<documentID-CLB-Dataset="166059"ID-DOI="http://dx.doi.org/10.3897/zookeys.1124.85952"ID-GBIF-Dataset="431ef183-6958-4de6-955f-a995b4c1a2dd"ID-PMC="PMC9836578"ID-Pensoft-Pub="1313-2970-1124-59"ID-Pensoft-UUID="2EFCF735419B588081D1586C3D1A8827"ID-PubMed="36762360"ID-ZooBank="09D04DEB58C24007AA8656ACABDE7BE3"ModsDocID="1313-2970-1124-59"checkinTime="1665870646046"checkinUser="pensoft"docAuthor="Zhang, Qiuqiu, Li, Shuqiang & Lin, Yucheng"docDate="2022"docId="5B595A91F0625C57BFC37C7FCEB829AA"docLanguage="en"docName="ZooKeys 1124: 59-108"docOrigin="ZooKeys 1124"docPubDate="2022-10-10"docSource="http://dx.doi.org/10.3897/zookeys.1124.85952"docTitle="Mysmena biangulata"docType="treatment"docVersion="5"id="2EFCF735419B588081D1586C3D1A8827"lastPageNumber="59"masterDocId="2EFCF735419B588081D1586C3D1A8827"masterDocTitle="Taxonomic study on Mysmenidae spiders (Mysmenidae, Araneae) from Xishuangbanna of Yunnan, China"masterLastPageNumber="108"masterPageNumber="59"pageNumber="59"updateTime="1732738812133"updateUser="ExternalLinkService">
<mods:titleid="FAE7D165EFA285001935C3F628EB785C">Taxonomic study on Mysmenidae spiders (Mysmenidae, Araneae) from Xishuangbanna of Yunnan, China</mods:title>
<mods:affiliationid="294F71A708F85F3BC0CA96C05F45F417">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
<mods:affiliationid="1B48EC5132CFB5BE4C0BAC26EBDEE376">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
<taxonomicNameid="00434116892F280DDF78532AFE6C533E"ID-CoL="73T4N"LSID="5B595A91-F062-5C57-BFC3-7C7FCEB829AA"authority="(Lin & Li, 2008)"baseAuthorityName="Lin & Li"baseAuthorityYear="2008"class="Arachnida"family="Mysmenidae"genus="Mysmena"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Mysmena biangulata"order="Araneae"pageId="0"pageNumber="59"phylum="Arthropoda"rank="species"species="biangulata">Mysmena biangulata (Lin & Li, 2008)</taxonomicName>
<figureCitationid="B892530CA62D33E9D4D6622742402402"captionStart="Figure 19"captionStartId="F19"captionText="Figure 19. Mysmena biangulata A-C male habitus D-F female habitus A, E dorsal B, F ventral C, D lateral. Abbreviations: AP = abdominal protuberance; FS = femoral spot; MC = Metatarsal clasping spine. Scale bars: 0.50 mm."figureDoi="10.3897/zookeys.1124.85952.figure19"httpUri="https://binary.pensoft.net/fig/754711"pageId="0"pageNumber="59">Figs 19</figureCitation>
<figureCitationid="60A46B90882989DF7490AAEAEAA291B7"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">, 20</figureCitation>
<taxonomicNameid="9748A2DD5883A687F2E873CA91A9CA43"authorityName="Lin & Li"authorityYear="2008"class="Arachnida"family="Mysmenidae"genus="Calodipoena"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Calodipoena biangulata"order="Araneae"pageId="0"pageNumber="59"phylum="Arthropoda"rank="species"species="biangulata">Calodipoena biangulata</taxonomicName>
<bibRefCitationid="C109A4309327C622EA0457E7506446A2"DOI="https://doi.org/10.3161/000345408X364337"author="Lin, Y"journalOrPublisher="Annales Zoologici, Warszawa"pageId="0"pageNumber="59"pagination="487 - 520"refId="B14"refString="Lin, Y, Li, S, 2008. Mysmenid spiders of China (Araneae: Mysmenidae). Annales Zoologici, Warszawa 58 (3): 487 - 520, DOI: https://doi.org/10.3161/000345408X364337"title="Mysmenid spiders of China (Araneae: Mysmenidae)."url="https://doi.org/10.3161/000345408X364337"volume="58"year="2008">Lin and Li 2008</bibRefCitation>
<taxonomicNameid="E087B4E5AD9F1BC34EB5428522F2D014"baseAuthorityName="Lin & Li"baseAuthorityYear="2008"class="Arachnida"family="Mysmenidae"genus="Mysmena"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Mysmena biangulata"order="Araneae"pageId="0"pageNumber="59"phylum="Arthropoda"rank="species"species="biangulata">Mysmena biangulata</taxonomicName>
<bibRefCitationid="16773F2D5F42EA7BA38614E905B6E88B"DOI="https://doi.org/10.1111/zoj.12199"author="Lopardo, L"journalOrPublisher="Zoological Journal of the Linnean Society"pageId="0"pageNumber="59"pagination="527 - 786"refId="B19"refString="Lopardo, L, Hormiga, G, 2015. Out of the twilight zone: Phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea). Zoological Journal of the Linnean Society 173 (3): 527 - 786, DOI: https://doi.org/10.1111/zoj.12199"title="Out of the twilight zone: Phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea)."url="https://doi.org/10.1111/zoj.12199"volume="173"year="2015">Lopardo and Hormiga 2015</bibRefCitation>
<figureCitationid="93D6BAC0583A11A4A0199E99544C1C32"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20B</figureCitation>
<bibRefCitationid="48E44C8B7E4C82B686B43405A3B664DE"author="Baert, L"journalOrPublisher="Bulletin - British Arachnological Society"pageId="0"pageNumber="59"pagination="303 - 308"refId="B1"refString="Baert, L, 1982. Spiders (Araneae) from Papua New Guinea III. Mysmenidae (Symphytognathoidea). Bulletin - British Arachnological Society 5: 303 - 308"title="Spiders (Araneae) from Papua New Guinea III. Mysmenidae (Symphytognathoidea)."volume="5"year="1982">Baert 1982</bibRefCitation>
<bibRefCitationid="C0529DFA033291C38F308CD030C7B736"author="Baert, L"journalOrPublisher="Indo-Malayan Zoology"pageId="0"pageNumber="59"pagination="225 - 244"refId="B2"refString="Baert, L, 1984. Spiders (Araneae) from Papua New Guinea. IV. Ochyroceratidae, Telemidae, Hadrotarsidae and Mysmenidae. Indo-Malayan Zoology 2: 225 - 244"title="Spiders (Araneae) from Papua New Guinea. IV. Ochyroceratidae, Telemidae, Hadrotarsidae and Mysmenidae."volume="2"year="1984">Baert 1984</bibRefCitation>
<figureCitationid="CD04CB167DA1C96AC88DE0C2E9AA1AA6"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20E</figureCitation>
<bibRefCitationid="77385CBE91679EF969769864E67D563F"DOI="https://doi.org/10.3161/000345408X364337"author="Lin, Y"journalOrPublisher="Annales Zoologici, Warszawa"pageId="0"pageNumber="59"pagination="487 - 520"refId="B14"refString="Lin, Y, Li, S, 2008. Mysmenid spiders of China (Araneae: Mysmenidae). Annales Zoologici, Warszawa 58 (3): 487 - 520, DOI: https://doi.org/10.3161/000345408X364337"title="Mysmenid spiders of China (Araneae: Mysmenidae)."url="https://doi.org/10.3161/000345408X364337"volume="58"year="2008">Lin and Li 2008</bibRefCitation>
<taxonomicNameid="F9D53F019EC6A094D47E67A4AB764FEE"baseAuthorityName="Lin & Li"baseAuthorityYear="2008"class="Arachnida"family="Mysmenidae"genus="Mysmena"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Mysmena biangulata"order="Araneae"pageId="0"pageNumber="59"phylum="Arthropoda"rank="species"species="biangulata">
<figureCitationid="82E8E165FD94B5DF35BD390998469191"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20A, B</figureCitation>
<bibRefCitationid="C66AAE67C5AF2045D1A15C0438E5B87C"author="Baert, L"journalOrPublisher="Bulletin - British Arachnological Society"pageId="0"pageNumber="59"pagination="303 - 308"refId="B1"refString="Baert, L, 1982. Spiders (Araneae) from Papua New Guinea III. Mysmenidae (Symphytognathoidea). Bulletin - British Arachnological Society 5: 303 - 308"title="Spiders (Araneae) from Papua New Guinea III. Mysmenidae (Symphytognathoidea)."volume="5"year="1982">Baert 1982</bibRefCitation>
). Females can be distinguished by the coiled spermathecae with modified glandulous sac and the directly opposite basal partition of copulatory ducts (Fig.
<figureCitationid="473DFA16670D1EB0034EBB32AAD9F8C3"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20D, E</figureCitation>
<taxonomicNameid="6E7B217C349D504F04C802D11256144B"baseAuthorityName="Lin & Li"baseAuthorityYear="2008"class="Arachnida"family="Mysmenidae"genus="Mysmena"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Mysmena biangulata"order="Araneae"pageId="0"pageNumber="59"phylum="Arthropoda"rank="species"species="biangulata">
<bibRefCitationid="AB89BAE36371251835980C0A7DBC0D5E"DOI="https://doi.org/10.3161/000345408X364337"author="Lin, Y"journalOrPublisher="Annales Zoologici, Warszawa"pageId="0"pageNumber="59"pagination="487 - 520"refId="B14"refString="Lin, Y, Li, S, 2008. Mysmenid spiders of China (Araneae: Mysmenidae). Annales Zoologici, Warszawa 58 (3): 487 - 520, DOI: https://doi.org/10.3161/000345408X364337"title="Mysmenid spiders of China (Araneae: Mysmenidae)."url="https://doi.org/10.3161/000345408X364337"volume="58"year="2008">Lin and Li 2008</bibRefCitation>
<figureCitationid="EB3C484F756F70A1C0516A9380533ED7"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20A, B</figureCitation>
): light-yellow; tibia big, about 2/3 volume of the bulb, cup-shaped; Except for retrolateral region, a row of long setae almost encircling the distal brim (Fig.
<figureCitationid="280A2F09EA92B9F8D0FACBC015C9B9FA"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20B</figureCitation>
). Cymbium nearly transparent; the cymbial conductor lateral bending, parallel to the cymbial process; the cymbial fold long and sclerotized, bears a row of ordered setae (Fig.
<figureCitationid="259A035A7576D5688604BFF0AE5A7BF9"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20A, B</figureCitation>
<figureCitationid="5919CDB99C9EB7C949A1AEC45E570DE4"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20A, B</figureCitation>
<taxonomicNameid="3938E53D3A893AD17719EB40D170528F"baseAuthorityName="Lin & Li"baseAuthorityYear="2008"class="Arachnida"family="Mysmenidae"genus="Mysmena"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Mysmena biangulata"order="Araneae"pageId="0"pageNumber="59"phylum="Arthropoda"rank="species"species="biangulata">
<figureCitationid="16068EDAACBE5296333639431B976CC6"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20C-E</figureCitation>
<figureCitationid="5344866BE69C48175CA0398BFC3FA724"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20C-E</figureCitation>
<figureCitationid="EAE3C35AC350917615C1F8EE9B9850F9"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20D</figureCitation>
). Fertilization ducts and copulatory ducts slightly sclerotized, coiling around each other; the fertilization ducts opening to both edges of epigyne; anterior copulatory ducts sclerotized, flow-shaped; two openings converge toward the centre of epigyne (Fig.
<figureCitationid="822ED5AE4EA21079EC5E3AC85517736C"captionStart="Figure 20"captionStartId="F20"captionText="Figure 20. Mysmena biangulata A, B male palp C epigyne D, E vulva. A prolateral B retrolateral C, D ventral E dorsal. Abbreviations: CD = copulatory duct; Cy = cymbium; CyC = cymbial conductor; CyF = cymbial fold; CyFs = setae on cymbial fold; CyP = cymbial process; E = embolus; FD = fertilization duct; S = spermatheca; SD = spermatic duct; Sp = scape; Te = tegulum; Ti = palpal tibia. Scale bars: 0.20 mm (A, B); 0.10 mm (C-E)."figureDoi="10.3897/zookeys.1124.85952.figure20"httpUri="https://binary.pensoft.net/fig/754712"pageId="0"pageNumber="59">20D, E</figureCitation>
