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383 lines
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<document id="61C35DF310AEE9F766BD54499081EA2C" ID-DOI="http://dx.doi.org/10.3897/zookeys.394.6446" ID-GBIF-Dataset="8534078f-1f99-4b9e-866b-80609e341172" ID-PMC="PMC3978267" ID-Pensoft-Pub="1313-2970-394-1" ID-PubMed="24715784" ID-ZooBank="32F5211696ED46C68BF9BE59C09F0B1A" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-394-1" ModsDocOrigin="ZooKeys 394" ModsDocTitle="A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)" checkinTime="1451246058701" checkinUser="pensoft" docAuthor="Yoshimura, Masashi & Fisher, Brian L." docDate="2014" docId="419ECA5DB47A5ACC938F26ADCCC42080" docLanguage="en" docName="ZooKeys 394: 1-99" docOrigin="ZooKeys 394" docSource="http://dx.doi.org/10.3897/zookeys.394.6446" docTitle="Mystrium rogeri Forel 1899" docType="treatment" docVersion="4" lastPageNumber="51" masterDocId="FF955F09FFA4FFC3FFC5FFA24014FFB0" masterDocTitle="A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)" masterLastPageNumber="99" masterPageNumber="1" pageNumber="45" updateTime="1701316919487" updateUser="plazi">
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<mods:title id="5E6AD5BA5ACC2B3E60B7813639A05035">A revision of the ant genus Mystrium in the Malagasy region with description of six new species and remarks on Amblyopone and Stigmatomma (Hymenoptera, Formicidae, Amblyoponinae)</mods:title>
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<mods:namePart id="CE15532E8A035CBFA94D1BFA2AA1E224">Yoshimura, Masashi</mods:namePart>
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<mods:namePart id="36222A34F5755E830F4E2FBF1CB4EAD7">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="074F6E108152AAC801C6F39068D98C26">text</mods:typeOfResource>
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<mods:title id="89A16D8B4D3F1EBB3F94ADE87F35F6CC">ZooKeys</mods:title>
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<mods:date id="8B3F40DC17624D84A0EF1111E79DF4BD">2014</mods:date>
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<mods:detail id="CC0D26B5A85BE1D1FF1F4C30C511C4A3" type="volume">
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<mods:number id="28479D69C5F08C82222D8238A19E906A">394</mods:number>
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<mods:start id="5E26C192B0C5586CA7B55EE890DB2DBB">1</mods:start>
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<mods:end id="01A22F457BBC04B78FD1D7AB091FA851">99</mods:end>
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<mods:url id="B45102AB220E26144135C1F21AFDEB12">http://dx.doi.org/10.3897/zookeys.394.6446</mods:url>
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<mods:classification id="5E636D2110DD40557DE64725FC1235AE">journal article</mods:classification>
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<mods:identifier id="842817D9E87B6EFE6568205329F4E0CE" type="DOI">http://dx.doi.org/10.3897/zookeys.394.6446</mods:identifier>
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<mods:identifier id="E5C7B81B80FB5B4AA60EE9409FD83311" type="Pensoft-Pub">1313-2970-394-1</mods:identifier>
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</mods:mods>
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<treatment id="419ECA5DB47A5ACC938F26ADCCC42080" ID-GBIF-Taxon="152052268" LSID="urn:lsid:plazi:treatment:419ECA5DB47A5ACC938F26ADCCC42080" httpUri="http://treatment.plazi.org/id/419ECA5DB47A5ACC938F26ADCCC42080" lastPageId="50" lastPageNumber="51" pageId="44" pageNumber="45">
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<subSubSection id="69EC19411B33EE9B722876F4F79F4968" pageId="44" pageNumber="45" type="nomenclature">
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<paragraph id="89E480D93402033DB8E9E2625982A6A9" pageId="44" pageNumber="45">
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<taxonomicName id="E369C7B87CCFA069EEA072EDD0AF575A" ID-CoL="45FPV" LSID="http://species-id.net/wiki/Mystrium_rogeri" authority="Forel, 1899" authorityName="Forel" authorityYear="1899" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri Forel, 1899</taxonomicName>
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Figs 4A, 8A, 9B, 9D, 12B, 29B, 37C, 38C, 39C, 40C, 41C, 43B, 43D, 44B, 44D, 45B, 45D, 46B, 46D, 47B, 47D, 48B, 48D, 49B
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</paragraph>
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</subSubSection>
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<subSubSection id="9685743C76E559F1D42BEEAD6C28A7FF" pageId="44" pageNumber="45" type="reference_group">
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<paragraph id="80E63F33431021618A410CBDB839DD05" pageId="44" pageNumber="45">
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<taxonomicName id="43BB39721A153B7FC2A52754211B2AE1" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
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Forel, 1899 (in part: worker only, not male). MADAGASCAR. Syntypes: worker and male [The lectotype is designated below].
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</paragraph>
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<paragraph id="7945A03844AC2830C706D4F4953195EA" pageId="44" pageNumber="45">
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<taxonomicName id="62C2C8D70708B46D5B7147CD2530A61E" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
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:
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<bibRefCitation id="EABE36C055E26E977C5CA719367EC960" author="Menozzi, C" journalOrPublisher="Zoologischer Anzeiger" pageId="76" pageNumber="77" pagination="518 - 536" title="Revisione delle formiche del genere Mystrium Roger." volume="82" year="1929">Menozzi 1929</bibRefCitation>
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.
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</paragraph>
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<paragraph id="1C15976E176F09D2F3E5B4C94C04F341" pageId="44" pageNumber="45">
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<taxonomicName id="A2AEC116D54E9D81FB437C8BEA7C0763" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
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[Not Roger, 1862]:
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<bibRefCitation id="2AE6113C0469EDCB5D6D091B5D25DB67" author="Forel, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="76" pageNumber="77" pagination="516 - 535" title="Nouvelles especes de Formicides de Madagascar (recoltees par M. Sikora). Premiere serie." volume="36" year="1892">Forel 1892</bibRefCitation>
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, in part (worker only).
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</paragraph>
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<paragraph id="B4B1C81047CB2A3BCFF55CF55FF3DD7B" pageId="44" pageNumber="45">
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<taxonomicName id="2F6D3DA2536BCF7377152B4EF438AC72" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium stadelmanni" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="stadelmanni">Mystrium stadelmanni</taxonomicName>
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:
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<bibRefCitation id="F36E0DEF53E698F1C9F10E6AE4B52E75" author="Forel, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="76" pageNumber="77" pagination="243 - 251" title="Nouvelles fourmis de l'Imerina oriental (Moramanga etc.)." volume="39" year="1895">Forel 1895</bibRefCitation>
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, in part.
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</paragraph>
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</subSubSection>
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<subSubSection id="EE3398B70030AFE54C0F3842BE29F346" pageId="44" pageNumber="45" type="lectotype">
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<paragraph id="965E7235BA3E8425D101DA7E4150EF1A" pageId="44" pageNumber="45">Lectotype</paragraph>
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<paragraph id="FC54A1A0639AFC2B3F43C2C49353DB02" pageId="44" pageNumber="45">[here designated]. Worker: CASENT0101670, MADAGASCAR, Amparafaravantsiv Moramangoufer [MHNG: examined].</paragraph>
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</subSubSection>
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<subSubSection id="8C98CA83CAF5E92D074BE85EB1B4C7ED" pageId="44" pageNumber="45" type="worker">
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<paragraph id="58D49DEDD7A4EC43C3D4E82BEEA6B13B" pageId="44" pageNumber="45">Worker.</paragraph>
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<paragraph id="FC9B306FD81839AAB854ABEF7ADE1BE4" pageId="44" pageNumber="45">Description. Measurements: lectotype. HL 1.87, HW 1.85, SL 1.13, ML 1.72, HD 1.24, WL 2.09, PnW 1.14, PpW 0.98, PtW 0.92, PtL 0.53, CI 98.7, SI 61.4, MI 93.3, PpI 86.5, PtI 174.6.</paragraph>
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<paragraph id="72DF11F19847E71122ACB9EF41D70F34" pageId="44" pageNumber="45">HL 1.19-2.27, HW 1.12-2.28, SL 0.70-1.37, ML 1.18-2.38, HD 0.83-1.47, WL 0.18-2.52, PnW 0.67-1.23, PpW 0.57-1.05, PtW 0.51-1.04, PtL 0.33-0.65, CI 93.9-101.4, SI 59.2-64.6, MI 90.9-105.7, PpI 79.3-90.1, PtI 153.1-182.6 (10 specimens measured).</paragraph>
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<paragraph id="7116445E2508B203950DE9E49D84F41E" pageId="44" pageNumber="45">
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Posterolateral corner of head strongly expanding posteriorly. Posterior face of vertex forming slightly blunt angle with dorsal face on median line of head, so that declivity of vertex on lateral part slightly steeper than on median part. Ventral half of vertex smooth and not sculptured. Eye small to moderately small. Anterior margin of clypeus straight or weakly concave and with small conical setae. Genal tooth of head undeveloped, but anterolateral corner of head angulate. Masticatory margin of mandible narrowly visible in full-face view, and difference in width of dorsal surface of mandible relatively small between mandibular shaft and distal portion. Second maxillary palpomere longer than third. First flagellomere (third antennal segment) as long as pedicel (second antennal segment). Shallow and fine longitudinal striae irregularly impressed on central part of pronotal dorsum, and sometimes shallowly reticulated around those striae. On lateral surface of pronotum, deep, fine longitudinal striae impressed and wide and deep punctures arranged on central horizontal line. Mesonotum differentiated from propodeum in dorsal view, length as long as that of propodeum in large individuals, shorter than propodeum in small individuals. Metanotal groove shallowly impressed, mesonotum in lateral view as high as pronotum in large individuals, higher than pronotum in small individuals. Metanotum weakly developed in largest individuals. Metapleural gland bulla developed, so that propodeal declivity in lateral view weakly convex posteriorly on ventral portion. Petiole in dorsal view moderately narrow, relatively narrower than that of
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<taxonomicName id="1D1A6E1D61666B5CCD087C44A685281B" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
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.
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</paragraph>
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<paragraph id="32B6E563D2E574DEBCDFD0D925BC50A5" pageId="44" pageNumber="45">Body color brown to black.</paragraph>
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</subSubSection>
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<subSubSection id="EDEB715849B9FDC9BBFF39B46C68BC41" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="queen">
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<paragraph id="4EEADE1249F6EB9B13B2A76C97A75E20" pageId="44" pageNumber="45">Queen.</paragraph>
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<paragraph id="8CC0C05E5AB29BD35D0A2BB904753153" pageId="44" pageNumber="45">Description. Measurements: HL 1.94-2.31, HW 1.99-2.40, SL 1.20-1.42, ML 1.87-2.48, HD 1.29-1.51, WL 2.78-3.31, MnW 1.42-1.83, PtW 1.03-1.32, PtL 0.61-0.78, CI 99.8-104.2, SI 58.1-62.8, MI 90.4-103.1, MnI 68.1-76.1, PtI 155.3-185.1 (10 specimens measured).</paragraph>
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<paragraph id="CBB40661A018EBD064F78F5AF91A1091" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
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Wings usually present and well developed; but lacking in intercaste. Wing sclerites fully developed even if wings have dropped off in queen; developed to undeveloped in variable degrees in intercaste. Posterolateral corner of head strongly expanding posteriorly, expansion not differentiated from that of workers. Posterior face of vertex forming slightly blunt angle with dorsal face on median line of head, so that declivity of vertex on lateral part slightly steeper than on median part. Ventral half of vertex smooth and not sculptured. Eye well developed. Both anterior and lateral ocelli clearly present in queen; ocelli varied from absent to developed in intercaste. Anterior margin of clypeus straight or weakly concave with small conical setae. Anterolateral portion of
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<pageBreakToken id="6C5D3212657033C58B957858C90E4F96" pageId="45" pageNumber="46" start="start">head</pageBreakToken>
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weakly to strongly angulate or with short spine. Masticatory margin of mandible narrowly, slightly visible in full-face view, dorsal surface on distal portion moderately wider than on mandibular shaft, difference in width much smaller than that in
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<taxonomicName id="886C5A7D7C2121BED3B3DD95F84DA30F" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
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. Teeth always present on whole mandibular shaft. Spatulate seta present on basal side of each basal denticle on masticatory margin of mandible. First flagellar segment on antenna as long as pedicel. Setae on pronotum spatulate, widened distally with sharp or blunt apex. Propodeal declivity in lateral view almost straight and forming right to slightly obtuse angle with its dorsal margin, of which ventral portion with small convexity by metapleural gland. Petiole relatively long in dorsal view, about 0.6
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<normalizedToken id="A35FFFCEEA6C854F4B7C66076F776CDF" originalValue="×">x</normalizedToken>
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length of abdominal segment III.
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</paragraph>
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<paragraph id="722450E1949FA1B71784C4B823272421" pageId="45" pageNumber="46">Body color black.</paragraph>
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</subSubSection>
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<subSubSection id="4B9086A4F6513FCE178B8763AF7D7C7F" pageId="45" pageNumber="46" type="male">
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<paragraph id="0A6191143B6AB18358BC1CC28C748816" pageId="45" pageNumber="46">Male.</paragraph>
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<paragraph id="7D8B6803F5B7730E06CA2472859CF94E" pageId="45" pageNumber="46">Description. Measurements: HL 1.10-1.41, HW 1.43-1.85, SL 0.25-0.34, EL 0.59-0.76, WL 2.27-3.03, MnW 1.46-1.96, CI 127.9-132.0, SI 16.2-20.0, EI 49.9-54.1, MnI 101.4-107.6 (6 specimens measured).</paragraph>
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<paragraph id="2990BC306160A5C54FB1AFB3EE8FCCDC" pageId="45" pageNumber="46">
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Eye relatively small, occupying about half of head length. Ocelli relatively distant from dorsal margin of head in full-face view, or just failing to reach dorsal margin. Dorsal margin of head in full-face view rounded. Both anterior and lateral ocelli small. Distance between lateral ocellus and eye long, 3
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<normalizedToken id="93823E80047EB1467315EB844018B028" originalValue="×">x</normalizedToken>
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longer than diameter of lateral ocellus. Posterior half of vertex clearly differentiated from dorsal half, dorsal face almost as long as posterior face. Palpal formula 4,3. First segment of maxillary palp flattened and distinctly wider than second segment. Second maxillary palpomere longer than third. Notauli clearly impressed on mesoscutum. Petiole in dorsal view thin, its length about 0.5-0.65
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<normalizedToken id="76E054F1A6B67A04F5BB092C70B137E7" originalValue="×">x</normalizedToken>
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that of abdominal tergite III. Petiolar dorsum covered with rough, deep punctures. Abdominal tergum VIII roughly and deeply punctured.
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</paragraph>
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<paragraph id="107F028C8A9D9CA3ED6D05F6ABFB9A2C" pageId="45" pageNumber="46">Abdominal sternum IX punctured on its distal portion. Basal ring short, not extending basally. Telomere extending slightly further distally than digitus. Basoventral expansion of aedeagus less developed basoventrally, as long as dorsal extension. Ventral margin of aedeagus gently curved ventrally in lateral view. Aedeagus weakly narrowing distally, its distal portion widely rounded.</paragraph>
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<paragraph id="BAAA114C8B60C3D0D2EDE1BCDF270D5A" pageId="45" pageNumber="46">On forewing, cu-a located at junction of Media (M) and Cubitus (Cu).</paragraph>
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<paragraph id="150F6BD3C5F1D04D312FE2C4B8EE17C4" pageId="45" pageNumber="46">Body color reddish brown to black.</paragraph>
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</subSubSection>
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<subSubSection id="FA709A6F418264AA499B32993926A42F" pageId="45" pageNumber="46" type="distribution">
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<paragraph id="7C2A2AE68DB297D7C28D9868DF7FDB7A" pageId="45" pageNumber="46">Distribution.</paragraph>
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<paragraph id="3FB336E3D2040B047312792E92DA46EE" pageId="45" pageNumber="46">MADAGASCAR and COMOROS: as in Figure 49B.</paragraph>
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</subSubSection>
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<subSubSection id="85D58D27EE547B99DAD98140440F5D1E" lastPageId="49" lastPageNumber="50" pageId="45" pageNumber="46" type="additional material examined">
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<paragraph id="B09CDA9AD130FE3BBB5CF41D11264658" pageId="45" pageNumber="46">Additional material examined.</paragraph>
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<paragraph id="5CF4702869314FEE873BDF9BEE07E5AF" lastPageId="49" lastPageNumber="50" pageId="45" pageNumber="46">
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In addition to the type material, specimens from the following localities were examined in this study: MADAGASCAR. Diego-Suarez. Sakalava Beach [vegetated beach dunes] (-12.26278°, 49.3975°), across sandy trail in dwarf littoral forest, 10 m alt.; 7 km N Joffreville [camp 2 of Fisher] (-12.33333°, 49.25°), in dry forest, 360 m alt.; Parc National Montagne
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<normalizedToken id="000A94217F031FB7EB5DD46B6FAB95CB" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
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[lemur trail] (-12.51667°, 49.18333°), rainforest, 975 m alt.; Parc National Montagne
|
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<normalizedToken id="85C029C8AE3DB118448BBEB50718675B" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
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[Petit Lac road] (-12.52028°, 49.17917°), rainforest, 1125 m alt.; Antsiranana. Parc National Montagne
|
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<normalizedToken id="B1BDD03FE6D41C0CF22860773AED2A10" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
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, Antomboka (-12.51269°, 49.17807°), montane rainforest, 970 m alt.; Parc National Montagne
|
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<normalizedToken id="7DF93107DBC345590C693CE483563EFE" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
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(-12.51389°, 49.17784°), montane rainforest, 984 m alt.; Pic Bades (-12.5186°, 49.18625°), montane rainforest, 900 m alt.; (-12.52306°, 49.17901°), montane rainforest, 1100 m alt.; Roussettes (-12.52574°, 49.17238°)
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<pageBreakToken id="4803753520411BB8E59A061B2344D611" pageId="46" pageNumber="47" start="start">,</pageBreakToken>
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montane rainforest, 1025 m alt.; Roussettes Camp 7 km SW Park entrance (-12.51444°, 49.18139°), rainforest, 960 m alt.; Mahasarika (-12.53176°, 49.17662°), montane rainforest, 1135 m alt.; (-12.53417°, 49.17607°), montane rainforest, 1325 m alt.; 3.6 km 235° SW Joffreville (-12.53444°, 49.1795°), montane rainforest, 925 m alt.; Petit lac (-12.53664°, 49.17412°), montane rainforest, 1130 m alt.; Lac Maudit (-12.58502°, 49.15147°), montane rainforest, 1250 m alt.; 12.2 km 211° SSW Joffreville (-12.59639°, 49.1595°), montane rainforest, 1300 m alt.;
|
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<normalizedToken id="429A2917977DCB7BECFB4E1279706810" originalValue="Forêt">Foret</normalizedToken>
|
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de Binara, 9.1 km 233° SW Daraina (-13.26333°, 49.60333°), rainforest, 800 m alt.; (-13.26333°, 49.6°), montane rainforest, 1100 m alt.; Nosy Faly (-13.3624°, 48.49101°), open secondary vegetation, 15 m alt.; Galoko chain, Mont Galoko (-13.58487°, 48.71818°), rainforest, 520 m alt.; Ampasindava,
|
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<normalizedToken id="699E0C0D9FCB618A716E166E7B8B4BD8" originalValue="Forêt">Foret</normalizedToken>
|
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|
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<normalizedToken id="9D3643240B98EF37DAC834820DCFCCF0" originalValue="d’Ambilanivy">d'Ambilanivy</normalizedToken>
|
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, 3.9 km 181° S Ambaliha (-13.79861°, 48.16167°), rainforest, 600 m alt.; R.S. Manongarivo, 12.8 km 228° SW Antanambao (-13.97667°, 48.42333°), rainforest, 780 m alt.; 14.5 km 220° SW Antanambao (-13.99833°, 48.42833°), montane rainforest, 1175 m alt.; Parc National de Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina (-14.435°, 49.76°), rainforest, 775 m alt.; 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina (-14.43667°, 49.775°), rainforest, 450 m alt.; Antranohofa, 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina (-14.44333°, 49.74333°), montane rainforest, 1325 m alt.; Betaolana Forest, along Bekona River (-14.52996°, 49.44039°), rainforest, 880 m alt.;
|
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<normalizedToken id="85D9E508EFBBE2D2FF4CB611FC5B41F6" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Ambanitaza, 26.1 km 347° Antalaha (-14.67933°, 50.18367°), rainforest, 240 m alt.; Toamasina. Montagne
|
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<normalizedToken id="6D052379C7AE6274C37EA78583DDDB15" originalValue="d’Anjanaharibe">d'Anjanaharibe</normalizedToken>
|
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|
, 18.0 km 21° NNE Ambinanitelo (-15.18833°, 49.615°), rainforest, 470 m alt.; Res. Ambodiriana, 4.8 km 306°Manompana, along Manompana river (-16.67233°, 49.70117°), rainforest, 125 m alt.;
|
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<normalizedToken id="59BBABC6BF3E492F0F529CB00593C227" originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
<normalizedToken id="7BA7571802291D5BB4922FE60BCCC265" originalValue="Spéciale">Speciale</normalizedToken>
|
|||
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Ambatovaky, Sandrangato river (-16.7633°, 49.26692°), rainforest, 520 m alt.; (-16.7702°, 49.26638°), rainforest, 470 m alt.; (-16.77274°, 49.26551°), rainforest, 450 m alt.; Manakambahiny, near Vavatenina Forest (-17.46667°, 49.35°), alt.; Parc National de Zahamena (-17.73359°, 48.72625°), rainforest, 950 m alt.; Tetezambatana forest, near junction of Nosivola and Manakambahiny Rivers (-17.74298°, 48.72936°), rainforest, 860 m alt.; Onibe River (-17.75908°, 48.85468°), rainforest, 780 m alt.; Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina (-17.88667°, 49.2025°), rainforest, 520 m alt.;
|
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<normalizedToken id="FA05C79676B56CF3FB7AB467A3D584B3" originalValue="Réserve">Reserve</normalizedToken>
|
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|
Nationale
|
|||
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|
<normalizedToken id="7B30CF5388393D06CA85991D525BAD7A" originalValue="Intégrale">Integrale</normalizedToken>
|
|||
|
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Betampona, Betampona 35.1 km NW Toamasina (-17.91801°, 49.20074°), rainforest, 500 m alt.;
|
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<normalizedToken id="F9F0BE1FE69B793E9E7138AD27436917" originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
Naturelle Betampona, 34.08 km 332° Toamasina (-17.91977°, 49.20039°), rainforest, 525 m alt.; Reserve Betampona, Camp Rendrirendry 34.1 km 332° Toamasina (-17.924°, 49.19967°), rainforest, 390 m alt.; F.C. Sandranantitra (-18.04833°, 49.09167°), rainforest, 450 m alt.; Station
|
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<normalizedToken id="BD73F31E5FC0907AC77413E3B5F933E1" originalValue="forestière">forestiere</normalizedToken>
|
|||
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|
Analamazaotra, Analamazaotra 1.3 km S Andasibe (-18.38466°, 48.41271°), montane rainforest, 980 m alt.; Ankerana (-18.40062°, 48.81311°), rainforest, 865 m alt.; Ankerana (-18.4017°, 48.80605°), montane forest, 1035 m alt.; (-18.4061°, 48.82029°), rainforest, 725 m alt.; (-18.40829°, 48.82107°), rainforest, 750 m alt.; (-18.4104°, 48.8189°), rainforest, 855 m alt.; F.C. Andriantantely (-18.695°, 48.81333°), rainforest, 530 m alt.; Corridor Forestier Analamay-Mantadia, Tsaravoniana (-18.75641°, 48.42195°), rainforest, 1036 m alt.; (-18.75737°, 48.42302°), rainforest, 1018 m alt.; (-18.76124°,
|
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<pageBreakToken id="FAD696D80B1F9145D1FD93448D16A977" pageId="47" pageNumber="48" start="start">48.42134</pageBreakToken>
|
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°), rainforest, 939 m alt.; (-18.76465°, 48.41938°), rainforest, 1039 m alt.; Ambohibolakely (-18.76087°, 48.37128°), rainforest, 1044 m alt.; (-18.76131°, 48.36437°), rainforest, 983 m alt.; (-18.77898°, 48.36375°), rainforest, 918 m alt.; Ambohibolakely (-18.77908°, 48.36628°), rainforest, 1014 m alt.; Ambatoharanana (-18.79956°, 48.4028°), rainforest, 1058 m alt.; (-18.80388°, 48.40506°), rainforest, 1013 m alt.; (-18.80398°, 48.40358°), rainforest, 1064 m alt.; (-18.80424°, 48.40081°), rainforest, 968 m alt.; (-18.80438°, 48.40735°), rainforest, 960 m alt.; Bevolota 17.1 km N Andasibe (-18.77071°, 48.43164°), montane rainforest, 995 m alt.; Analamay (-18.80623°, 48.33707°), montane rainforest, 1068 m alt.; Parc National
|
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|
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<normalizedToken id="88FC4F3C37F3AB7AA0A5B368F729EABF" originalValue="d´">d'</normalizedToken>
|
|||
|
|
Andasibe-Mantadia,
|
|||
|
|
<normalizedToken id="584A656FA63C2AA2B69DADD358F67153" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
de Mantadia, 25.7 km 248° Moramanga (-18.81402°, 48.43028°), rainforest, 1040 m alt.; Ambatovy, 12.4 km NE Moramanga (-18.83937°, 48.30842°), montane rainforest, 1080 m alt.;
|
|||
|
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<normalizedToken id="3DB185D259B157CF12488F0A82A37ABD" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Ambatovy, 14.3 km 57° Moramanga (-18.85083°, 48.32°), montane rainforest, 1075 m alt.; Torotorofotsy (-18.87082°, 48.34737°), montane rainforest, marsh edge, 1070 m alt.; Toliara.
|
|||
|
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<normalizedToken id="36E300932CB279451CEB9DFC90AB41EC" originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
<normalizedToken id="0D733DAF6B23B2A5D260BDA8CE4F3E88" originalValue="Spéciale">Speciale</normalizedToken>
|
|||
|
|
<normalizedToken id="9808868C509361053682E68131A0236B" originalValue="d’Ambohijanahary">d'Ambohijanahary</normalizedToken>
|
|||
|
|
,
|
|||
|
|
<normalizedToken id="F1459E60A35BDF1542D0F51A1EEFB3B2" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
<normalizedToken id="7968866B91AEDE57C144CB3238BC3B63" originalValue="d’Ankazotsihitafototra">d'Ankazotsihitafototra</normalizedToken>
|
|||
|
|
, 35.2 km 312° NW Ambaravaranala (-18.26667°, 45.40667°), montane rainforest, 1050 m alt.; Makay Mts. (-21.20978°, 45.34184°), Gallery forest on sandy soil, 525 m alt.; (-21.21013°, 45.35462°), Gallery forest with bamboo, 540 m alt.; (-21.22336°, 45.32628°), Gallery forest on sandy soil, 480 m alt.; (-21.227°, 45.33222°), gallery forest on sandy soil, 475 m alt.;
|
|||
|
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<normalizedToken id="B7970B0C359CC26CC95E000D4115B8A3" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Ivohibe 55.6 km N Tolagnaro (-24.56167°, 47.20017°), rainforest, 650 m alt.; 11 km NW Enkara,
|
|||
|
|
<normalizedToken id="6FE5F6A56512AB7BD4609D23516B0280" originalValue="Rés">Res</normalizedToken>
|
|||
|
|
Andohahela (-24.56667°, 46.81667°), rainforest, 950 m alt.;
|
|||
|
|
<normalizedToken id="830D0752B2C01CA09DC2A423EDD37A55" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Ivohibe 55.0 km N Tolagnaro (-24.569°, 47.204°), rainforest, 200 m alt.; 6 km SSW Eminiminy, Res. Andohahela (-24.73333°, 46.8°), wet forest, 330 m alt.;
|
|||
|
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<normalizedToken id="6CDA0475C4FF6B7A44D95B4AB8C914B8" originalValue="Rés">Res</normalizedToken>
|
|||
|
|
. Andohahela, 6 km SSW Eminiminy (-24.73333°, 46.8°), rainforest, 330 m alt.; Parc National Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro (-24.7585°, 46.85367°), rainforest, 275 m alt.; Manampanihy River, 5.4 km 113° ESE Mahamavo, 36.7 km 343° NNW Tolagnaro (-24.76389°, 46.76683°), rainforest, 650 m alt.; Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro (-24.76389°, 46.75167°), montane rainforest, 900 m alt.;
|
|||
|
|
<normalizedToken id="764C911B67A465C964AF9D0DCC5BC709" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Mandena 8.5 km N Tolagnaro (-24.95267°, 47.0025°), littoral rainforest, 20 m alt.; Grand Lavasoa, 25.9 km W Tolagnaro (-25.08767°, 46.749°), rainforest, 450 m alt.; Toamasina. Res. Ambodiriana, 4.8 km 306° Manompana, along Manompana river (-16.67233°, 49.70117°), rainforest, 125 m alt.;
|
|||
|
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<normalizedToken id="F067982EFDCBDA1C1712A816FDCEA689" originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
<normalizedToken id="0CC16F7C3892F09D17D92C9E90B24609" originalValue="Spéciale">Speciale</normalizedToken>
|
|||
|
|
Ambatovaky, Sandrangato river (-16.7633°, 49.26692°), rainforest, 520 m alt.; (-16.7702°, 49.26638°), rainforest, 470 m alt.; (-16.77274°, 49.26551°), rainforest, 450 m alt.; Manakambahiny, near Vavatenina Forest (-17.46667°, 49.35°), alt.; Parc National de Zahamena (-17.73359°, 48.72625°), rainforest, 950 m alt.; Tetezambatana forest, near junction of Nosivola and Manakambahiny Rivers (-17.74298°, 48.72936°), rainforest, 860 m alt.; Onibe River (-17.75908°, 48.85468°), rainforest, 780 m alt.; Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina (-17.88667°, 49.2025°), rainforest, 520 m alt.;
|
|||
|
|
<normalizedToken id="28B90E9937CD80E67D56940B1E731F32" originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
Nationale
|
|||
|
|
<normalizedToken id="A06CF68C79A9D1EEBB04BFEFB7541459" originalValue="Intégrale">Integrale</normalizedToken>
|
|||
|
|
Betampona, Betampona 35.1 km NW Toamasina (-17.91801°, 49.20074°), rainforest, 500 m alt.;
|
|||
|
|
<normalizedToken id="7E57A7523E48710FE363BD5981E257EE" originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
Naturelle Betampona, 34.08 km 332° Toamasina (-17.91977°, 49.20039°), rainforest, 525 m alt.; Reserve Betampona, Camp Rendrirendry 34.1 km 332° Toamasina
|
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<pageBreakToken id="DDE527A4C01E3900516313AE70EFA057" pageId="48" pageNumber="49" start="start">(</pageBreakToken>
|
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-17.924°, 49.19967°), rainforest, 390 m alt.; F.C. Sandranantitra (-18.04833°, 49.09167°), rainforest, 450 m alt.; Station
|
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|
<normalizedToken id="9CAEC03C444C0D195E0C29725DE04678" originalValue="forestière">forestiere</normalizedToken>
|
|||
|
|
Analamazaotra, Analamazaotra 1.3 km S Andasibe (-18.38466°, 48.41271°), montane rainforest, 980 m alt.; Ankerana (-18.40062°, 48.81311°), rainforest, 865 m alt.; Ankerana (-18.4017°, 48.80605°), montane forest, 1035 m alt.; (-18.4061°, 48.82029°), rainforest, 725 m alt.; (-18.40829°, 48.82107°), rainforest, 750 m alt.; (-18.4104°, 48.8189°), rainforest, 855 m alt.; F.C. Andriantantely (-18.695°, 48.81333°), rainforest, 530 m alt.; Corridor Forestier Analamay-Mantadia, Tsaravoniana (-18.75641°, 48.42195°), rainforest, 1036 m alt.; (-18.75737°, 48.42302°), rainforest, 1018 m alt.; (-18.76124°, 48.42134°), rainforest, 939 m alt.; (-18.76465°, 48.41938°), rainforest, 1039 m alt.; Ambohibolakely (-18.76087°, 48.37128°), rainforest, 1044 m alt.; (-18.76131°, 48.36437°), rainforest, 983 m alt.; (-18.77898°, 48.36375°), rainforest, 918 m alt.; Ambohibolakely (-18.77908°, 48.36628°), rainforest, 1014 m alt.; Ambatoharanana (-18.79956°, 48.4028°), rainforest, 1058 m alt.; (-18.80388°, 48.40506°), rainforest, 1013 m alt.; (-18.80398°, 48.40358°), rainforest, 1064 m alt.; (-18.80424°, 48.40081°), rainforest, 968 m alt.; (-18.80438°, 48.40735°), rainforest, 960 m alt.; Bevolota 17.1 km N Andasibe (-18.77071°, 48.43164°), montane rainforest, 995 m alt.; Analamay (-18.80623°, 48.33707°), montane rainforest, 1068 m alt.; Parc National
|
|||
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|
<normalizedToken id="0D6A28190357827E86F4447A4F9D002D" originalValue="d´">d'</normalizedToken>
|
|||
|
|
Andasibe-Mantadia,
|
|||
|
|
<normalizedToken id="DE0BDA37C986858DEBEDF00EB8C5CF16" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
de Mantadia, 25.7 km 248° Moramanga (-18.81402°, 48.43028°), rainforest, 1040 m alt.; Ambatovy, 12.4 km NE Moramanga (-18.83937°, 48.30842°), montane rainforest, 1080 m alt.;
|
|||
|
|
<normalizedToken id="8C08353760D397C51D4C15C9A3A6A6CF" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Ambatovy, 14.3 km 57° Moramanga (-18.85083°, 48.32°), montane rainforest, 1075 m alt.; Torotorofotsy (-18.87082°, 48.34737°), montane rainforest, marsh edge, 1070 m alt.; Toliara.
|
|||
|
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<normalizedToken id="BEFBE8B72177353B46D13339A5E64AD1" originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
<normalizedToken id="1D7DDECF963D671505BD08201FAEE85C" originalValue="Spéciale">Speciale</normalizedToken>
|
|||
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|
<normalizedToken id="51B2BE6017C028163119A413DC4930EA" originalValue="d’Ambohijanahary">d'Ambohijanahary</normalizedToken>
|
|||
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,
|
|||
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|
<normalizedToken id="9294E33C03AC483F8936506A59394240" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
<normalizedToken id="46E6E91C74E7BC898361137E38BEE906" originalValue="d’Ankazotsihitafototra">d'Ankazotsihitafototra</normalizedToken>
|
|||
|
|
, 35.2 km 312° NW Ambaravaranala (-18.26667°, 45.40667°), montane rainforest, 1050 m alt.; Makay Mts. (-21.20978°, 45.34184°), gallery forest on sandy soil, 525 m alt.; (-21.21013°, 45.35462°), gallery forest with bamboo, 540 m alt.; (-21.22336°, 45.32628°), gallery forest on sandy soil, 480 m alt.; (-21.227°, 45.33222°), gallery forest on sandy soil, 475 m alt.;
|
|||
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<normalizedToken id="96E4266EA1B487AAA6C9CA45A4CF0B04" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Ivohibe 55.6 km N Tolagnaro (-24.56167°, 47.20017°), rainforest, 650 m alt.; 11 km NW Enkara,
|
|||
|
|
<normalizedToken id="01A1B3F344C8A8F42F95153AA78C45E3" originalValue="Rés">Res</normalizedToken>
|
|||
|
|
Andohahela (-24.56667°, 46.81667°), rainforest, 950 m alt.;
|
|||
|
|
<normalizedToken id="4B1ADE3BB652BE4BED38959BBE200F8A" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
Ivohibe 55.0 km N Tolagnaro (-24.569°, 47.204°), rainforest, 200 m alt.; 6 km SSW Eminiminy, Res. Andohahela (-24.73333°, 46.8°), wet forest, 330 m alt.;
|
|||
|
|
<normalizedToken id="58021DE599FA462690F137FE2B17E512" originalValue="Rés">Res</normalizedToken>
|
|||
|
|
. Andohahela, 6 km SSW Eminiminy (-24.73333°, 46.8°), rainforest, 330 m alt.; Parc National Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro (-24.7585°, 46.85367°), rainforest, 275 m alt.; Manampanihy River, 5.4 km 113° ESE Mahamavo, 36.7 km 343° NNW Tolagnaro (-24.76389°, 46.76683°), rainforest, 650 m alt.; Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro (-24.76389°, 46.75167°), montane rainforest, 900 m alt.;
|
|||
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<normalizedToken id="CC3EF3C346D2AEA7E17F8936CD2F5544" originalValue="Forêt">Foret</normalizedToken>
|
|||
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|
Mandena 8.5 km N Tolagnaro (-24.95267°, 47.0025°), littoral rainforest, 20 m alt.; Grand Lavasoa, 25.9 km W Tolagnaro (-25.08767°, 46.749°), rainforest, 450 m alt. Mahajanga.
|
|||
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<normalizedToken id="811045C54E3A9ED2891052907F8FF3B4" originalValue="Réserve">Reserve</normalizedToken>
|
|||
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|
<normalizedToken id="280319C3C6BD26C4725BFA604880816D" originalValue="Spéciale">Speciale</normalizedToken>
|
|||
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Marotandrano, Marotandrano 48.3 km S Mandritsara (-16.28322°, 48.81443°), transition humid forest, 865 m alt. Fianarantsoa. Ranomafana Nat. Park, Miaranony Forest (-21.25°, 47.41667°), montane forest, 700 m alt.; 7 km W Ranomafana (-21.26667°, 47.41667°), montane rainforest, 1000
|
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<pageBreakToken id="4D08B7ED7E715BFBB50985A468A2C5CE" pageId="49" pageNumber="50" start="start">m</pageBreakToken>
|
|||
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|
alt.; Parc National de Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana (-21.29°, 47.43333°), montane rainforest, 1100 m alt.; 7.6 km 122° Kianjavato,
|
|||
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<normalizedToken id="AF7CCB133FD9C411E9F665E5DB93D832" originalValue="Forêt">Foret</normalizedToken>
|
|||
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|
<normalizedToken id="3E0D17F483C36728F8D7597D343D654B" originalValue="Classée">Classee</normalizedToken>
|
|||
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|
Vatovavy (-21.4°, 47.94°), rainforest, 175 m alt.; Andrambovato along river Tatamaly (-21.50967°, 47.40762°), cultivated land (tavy), 984 m alt.; (-21.51089°, 47.40987°), rainforest, 994 m alt.; 2 km W Andrambovato, along river Tatamaly (-21.51167°, 47.41°), montane rainforest, 1075 m alt.; 45 km S. Ambalavao (-22.21667°, 47.01667°), rainforest, 785 m alt.; 43 km S Ambalavao,
|
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<normalizedToken id="D970A3D48C1FE6D5AD1694334DE440F6" originalValue="Rés">Res</normalizedToken>
|
|||
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|
. Andringitra (-22.23333°, 47°), rainforest, 825 m alt.; 9.0 km NE Ivohibe (-22.42667°, 46.93833°), montane rainforest, 900 m alt.; Parc National
|
|||
|
|
<normalizedToken id="0E3ECB614EA38ED145D5239F3E642F5E" originalValue="d’Isalo">d'Isalo</normalizedToken>
|
|||
|
|
, 9.1 km 354° N Ranohira (-22.48167°, 45.46167°), gallery forest, 725 m alt.; R.S. Ivohibe 8.0 km E Ivohibe (-22.48333°, 46.96833°), montane rainforest, 1200 m alt.;
|
|||
|
|
<normalizedToken id="2D56E77146BAB6687764AD1338061829" originalValue="Forêt">Foret</normalizedToken>
|
|||
|
|
de Vevembe, 66.6 km 293° Farafangana (-22.791°, 47.18183°), rainforest, transition to montane forest, 600 m alt.;
|
|||
|
|
<normalizedToken id="F3FC5BC2E1138C323C97A92737EDC1A1" originalValue="Réserve">Reserve</normalizedToken>
|
|||
|
|
Speciale Manombo 24.5 km 228° Farafangana (-23.01583°, 47.719°), rainforest, 30 m alt.; Parc National Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango (-23.83517°, 46.96367°), rainforest, 940 m alt.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="67C4952A69F1BDD34855E52F200BF3F0" pageId="49" pageNumber="50">
|
|||
|
|
COMOROS.
|
|||
|
|
<normalizedToken id="B3A60E892E2783E2EA54CE9365F2DC68" originalValue="Mohéli">Moheli</normalizedToken>
|
|||
|
|
, Ouallah (-12.32717°, 43.65952°), coastal scrub, 10 m alt.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="793A1658410EEDA2F01BA5F6FC6C99DA" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="remarks">
|
|||
|
|
<paragraph id="1E2A37CF04F6694EE37BED3AE44B4629" pageId="49" pageNumber="50">Remarks.</paragraph>
|
|||
|
|
<paragraph id="C6D0ED75B4BC8B3DFEBB9B1C35F111D1" pageId="49" pageNumber="50">
|
|||
|
|
The females of
|
|||
|
|
<taxonomicName id="D5D7CBAAEAE81A814D5D1E52CF1F6A22" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
can be distinguished easily from other
|
|||
|
|
<taxonomicName id="409820D8442758E1530D6A3B14035F1F" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Mystrium</taxonomicName>
|
|||
|
|
females by the combination of a distinct longitudinal carina on the central portion of the labrum (Fig. 8A) or a single lateral spine on abdominal sternum VII (Fig. 8C), and the straight anterior margin of the clypeus (Fig. 9D). In addition to these diagnostic characters, a combination of unsculptured and smooth areas on the ventral portion of the vertex (as in Fig. 11A) and the straight anterior clypeal margin separates
|
|||
|
|
<taxonomicName id="A258344AC7F8E0B88C1146851EC394C1" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
from the other
|
|||
|
|
<taxonomicName id="D73735DD2055ECE037F9162D28804CCE" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Mystrium</taxonomicName>
|
|||
|
|
species workers; and a larger-sized alate queen (HW>1.99) with spatulate setae on the pronotal dorsum (Fig. 9B) separate
|
|||
|
|
<taxonomicName id="73D7BB8B454CB7B1637992C870C5D646" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
from the queens of other species. For males, the small eye and small ocelli (as in Fig. 26B) that are separated from each other by a distance of more than 3
|
|||
|
|
<normalizedToken id="4E8125D780B113287A4B295C6AF9ACC4" originalValue="×">x</normalizedToken>
|
|||
|
|
the maximum diameter of lateral ocellus, petiolar dorsum (as in Fig. 28C) and abdominal tergum VIII (as in Fig. 28A) with rough and deep punctures, and less developed basoventral expansion of aedeagus (Fig. 29B) separate
|
|||
|
|
<taxonomicName id="AEB0E9BB818BDE1E667E392A286FEF61" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
from the other
|
|||
|
|
<taxonomicName id="9021A375F4E2F3D74B746459250AE725" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Mystrium</taxonomicName>
|
|||
|
|
males in the Malagasy region.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="7E2D5F5FF9B8B58CD4BF53DB619F8B8C" pageId="49" pageNumber="50">
|
|||
|
|
We propose a new set of diagnostic characters in this study to correctly identify
|
|||
|
|
<taxonomicName id="78F07E921276863D15699A5691F83141" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
. The diagnostic characters listed above are constant in females regardless of their body size. Because of the remarkable morphological variation in each species, most diagnostic characters for
|
|||
|
|
<taxonomicName id="9CC9424643E224A0D208B17D4D5961F0" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
proposed in previous papers (e.g.
|
|||
|
|
<bibRefCitation id="BF33BD4B0170C59B1ECFB05C95A5FE28" author="Emery, C" journalOrPublisher="Bullettino della Societa Entomologica Italiana" pageId="76" pageNumber="77" pagination="263 - 290" title="Formiche di Madagascar raccolte dal Sig. A. Mocquerys nei pressi della Baia di Antongil (1897 - 1898)." volume="31" year="1899">Emery 1899</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="3B5B5E5B108B00D0EFDC60F1C006DFD4" author="Menozzi, C" journalOrPublisher="Zoologischer Anzeiger" pageId="76" pageNumber="77" pagination="518 - 536" title="Revisione delle formiche del genere Mystrium Roger." volume="82" year="1929">Menozzi 1929</bibRefCitation>
|
|||
|
|
) cannot separate
|
|||
|
|
<taxonomicName id="435CF03F07C93C4490B6321C03F94A48" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
from the other
|
|||
|
|
<taxonomicName id="4B3F2CC94F1F840D37FFC2DF35DBB4DF" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">Mystrium</taxonomicName>
|
|||
|
|
species, especially from
|
|||
|
|
<taxonomicName id="62A33F9FC5FBFB232C5803F871DFBD30" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="9221D5DE745E523A2F7E2CCABC824DD3" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
also displays distinct variation in body size, color, relative mandible length, and shape of the petiole. The difference in the petiolar shape (
|
|||
|
|
<bibRefCitation id="915A9789EBE4F1B42498009CDD6B46C3" author="Emery, C" journalOrPublisher="Bullettino della Societa Entomologica Italiana" pageId="76" pageNumber="77" pagination="263 - 290" title="Formiche di Madagascar raccolte dal Sig. A. Mocquerys nei pressi della Baia di Antongil (1897 - 1898)." volume="31" year="1899">Emery 1899</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="6AC627DC3DC1C3A826BA212DF9C43447" author="Menozzi, C" journalOrPublisher="Zoologischer Anzeiger" pageId="76" pageNumber="77" pagination="518 - 536" title="Revisione delle formiche del genere Mystrium Roger." volume="82" year="1929">Menozzi 1929</bibRefCitation>
|
|||
|
|
) cannot be used for complete separation between
|
|||
|
|
<taxonomicName id="E22D095FAAB5CAFC84986B99B306E179" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="B2D802DD71EDF49156739C93CF34D35E" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
(PtI 153-183 vs. PtI 146-168). As mentioned in their discussion of
|
|||
|
|
<taxonomicName id="FA73E22363DECF87A337B8FAAF95F992" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="65ABB9A21E13199149CEF7B41AD72186" author="Molet, M" journalOrPublisher="American Naturalist" pageId="76" pageNumber="77" pagination="328 - 341" title="Evolution of novel mosaic castes in ants: modularity, phenotypic plasticity, and colonial buffering." url="10.1086/667368" volume="180" year="2012">Molet et al. (2012)</bibRefCitation>
|
|||
|
|
reported "mosaic monster" intermediates between queen and worker. The existence of this intercaste is not unique to
|
|||
|
|
<taxonomicName id="F375F5D114EF4C5F23A529E81FA9F956" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
, but rather a character of the mysticum species group that we confirmed in
|
|||
|
|
<taxonomicName id="640FE776D060991D1566DE8B14CFAF2E" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
as well. In this study, the intercastes are keyed out with either of the workers or queens of each species.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="A8B79FB7008BFF0602E8BE3AFBA17A49" pageId="50" pageNumber="51">
|
|||
|
|
<pageBreakToken id="E9113B044F686A04DB50DC73CA069A81" pageId="50" pageNumber="51" start="start">As</pageBreakToken>
|
|||
|
|
we discussed in
|
|||
|
|
<taxonomicName id="F345F7D3145006723267FB1560DC9B6D" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
, further studies are necessary to better separate the males of
|
|||
|
|
<taxonomicName id="2BCCB6324E8A36E21C26FD4033D9E820" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="51FBBD298BBDD9053F720682E7DDFC72" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
. Here we propose only an aedeagal character to separate these very similar males (Fig. 29).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="AE31E2F99F61C69E551303596CBCC0DE" pageId="50" pageNumber="51">
|
|||
|
|
The lectotype for
|
|||
|
|
<taxonomicName id="7ECF7520501FBF8987D2980C5777DDF9" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
is designated.
|
|||
|
|
<taxonomicName id="B8CF7D41602829E16ADCB2D381EC1478" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
was described by Forel in 1899 using at least two workers and one male collected in Madagascar, and we confirm one worker and one male in his collection. Although we could not find any information to identify which type specimens were used in his original description (
|
|||
|
|
<bibRefCitation id="711D3FD7957D55287B4A3E5834F9E6CA" author="Forel, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="76" pageNumber="77" pagination="303 - 310" title="Trois notices myrmecologiques." volume="43" year="1899">Forel 1899</bibRefCitation>
|
|||
|
|
), Forel mentioned that the workers he determined as
|
|||
|
|
<taxonomicName id="839E11BCA72A2D9461E9B1886213214D" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
were the same specimens he determined as
|
|||
|
|
<taxonomicName id="4CEDE187347C8C666A2B79BDB1540E0D" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="8BCA11BD2B249258E3527C3668F6611E" author="Forel, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="76" pageNumber="77" pagination="516 - 535" title="Nouvelles especes de Formicides de Madagascar (recoltees par M. Sikora). Premiere serie." volume="36" year="1892">Forel 1892</bibRefCitation>
|
|||
|
|
), and which Emery had sent him. According to the description in
|
|||
|
|
<bibRefCitation id="BF4180D74150B91E60D42BE473492A86" author="Forel, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="76" pageNumber="77" pagination="516 - 535" title="Nouvelles especes de Formicides de Madagascar (recoltees par M. Sikora). Premiere serie." volume="36" year="1892">Forel (1892)</bibRefCitation>
|
|||
|
|
, the worker of
|
|||
|
|
<taxonomicName id="967FDC1568B988FE0EE6F093081A5AC1" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
is smaller and stouter than the worker of
|
|||
|
|
<taxonomicName id="475F9A355FA21F779045DC5BBE90A6CE" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
; however, we could not deduce how he determined the boundary of "
|
|||
|
|
<taxonomicName id="E0AC3AEBC4D2F4031A34EC6C0C76D8E0" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
" in the worker caste. To further complicate the story, when Forel described
|
|||
|
|
<taxonomicName id="ED15A838537A87E4C3A452317804DA8F" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
in 1899, he probably determined workers of actual
|
|||
|
|
<taxonomicName id="FBD5D6952C11EE0C3C9A81BD011305B1" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
as
|
|||
|
|
<taxonomicName id="E40D00EF4F29581862C5029586CB88B9" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium stadelmanni" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="stadelmanni">Mystrium stadelmanni</taxonomicName>
|
|||
|
|
(see remarks for
|
|||
|
|
<taxonomicName id="D7D71D76AF32155B68493FCA2EC0EBB1" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
). Moreover, once he determined
|
|||
|
|
<normalizedToken id="C1EC4A57DD0DFAE16349AFB4D7A8AFC6" originalValue="Emery’s">Emery's</normalizedToken>
|
|||
|
|
material as
|
|||
|
|
<taxonomicName id="9C0C68447E35B10481BCD469128402C3" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
|
|||
|
|
, the worker caste of
|
|||
|
|
<taxonomicName id="79B720D49C7E1C4E5056113BB8B3DB5C" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
|
|||
|
|
should have been considered
|
|||
|
|
<normalizedToken id="C62B9EACA23E331E0FF391E199140E7F" originalValue="“unknown”">"unknown"</normalizedToken>
|
|||
|
|
once again. In addition to the misidentification of the worker caste, his description did not provide any details to identify which male specimens he studied. We also confirm
|
|||
|
|
<normalizedToken id="6605A10F434980DA6E41E05CF175ED0B" originalValue="Forel’s">Forel's</normalizedToken>
|
|||
|
|
suspicions (
|
|||
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<bibRefCitation id="BE46D7514D4111D07D5DB8E01E6B70DF" author="Forel, A" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="76" pageNumber="77" pagination="303 - 310" title="Trois notices myrmecologiques." volume="43" year="1899">Forel 1899</bibRefCitation>
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) that the male described as
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<taxonomicName id="DF67E6B5962E1ED63BBB2F35563B35EA" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
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is not conspecific with the worker. We verify that the male specimen in his collection, which was probably used for the description of
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<taxonomicName id="97A0F8BDEEFAFFD5A4CE99170188BF37" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
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, is
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<taxonomicName id="9094B0B525DA3CF22569A97429631D06" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium oberthueri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="oberthueri">Mystrium oberthueri</taxonomicName>
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.
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</paragraph>
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<paragraph id="7000DA139DFA3397700D18B03BD6B025" pageId="50" pageNumber="51">
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<taxonomicName id="5C6DB7E64CDDFA27A950036432A18788" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
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is one of the most common
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<taxonomicName id="4E93521E1BBBCABDC37CAD6A28709372" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">Mystrium</taxonomicName>
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species used in recent ecological and phylogenetic studies (
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<bibRefCitation id="C470DDA491C8E2003E64663EE17E3B34" author="Molet, M" journalOrPublisher="Biological journal of the Linnean Society" pageId="76" pageNumber="77" pagination="198 - 207" title="Shift from independent to dependent colony foundation and evolution of ' multi-purpose' ergatoid queens in Mystrium ants (subfamily Amblyoponinae)." url="10.1111/j.1095-8312.2009.01257.x" volume="98" year="2009">Molet et al. 2009</bibRefCitation>
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;
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<bibRefCitation id="AB199C44EC7BFF0FC0688BBB0CB72288" author="Molet, M" journalOrPublisher="Naturwissenschaften" pageId="76" pageNumber="77" pagination="280 - 287" title="Winged queens replaced by reproductives smaller than workers in Mystrium ants." url="10.1007/s00114-006-0190-2" volume="94" year="2007 a">Molet et al. 2007a</bibRefCitation>
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;
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<bibRefCitation id="9433D1730AC1F1803DAFE82042402B6E" author="Molet, M" journalOrPublisher="American Naturalist" pageId="76" pageNumber="77" pagination="328 - 341" title="Evolution of novel mosaic castes in ants: modularity, phenotypic plasticity, and colonial buffering." url="10.1086/667368" volume="180" year="2012">Molet et al. 2012</bibRefCitation>
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;
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<bibRefCitation id="8C2EA8DDD027E26EA296E86F048001B3" author="Moreau, CS" journalOrPublisher="Science" pageId="77" pageNumber="78" pagination="101 - 104" title="Phylogeny of the ants: diversification in the age of angiosperms." url="10.1126/science.1124891" volume="312" year="2006">Moreau et al. 2006</bibRefCitation>
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;
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<bibRefCitation id="262A2BD6E4F745A1365468244893BCF7" author="Saux, C" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="77" pageNumber="78" pagination="457 - 468" title="Dracula ant phylogeny as inferred by nuclear 28 S rDNA sequences and implications for ant systematics (Hymenoptera: Formicidae: Amblyoponinae)." url="10.1016/j.ympev.2004.06.017" volume="33" year="2004">Saux et al. 2004</bibRefCitation>
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). However, the identification of their voucher material is now reassessed: part of the material identified as
|
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<taxonomicName id="9295A28FF8479FCA2ACD0F394B3B6657" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
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in Molet et al. (
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<bibRefCitation id="D579FE2CE9058ECE0E9E7F80B7376EA5" author="Molet, M" journalOrPublisher="Biological journal of the Linnean Society" pageId="76" pageNumber="77" pagination="198 - 207" title="Shift from independent to dependent colony foundation and evolution of ' multi-purpose' ergatoid queens in Mystrium ants (subfamily Amblyoponinae)." url="10.1111/j.1095-8312.2009.01257.x" volume="98" year="2009">Molet et al. 2009</bibRefCitation>
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;
|
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<bibRefCitation id="718128A87AE7C126165DCE55621F622D" author="Molet, M" journalOrPublisher="Naturwissenschaften" pageId="76" pageNumber="77" pagination="280 - 287" title="Winged queens replaced by reproductives smaller than workers in Mystrium ants." url="10.1007/s00114-006-0190-2" volume="94" year="2007 a">Molet et al. 2007a</bibRefCitation>
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;
|
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<bibRefCitation id="1CB557897C61C9597207079E24A5671F" author="Molet, M" journalOrPublisher="American Naturalist" pageId="76" pageNumber="77" pagination="328 - 341" title="Evolution of novel mosaic castes in ants: modularity, phenotypic plasticity, and colonial buffering." url="10.1086/667368" volume="180" year="2012">Molet et al. 2012</bibRefCitation>
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) is not
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<taxonomicName id="53E1BE768280FA8623ACB7209353537A" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium rogeri" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="rogeri">Mystrium rogeri</taxonomicName>
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(see remarks under
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<taxonomicName id="E1B73E8E9FC2BAFC0C43B5319CBC6798" class="Insecta" family="Formicidae" genus="Mystrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mystrium mysticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="mysticum">Mystrium mysticum</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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