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<mods:title id="1D73EC192E5FFCADEF036901B4D0BFF0">Decapod assemblage from the late Miocene (early-middle Messinian) of the Romagna Apennines nearby Brisighella, Emilia-Romagna (N Italy)</mods:title>
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<taxonomicName id="0EAD4D5AFFC0314EA327B012FDA1F9F2" ID-CoL="BTRNJ" authority="Delle Cave, 1988" authorityName="Delle Cave" authorityYear="1988" box="[113,567,1581,1607]" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA327B012FEFBF9F2" box="[113,365,1581,1607]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
Delle Cave, 1988
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Delle Cave, 1988: 123-126
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, Pl. 1, figs. 1, 2, Pl. 2, figs. 1-5.
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<emphasis id="FBD9EACBFFC0314EA327B0DEFEF3F94E" box="[113,357,1761,1787]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
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– De Angeli &amp; Garassino 2006: 74. — De Angeli 
<emphasis id="FBD9EACBFFC0314EA258B13FFED0F8AC" box="[270,326,1791,1817]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2009: 185, 195, fig. 16. — 
<treatmentCitation id="480C10C8FFC0314EA12DB0C0FEB6F88D" author="Schweitzer C. E. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." page="126" pageId="3" pageNumber="30" year="2010">
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Schweitzer 
<emphasis id="FBD9EACBFFC0314EA327B121FF3EF88D" box="[113,168,1822,1848]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2010: 126
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. — 
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<bibRefCitation id="AD3C4B28FFC0314EA21BB121FDF7F88D" author="Garassino A. &amp; Pasini G. &amp; De Angeli A. &amp; Charbonnier S. &amp; Famiani F. &amp; Baldanza A. &amp; Bizzarri R." box="[333,609,1821,1848]" pageId="3" pageNumber="30" pagination="1 - 61" refId="ref5024" refString="Garassino A., Pasini G., De Angeli A., Charbonnier S., Famiani F., Baldanza A. &amp; Bizzarri R., 2012 - The decapod community from the Early Pliocene (Zanclean) of &quot; La Serra &quot; quarry (San Miniato, central Italy): sedimentology, systematics, and palaeoenvironmental implications. Annales de Paleontologie, 98: 1 - 61." type="journal article" year="2012">
Garassino 
<emphasis id="FBD9EACBFFC0314EA297B121FE6EF88D" box="[449,504,1822,1848]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2012: 52
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. — Baldanza 
<emphasis id="FBD9EACBFFC0314EA327B102FF3EF8E3" box="[113,168,1852,1878]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2013: 347, 348, fig. 13. — 
<treatmentCitation id="480C10C8FFC0314EA284B103FF0AF8C0" author="Pasini G. &amp; Garassino A." page="325" pageId="3" pageNumber="30" year="2013">
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, 326, fig. 5. — 
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<bibRefCitation id="AD3C4B28FFC0314EA215B165FD04F8C1" author="Pasini G. &amp; Garassino A." box="[323,658,1882,1908]" pageId="3" pageNumber="30" pagination="343 - 346" refId="ref5426" refString="Pasini G. &amp; Garassino A., 2013 b - New record of crabs (Decapoda, Brachyura) from the early Pliocene of Cassine (Piedmont, northern Italy). Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen, 270 (3): 343 - 346." type="journal article" year="2013">Pasini &amp; Garassino 2013b: 344</bibRefCitation>
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, 345, fig. 2. — 
<treatmentCitation id="480C10C8FFC0314EA3FBB147FE09F826" author="Pasini G. &amp; Garassino A. &amp; Hyzny M. &amp; Baldanza A. &amp; Bizzarri R. &amp; Famiani F." box="[173,415,1912,1939]" page="254" pageId="3" pageNumber="30" year="2014">
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Pasini 
<emphasis id="FBD9EACBFFC0314EA3A3B146FEBCF827" box="[245,298,1912,1938]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2014: 254
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, 255, fig. 9B. — Baldanza 
<emphasis id="FBD9EACBFFC0314EA1E9B146FD62F827" box="[703,756,1912,1938]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2017: 60, 61, fig. 15C. — 
<treatmentCitation id="480C10C8FFC0314EA2DBB1A9FDE5F805" author="Pasini G. &amp; Garassino A. &amp; Sami M." box="[397,627,1942,1968]" httpUri="http://treatment.plazi.org/id/33248510FF98EB0DFFB488C5C267060E" page="31" pageId="3" pageNumber="30" year="2018">
<bibRefCitation id="AD3C4B28FFC0314EA2DBB1A9FDE5F805" author="Pasini G. &amp; Garassino A. &amp; Sami M." box="[397,627,1942,1968]" pageId="3" pageNumber="30" refId="ref5541" refString="Pasini G., Garassino A. &amp; Sami M., 2018 - Decapod crustaceans from the late Pliocene (Placentian) nearby Faenza (Emilia-Romagna, N Italy). Natural History Sciences. Atti della Societa italiana di" type="book" year="2018">
Pasini 
<emphasis id="FBD9EACBFFC0314EA283B1A8FD9AF805" box="[469,524,1942,1968]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
2018: 31
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, fig. 3A.
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<emphasis id="FBD9EACBFFC0314EA033B4DFFB21FD4E" bold="true" box="[869,1207,736,763]" pageId="3" pageNumber="30">Material and measurements</emphasis>
: 13 carapaces in dorsal view (
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2300 – lcxp: 
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); 
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2307 
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lcxp: 
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, wcxp: 
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; 
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1154 
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lcxp: 
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, wcxp: 
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; 
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2309 
<emphasis id="FBD9EACBFFC0314EA79FB504FB41FCE0" bold="true" box="[1225,1239,827,853]" pageId="3" pageNumber="30">–</emphasis>
lcxp: 
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, wcxp: 
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; 
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2315 
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lcxp: 
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, wcxp: 
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2318 – lcxp: 
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, wcxp: 
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; 
<collectionCode id="AFBCAE1CFFC0314EA79AB547FA92FC27" box="[1228,1284,888,914]" country="Switzerland" httpUri="http://grbio.org/cool/7f85-3xtv" name="Sauriermuseum Frick" pageId="3" pageNumber="30">MSF</collectionCode>
2325 – lcxp: 
<quantity id="0E559B3CFFC0314EA6F0B547FCFBFC05" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="30" unit="mm" value="10.0">10 mm</quantity>
, wcxp: 
<quantity id="0E559B3CFFC0314EA09FB5A9FB8BFC05" box="[969,1053,918,944]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="30" unit="mm" value="13.0">13 mm</quantity>
; 
<collectionCode id="AFBCAE1CFFC0314EA77DB5A9FBF5FC05" box="[1067,1123,918,944]" country="Switzerland" httpUri="http://grbio.org/cool/7f85-3xtv" name="Sauriermuseum Frick" pageId="3" pageNumber="30">MSF</collectionCode>
2326 – lcxp: 
<quantity id="0E559B3CFFC0314EA653B5A9FADDFC05" box="[1285,1355,918,944]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="30" unit="mm" value="9.0">9 mm</quantity>
, wcxp: 
<quantity id="0E559B3CFFC0314EA6F0B5A9FCF8FC7B" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="30" unit="mm" value="12.0">12 mm</quantity>
; 
<collectionCode id="AFBCAE1CFFC0314EA02FB58BFC27FC7B" box="[889,945,948,974]" country="Switzerland" httpUri="http://grbio.org/cool/7f85-3xtv" name="Sauriermuseum Frick" pageId="3" pageNumber="30">MSF</collectionCode>
2327 – lcxp: 
<quantity id="0E559B3CFFC0314EA71FB58BFB0FFC7B" box="[1097,1177,948,974]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="30" unit="mm" value="17.0">17 mm</quantity>
, wcxp: 
<quantity id="0E559B3CFFC0314EA7B9B58BFAD7FC7B" box="[1263,1345,948,974]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="30" unit="mm" value="24.0">24 mm</quantity>
; 
<collectionCode id="AFBCAE1CFFC0314EA61DB58BFA15FC7B" box="[1355,1411,948,974]" country="Switzerland" httpUri="http://grbio.org/cool/7f85-3xtv" name="Sauriermuseum Frick" pageId="3" pageNumber="30">MSF</collectionCode>
2328 – lcxp: 
<quantity id="0E559B3CFFC0314EA0C2B5EDFC73FC59" box="[916,997,978,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="30" unit="mm" value="12.0">12 mm</quantity>
, wcxp: 
<quantity id="0E559B3CFFC0314EA76AB5EDFB18FC59" box="[1084,1166,978,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="30" unit="mm" value="14.0">14 mm</quantity>
; 
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2329 – lcxp: 
<quantity id="0E559B3CFFC0314EA63AB5EDFA2BFC59" box="[1388,1469,978,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="30" unit="mm" value="13.0">13 mm</quantity>
, wcxp: 
<quantity id="0E559B3CFFC0314EA0D8B5CFFC72FBBF" box="[910,996,1008,1035]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="30" unit="mm" value="15.0">15 mm</quantity>
; 
<collectionCode id="AFBCAE1CFFC0314EA0A4B5CFFBBCFBBF" box="[1010,1066,1008,1034]" country="Switzerland" httpUri="http://grbio.org/cool/7f85-3xtv" name="Sauriermuseum Frick" pageId="3" pageNumber="30">MSF</collectionCode>
2330 – lcxp: 
<quantity id="0E559B3CFFC0314EA784B5CFFA8EFBBF" box="[1234,1304,1008,1034]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="30" unit="mm" value="7.0">7 mm</quantity>
, wcxp: 
<quantity id="0E559B3CFFC0314EA620B5CFFA28FBBF" box="[1398,1470,1008,1034]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="30" unit="mm" value="9.0">9 mm</quantity>
; 
<collectionCode id="AFBCAE1CFFC0314EA069B231FCE1FB9D" box="[831,887,1038,1064]" country="Switzerland" httpUri="http://grbio.org/cool/7f85-3xtv" name="Sauriermuseum Frick" pageId="3" pageNumber="30">MSF</collectionCode>
2331 – lcxp: 
<quantity id="0E559B3CFFC0314EA747B231FBC2FB9C" box="[1041,1108,1038,1065]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="30" unit="mm" value="6.0">6 mm</quantity>
, wcxp: 
<quantity id="0E559B3CFFC0314EA7FDB231FB66FB9C" box="[1195,1264,1038,1065]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="30" unit="mm" value="8.0">8 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="81B76552FFC03149A033B213FEE4FF75" lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30" type="discussion">
<paragraph id="C91236D9FFC0314EA033B213FB9FFA05" blockId="3.[831,1475,736,1969]" pageId="3" pageNumber="30">
<emphasis id="FBD9EACBFFC0314EA033B213FC75FBF3" bold="true" box="[869,995,1068,1094]" pageId="3" pageNumber="30">Discussion</emphasis>
. The studied specimens have been assigned to 
<taxonomicName id="0EAD4D5AFFC0314EA0DAB275FAC4FBD1" authority="Delle Cave, 1988" authorityName="Delle Cave" authorityYear="1988" box="[908,1362,1098,1124]" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA0DAB275FB1EFBD1" box="[908,1160,1098,1124]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
Delle Cave, 
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</taxonomicName>
in having a slightly convex suboctagonal carapace, wider than long; straight front margin, with a weak median incision; short convex anterolateral margin; long convergent posterolateral margin; posterior margin straight medially; dorsal region well marked by grooves, with wide raised epigastric lobes; well-marked suboval protogastric regions; subpentagonal mesogastric regions with narrow, elongate anterior process between protogastric regions; cardiac region well marked by branchiocardiac grooves; small poorly marked hepatic regions; and wide, well-marked, branchial regions.
</paragraph>
<paragraph id="C91236D9FFC03149A033B38BFEE4FF75" blockId="3.[831,1475,736,1969]" lastBlockId="4.[113,755,135,192]" lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
<taxonomicName id="0EAD4D5AFFC0314EA033B38BFBFEFA7B" authorityName="Delle Cave" authorityYear="1988" box="[869,1128,1460,1486]" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA033B38BFBFEFA7B" box="[869,1128,1460,1486]" italics="true" pageId="3" pageNumber="30">Monodaeus bortolottii</emphasis>
</taxonomicName>
has been previously reported from the Pliocene of Volterra (Delle Cave, 1988) and Grosseto (De Angeli 
<emphasis id="FBD9EACBFFC0314EA767B3CEFBFCF9BF" box="[1073,1130,1520,1546]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
, 2009). Later 
<bibRefCitation id="AD3C4B28FFC0314EA65AB3CFFC5CF99D" author="Pasini G. &amp; Garassino A." pageId="3" pageNumber="30" pagination="319 - 328" refId="ref5378" refString="Pasini G. &amp; Garassino A., 2013 a - Records of brachyuran crabs from the Pliocene (Piacenzian) of Reggio Emilia (Emilia Romagna, N Italy). Boletin de la Sociedad Geologica Mexicana, 65 (2): 319 - 328." type="journal article" year="2013">Pasini &amp; Garassino (2013a</bibRefCitation>
, b) and Garassino, Hyžný &amp; Pasini 
<emphasis id="FBD9EACBFFC0314EA624B030FA1EF99D" box="[1394,1416,1551,1576]" italics="true" pageId="3" pageNumber="30">in</emphasis>
Baldanza 
<emphasis id="FBD9EACBFFC0314EA0DFB012FC57F9F3" box="[905,961,1580,1606]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
(2013) reported this species from the Pliocene (Piacenzian) of Castellarano and Monticelli di Quattro Castella (Reggio Emilia, 
<collectingRegion id="0B69F83BFFC0314EA7D0B056FAD8F931" box="[1158,1358,1641,1668]" country="Italy" name="Emilia-Romagna" pageId="3" pageNumber="30">Emilia-Romagna</collectingRegion>
), from the early Pliocene of Cassine (Alessandria, 
<collectingRegion id="0B69F83BFFC0314EA644B0B8FA13F917" box="[1298,1413,1671,1698]" country="Italy" name="Piemonte" pageId="3" pageNumber="30">Piedmont</collectingRegion>
), and from the early Pleistocene of the Poggio i Sodi (Siena, 
<collectingRegion id="0B69F83BFFC0314EA069B0FBFC32F96B" box="[831,932,1732,1758]" country="Italy" name="Toscana" pageId="3" pageNumber="30">Tuscany</collectingRegion>
). Later 
<bibRefCitation id="AD3C4B28FFC0314EA0AEB0FCFB5BF968" author="Pasini G. &amp; Garassino A. &amp; Hyzny M. &amp; Baldanza A. &amp; Bizzarri R. &amp; Famiani F." box="[1016,1229,1731,1757]" pageId="3" pageNumber="30" refId="ref5477" refString="Pasini G., Garassino A., Hyzny M., Baldanza A., Bizzarri R. &amp; Famiani F., 2014 - The bathyal decapod crustacean community from the early Pleistocene of Volterra (Pisa, Tuscany, central Italy). Neues" type="book" year="2014">
Pasini 
<emphasis id="FBD9EACBFFC0314EA715B0FBFBEDF968" box="[1091,1147,1731,1757]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
(2014)
</bibRefCitation>
reported 
<taxonomicName id="0EAD4D5AFFC0314EA66EB0FBFCC1F94E" authorityName="Delle Cave" authorityYear="1988" class="Malacostraca" family="Xanthidae" genus="Monodaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="bortolottii">
<emphasis id="FBD9EACBFFC0314EA66EB0FBFCC1F94E" italics="true" pageId="3" pageNumber="30">M. bortolottii</emphasis>
</taxonomicName>
from the early Pleistocene of Volterra (Pisa, 
<collectingRegion id="0B69F83BFFC0314EA604B0DDFA20F949" box="[1362,1462,1762,1788]" country="Italy" name="Toscana" pageId="3" pageNumber="30">Tuscany</collectingRegion>
), close to the 
<typeStatus id="1616887BFFC0314EA081B13FFB9EF8AF" box="[983,1032,1792,1818]" pageId="3" pageNumber="30">type</typeStatus>
locality originally reported by Delle Cave (1988). Recently this species has been reported also from the early Pleistocene of Poggi Gialli (Sinalunga, 
<collectingRegion id="0B69F83BFFC0314EA069B165FC37F8C1" box="[831,929,1882,1908]" country="Italy" name="Toscana" pageId="3" pageNumber="30">Tuscany</collectingRegion>
) by De Angeli, Garassino &amp; Pasini 
<emphasis id="FBD9EACBFFC0314EA66CB165FAC6F8C6" box="[1338,1360,1882,1907]" italics="true" pageId="3" pageNumber="30">in</emphasis>
Baldanza 
<emphasis id="FBD9EACBFFC0314EA069B146FCEEF827" box="[831,888,1912,1938]" italics="true" pageId="3" pageNumber="30">et al.</emphasis>
(2017), and from the Pliocene of Faenza (Ravenna, 
<collectingRegion id="0B69F83BFFC0314EA069B1A9FB92F804" box="[831,1028,1942,1969]" country="Italy" name="Emilia-Romagna" pageId="3" pageNumber="30">Emilia-Romagna</collectingRegion>
) (
<bibRefCitation id="AD3C4B28FFC0314EA748B1A9FB6CF805" author="Pasini G. &amp; Garassino A. &amp; Sami M." box="[1054,1274,1942,1968]" pageId="3" pageNumber="30" refId="ref5541" refString="Pasini G., Garassino A. &amp; Sami M., 2018 - Decapod crustaceans from the late Pliocene (Placentian) nearby Faenza (Emilia-Romagna, N Italy). Natural History Sciences. Atti della Societa italiana di" type="book" year="2018">
Pasini 
<emphasis id="FBD9EACBFFC0314EA739B1A8FB24F805" box="[1135,1202,1942,1968]" italics="true" pageId="3" pageNumber="30">et al.,</emphasis>
2018
</bibRefCitation>
). This is the first record of this species in the Miocene, notably enlarging its stratigraphic range.
</paragraph>
</subSubSection>
</treatment>
</document>