954,1980]" pageId="10" pageNumber="11">
<emphasis id="7EDF170D140F5550D18DF81BFE90B1B1" bold="true" box="[234,359,1954,1980]" pageId="10" pageNumber="11">
<typeStatus id="931075BD140F5550D18DF81BFE90B1B1" box="[234,359,1954,1980]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4C14CB1F140F5550D1DAF87CFAD8B1D2" blockId="10.[189,1327,1954,2016]" box="[189,1327,1989,2016]" pageId="10" pageNumber="11">
<collectingCountry id="34BC8B8F140F5550D1DAF87CFD93B1D2" box="[189,612,1989,2015]" name="Central African Republic" pageId="10" pageNumber="11">CENTRAL AFRICAN REPUBLIC</collectingCountry>
:
<specimenCount id="5AAD0096140F5550D309F87CFD47B1ED" box="[622,688,1989,2016]" pageId="10" pageNumber="11" type="male">2 ♂♂</specimenCount>
with the same locality label as the
<typeStatus id="931075BD140F5550D52BF87CFB44B1D2" box="[1100,1203,1989,2015]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(
<collectionCode id="2ABA53DA140F5550D5A4F87CFAD3B1D2" box="[1219,1316,1989,2015]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="04B19894140E5556D1DAFEB2FE1DB361" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="4C14CB1F140E5551D1DAFEB2FEB9B728" blockId="11.[189,334,267,293]" box="[189,334,267,293]" pageId="11" pageNumber="12">
<heading id="175C7C73140E5551D1DAFEB2FEB9B728" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="7EDF170D140E5551D1DAFEB2FEB9B728" bold="true" box="[189,334,267,293]" pageId="11" pageNumber="12">Description</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140E5551D18DFE83FE26B759" blockId="11.[189,734,313,375]" box="[234,465,313,340]" pageId="11" pageNumber="12">
<emphasis id="7EDF170D140E5551D18DFE83FEABB759" bold="true" box="[234,348,314,340]" pageId="11" pageNumber="12">
<typeStatus id="931075BD140E5551D18DFE83FEABB759" box="[234,348,314,340]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="D490D79A140E5551D00BFE80FE3FB759" box="[364,456,313,340]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="11" pageNumber="12">Fig. 4A</figureCitation>
)
</paragraph>
<paragraph id="4C14CB1F140E5551D1DAFEE4FD29B77A" blockId="11.[189,734,313,375]" box="[189,734,349,375]" pageId="11" pageNumber="12">
Body length
<quantity id="8B5366FA140E5551D032FEE4FE5DB77A" box="[341,426,349,375]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="12" unit="mm" value="14.0">14 mm</quantity>
. Colour uniformly brown.
</paragraph>
<paragraph id="4C14CB1F140E5551D1DAFE27FDEDB448" blockId="11.[189,1399,414,581]" pageId="11" pageNumber="12">
Anterior margin of frontoclypeus slightly convex in middle, slightly sinuate each side of medial convexity, rounded laterally, with a narrow border (
<figureCitation id="D490D79A140E5551D24EFE78FC86B7D6" box="[809,881,449,475]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
H–I). Frontoclypeus with conical, tubercleshaped transverse process medially approximately in middle of line connecting anterior margins of eyes and anterior margin of frontoclypeus. Frontoclypeus somewhat rugose anteriad of process and coarsely punctate posteriad of process.
</paragraph>
<caption id="18D49B97140E5551D1DAF831FC92B1E7" ID-DOI="http://doi.org/10.5281/zenodo.3850315" ID-Zenodo-Dep="3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1928,1954]" targetBox="[194,1388,717,1886]" targetPageId="11">
<paragraph id="4C14CB1F140E5551D1DAF831FC92B1E7" blockId="11.[189,1399,1928,2026]" pageId="11" pageNumber="12">
<emphasis id="7EDF170D140E5551D1DAF831FEF8B1AE" bold="true" box="[189,271,1928,1955]" pageId="11" pageNumber="12">Fig. 4.</emphasis>
<taxonomicName id="8BABB09C140E5551D07CF830FE19B1AE" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[283,494,1929,1955]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ferrierei" status="sp. nov.">
<emphasis id="7EDF170D140E5551D07CF830FE19B1AE" box="[283,494,1929,1955]" italics="true" pageId="11" pageNumber="12">Orphnus ferrierei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA76140E5551D09DF830FDAEB1AE" box="[506,601,1929,1955]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EDF170D140E5551D302F831FD8DB1AF" bold="true" box="[613,634,1928,1954]" pageId="11" pageNumber="12">A</emphasis>
,
<emphasis id="7EDF170D140E5551D3EAF831FD31B1AF" bold="true" box="[653,710,1928,1954]" pageId="11" pageNumber="12">C–D</emphasis>
. Habitus.
<emphasis id="7EDF170D140E5551D22FF830FCABB1AE" bold="true" box="[840,860,1929,1955]" pageId="11" pageNumber="12">B</emphasis>
. Labels of holotype.
<emphasis id="7EDF170D140E5551D500F830FB8CB1AE" bold="true" box="[1127,1147,1929,1955]" pageId="11" pageNumber="12">E</emphasis>
. Abdomen in dorsal view.
<emphasis id="7EDF170D140E5551D063F815FEE1B1CB" bold="true" box="[260,278,1964,1990]" pageId="11" pageNumber="12">F</emphasis>
. Aedeagus in lateral view.
<emphasis id="7EDF170D140E5551D33CF815FD85B1CB" bold="true" box="[603,626,1964,1990]" pageId="11" pageNumber="12">G</emphasis>
. Parameres in dorsal view.
<emphasis id="7EDF170D140E5551D2DAF815FC19B1CB" bold="true" box="[957,1006,1964,1990]" pageId="11" pageNumber="12">H–I</emphasis>
. Head in dorsal and apical view.
<emphasis id="7EDF170D140E5551D1DAF876FF3BB1E4" bold="true" box="[189,204,1999,2025]" pageId="11" pageNumber="12">J</emphasis>
. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes.
</paragraph>
</caption>
<paragraph id="4C14CB1F14095556D1DAFEB4FDECB747" blockId="12.[189,1398,268,330]" pageId="12" pageNumber="13">Eyes relatively large: width about 1/5.6 distance between eyes in dorsal view. Antennae 10-segmented, without malformed segments.</paragraph>
<paragraph id="4C14CB1F14095556D1DAFEC8FB58B415" blockId="12.[189,1398,369,536]" pageId="12" pageNumber="13">
Pronotum widely rounded laterally, almost as wide as elytra, 1.6 times wider than length, 0.55 length of elytra (
<figureCitation id="D490D79A14095556D054FE2DFE64B7A3" box="[307,403,404,430]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="12" pageNumber="13">Fig. 1O</figureCitation>
). Anterior border wide, with almost smooth posterior margin. Basal border narrow, keel-shaped, separated from pronotal disc by a smooth groove. Pronotal disc anteromedially slightly flattened. Surface of pronotum covered with double puncturation composed of large rounded and minute punctures; large punctures cover sides of pronotum and a smaller area anteromedially.
</paragraph>
<paragraph id="4C14CB1F14095556D1DAFD86FCCDB454" blockId="12.[189,826,575,601]" box="[189,826,575,601]" pageId="12" pageNumber="13">Scutellum rounded apically, about 1/16 length of elytra.</paragraph>
<paragraph id="4C14CB1F14095556D1DAFD39FEAAB509" blockId="12.[189,1399,640,772]" pageId="12" pageNumber="13">Elytra 1.08 times wider than long, with feeble marked humeral humps. Elytra widest approximately in middle, with widely rounded lateral margins. First six striae faintly visible as very shallow grooves, without distinct rows of punctures. Elytra covered with sparse punctures becoming slightly coarser towards base.</paragraph>
<paragraph id="4C14CB1F14095556D1DAFC95FC4BB54B" blockId="12.[189,956,810,838]" box="[189,956,810,838]" pageId="12" pageNumber="13">Brachypterous. Wings vestigial, narrow, about ¾ length of elytra.</paragraph>
<paragraph id="4C14CB1F14095556D1DAFCD4FB9FB58A" blockId="12.[189,1128,875,903]" box="[189,1128,875,903]" pageId="12" pageNumber="13">
Stridulatory field: carinae separated by 1/30 length of field in central ¼
<superScript id="BBDE665714095556D28CFCD2FC0EB574" attach="left" box="[1003,1017,875,889]" fontSize="6" pageId="12" pageNumber="13">rd</superScript>
(
<figureCitation id="D490D79A14095556D561FCD4FBACB58A" box="[1030,1115,877,903]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="12" pageNumber="13">Fig. 4J</figureCitation>
).
</paragraph>
<paragraph id="4C14CB1F14095556D1DAFC17FA96B202" blockId="12.[189,1399,942,1039]" pageId="12" pageNumber="13">
Abdominal sternite 8 medially slightly longer than sternites 6 and 7 combined; sternites 6 slightly longer than sternite 7. Pygidium visible from above, rounded apically. Plectrum trapezoidal, elongated (
<figureCitation id="D490D79A14095556D477FC6BFA9DB5E1" box="[1296,1386,978,1004]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="12" pageNumber="13">Fig. 4E</figureCitation>
). Lateral plate of second abdominal sternite relatively small, shorter than plectrum, with rounded apex.
</paragraph>
<paragraph id="4C14CB1F14095556D1DAFB8FFBD6B279" blockId="12.[189,1398,1078,1140]" pageId="12" pageNumber="13">
Aedeagus with relatively long parameres (0.53 length of phallobase). Apices of parameres narrowly rounded in dorsal view and lateral view, without excavations (
<figureCitation id="D490D79A14095556D2F1FBE0FC2CB279" box="[918,987,1113,1140]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="12" pageNumber="13">Fig. 4</figureCitation>
F–G).
</paragraph>
<paragraph id="4C14CB1F14095556D18DFB22FEB2B2B8" blockId="12.[189,325,1179,1240]" box="[234,325,1179,1205]" pageId="12" pageNumber="13">
<emphasis id="7EDF170D14095556D18DFB22FEB2B2B8" bold="true" box="[234,325,1179,1205]" pageId="12" pageNumber="13">Female</emphasis>
</paragraph>
<paragraph id="4C14CB1F14095556D1DAFB07FECEB2D5" blockId="12.[189,325,1179,1240]" box="[189,313,1214,1240]" pageId="12" pageNumber="13">Unknown.</paragraph>
<paragraph id="4C14CB1F14095556D1DAFB46FEC4B314" blockId="12.[189,307,1279,1305]" box="[189,307,1279,1305]" pageId="12" pageNumber="13">
<heading id="175C7C7314095556D1DAFB46FEC4B314" bold="true" box="[189,307,1279,1305]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="7EDF170D14095556D1DAFB46FEC4B314" bold="true" box="[189,307,1279,1305]" pageId="12" pageNumber="13">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F14095556D1DAFA96FE1DB361" blockId="12.[189,1399,1326,1388]" pageId="12" pageNumber="13">
<typeStatus id="931075BD14095556D1DAFA96FED8B344" box="[189,303,1327,1353]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
(
<figureCitation id="D490D79A14095556D026FA97FE7FB345" box="[321,392,1326,1352]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="12" pageNumber="13">Fig. 4</figureCitation>
C–D) are lighter coloured than
<typeStatus id="931075BD14095556D399FA97FC9FB345" box="[766,872,1326,1352]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, with smaller frontoclypeal processes. Body length 10.0 and
<quantity id="8B5366FA14095556D01DFAE8FE11B361" box="[378,486,1361,1388]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="12" pageNumber="13" unit="mm" value="12.5">12.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="04B1989414095556D1DAFA2AFB00B02F" pageId="12" pageNumber="13" type="distribution">
<paragraph id="4C14CB1F14095556D1DAFA2AFEA1B3A0" blockId="12.[189,342,1427,1453]" box="[189,342,1427,1453]" pageId="12" pageNumber="13">
<heading id="175C7C7314095556D1DAFA2AFEA1B3A0" bold="true" box="[189,342,1427,1453]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="7EDF170D14095556D1DAFA2AFEA1B3A0" bold="true" box="[189,342,1427,1453]" pageId="12" pageNumber="13">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F14095556D1DAFA7BFB00B02F" blockId="12.[189,1399,1473,1570]" pageId="12" pageNumber="13">
<taxonomicName id="8BABB09C14095556D1DAFA7BFEB3B3D1" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[189,324,1474,1500]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ferrierei" status="sp. nov.">
<emphasis id="7EDF170D14095556D1DAFA7BFEB3B3D1" box="[189,324,1474,1500]" italics="true" pageId="12" pageNumber="13">O. ferrierei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA7614095556D02AFA7BFE5DB3D1" box="[333,426,1474,1500]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
is known from Haute-Sangha, modern
<collectingRegion id="8E6F05FD14095556D2E0FA78FB30B3D6" box="[903,1223,1473,1499]" country="Central African Republic" name="Mambere-Kadei" pageId="12" pageNumber="13">Mambéré-Kadéï prefecture</collectingRegion>
of the
<collectingCountry id="34BC8B8F14095556D447FA78FE70B3F3" name="Central African Republic" pageId="12" pageNumber="13">Central African Republic</collectingCountry>
(
<figureCitation id="D490D79A14095556D0F0FA5DFE29B3F2" box="[407,478,1508,1535]" captionStart="Fig" captionStartId="16.[189,232,1966,1992]" captionTargetBox="[194,1393,1011,1923]" captionTargetId="figure@16.[189,1398,1009,1925]" captionTargetPageId="16" captionText="Fig. 6. Orphnus (Phornus) spp. Locality map. Orphnus parastrangulatus sp. nov. is not mapped, because there are no more or less precise locality data available for this species." figureDoi="http://doi.org/10.5281/zenodo.3850323" httpUri="https://zenodo.org/record/3850323/files/figure.png" pageId="12" pageNumber="13">Fig. 6</figureCitation>
). The region is a low-elevation plateau and a transition zone between Northern Congolian forest-savannah mosaic and Northwestern Congolian lowland forests ecoregions.
</paragraph>
</subSubSection>
</treatment>
</document>
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