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<mods:title id="76FA0BE0D3FA9D7983710F0E3783D860">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). III. Consanguineum, crassum and reflexum groups, and an additional species of the fusculum group</mods:title>
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<mods:namePart id="8189650BD8D237DEE3C57ED6B79E3350">Shavrin, Alexey V.</mods:namePart>
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<treatment id="3706C715FFB9FFE9FF5B60C7FA04FC91" ID-DOI="http://doi.org/10.5281/zenodo.5921725" ID-GBIF-Taxon="159834452" ID-Zenodo-Dep="5921725" LSID="urn:lsid:plazi:treatment:3706C715FFB9FFE9FF5B60C7FA04FC91" httpUri="http://treatment.plazi.org/id/3706C715FFB9FFE9FF5B60C7FA04FC91" lastPageNumber="470" pageId="19" pageNumber="470">
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<paragraph id="BF107603FFB9FFE9FF5B60C7FD13FE3A" blockId="19.[151,650,477,504]" box="[151,650,477,504]" pageId="19" pageNumber="470">
<heading id="E458C16FFFB9FFE9FF5B60C7FD13FE3A" bold="true" box="[151,650,477,504]" fontSize="11" level="1" pageId="19" pageNumber="470" reason="1">
<emphasis id="8DDBAA11FFB9FFE9FF5B60C7FD13FE3A" bold="true" box="[151,650,477,504]" pageId="19" pageNumber="470">
Key to the species of the
<taxonomicName id="78AF0D80FFB9FFE9FE0760C7FDA1FE35" ID-CoL="5VCFR" baseAuthorityName="Reitter" baseAuthorityYear="1891" box="[459,568,477,504]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFB9FFE9FE0760C7FDA1FE35" bold="true" box="[459,568,477,504]" italics="true" pageId="19" pageNumber="470">reflexum</emphasis>
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group
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<subSubSection id="F7B52588FFB9FFE9FF5B6338FA04FC91" pageId="19" pageNumber="470" type="key">
<keyStep id="045B6EA6FFB9FFE9FF5B6338FA05FDBC" pageId="19" pageNumber="470">
<paragraph id="BF107603FFB9FFE9FF5B6338FA05FD98" blockId="19.[151,1437,546,861]" pageId="19" pageNumber="470">
<keyLead id="045ED536FFB9FFE9FF5B6338FA05FD98" pageId="19" pageNumber="470">
1 Lateral edges of pronotum without projections. Body yellow to reddish-brown. Body length:
<quantity id="7857DBE6FFB9FFE9FB976338FB71FDFA" box="[1115,1256,546,567]" metricMagnitude="-3" metricUnit="m" metricValue="3.195" metricValueMax="3.9" metricValueMin="2.49" pageId="19" pageNumber="470" unit="mm" value="3.195" valueMax="3.9" valueMin="2.49">2.49–3.90 mm</quantity>
. Aedeagus narrow, with long parameres (
<figureCitation id="27946A86FFB9FFE9FE7F6325FE62FD98" box="[435,507,575,597]" captionStart="FIGURES 78–81" captionStartId="22.[151,264,1975,2000]" captionTargetBox="[266,1322,176,1949]" captionTargetId="figure@22.[255,1333,166,1956]" captionTargetPageId="22" captionText="FIGURES 78–81. Aedeagus of Anthobium reflexum (78–79: Kazakhstan) and A. kashmiricum (80–81: India, Kashmir): 78, 80—parameral view, 79, 81—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517630" httpUri="https://zenodo.org/record/3517630/files/figure.png" pageId="19" pageNumber="470">Fig. 78</figureCitation>
). Habitus as in
<figureCitation id="27946A86FFB9FFE9FD436325FD4FFD99" box="[655,726,575,597]" captionStart="FIGURES 73–77" captionStartId="20.[151,264,1834,1859]" captionTargetBox="[165,1424,451,1805]" captionTargetId="figure@20.[157,1431,445,1815]" captionTargetPageId="20" captionText="FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517628" httpUri="https://zenodo.org/record/3517628/files/figure.png" pageId="19" pageNumber="470">Fig. 73</figureCitation>
. Central Asia...........................
<taxonomicName id="78AF0D80FFB9FFE9FB506325FA05FD98" ID-CoL="5VCFR" authority="(Reitter, 1891)" baseAuthorityName="Reitter" baseAuthorityYear="1891" box="[1180,1436,575,597]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="reflexum">
<emphasis id="8DDBAA11FFB9FFE9FB506325FA93FD99" box="[1180,1290,575,596]" italics="true" pageId="19" pageNumber="470">A. reflexum</emphasis>
(
<bibRefCitation id="DB3E0BF2FFB9FFE9FADA6325FA0CFD98" author="Reitter, E." box="[1302,1429,575,597]" pageId="19" pageNumber="470" pagination="195 - 199" refId="ref19463" refString="Reitter, E. (1891) Zweiter Beitrag zur Coleopteren-Fauna des russischen Reiches. Wiener Entomologische Zeitung, 10, 195 - 199." type="journal article" year="1891">Reitter, 1891</bibRefCitation>
)
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</paragraph>
<paragraph id="BF107603FFB9FFE9FF5B6347FA05FDBC" blockId="19.[151,1437,546,861]" box="[151,1436,604,626]" pageId="19" pageNumber="470">
<keyLead id="045ED536FFB9FFE9FF5B6347FA05FDBC" box="[151,1436,604,626]" pageId="19" pageNumber="470">- Lateral edges of pronotum with indistinct projections. Aedeagus wider, parameres shorter............................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="045B6EA6FFB9FFE9FF5B6360FA05FD2A" pageId="19" pageNumber="470">
<paragraph id="BF107603FFB9FFE9FF5B6360FA04FD61" blockId="19.[151,1437,546,861]" pageId="19" pageNumber="470">
<keyLead id="045ED536FFB9FFE9FF5B6360FA04FD61" pageId="19" pageNumber="470">
2 Elytra less than twice as long as pronotum. Body yellow-brown. Smaller:
<quantity id="7857DBE6FFB9FFE9FC786363FBDAFD42" box="[948,1091,633,655]" metricMagnitude="-3" metricUnit="m" metricValue="3.4699999999999998" metricValueMax="3.54" metricValueMin="3.4" pageId="19" pageNumber="470" unit="mm" value="3.4699999999999998" valueMax="3.54" valueMin="3.4">3.40–3.54 mm</quantity>
. Aedeagus as in
<figureCitation id="27946A86FFB9FFE9FB2B6360FAB7FD42" box="[1255,1326,633,656]" captionStart="FIGURES 83–86" captionStartId="28.[151,264,1973,1998]" captionTargetBox="[288,1302,176,1938]" captionTargetId="figure@28.[273,1314,166,1954]" captionTargetPageId="28" captionText="FIGURES 83–86. Aedeagus of Anthobium deplanatum (83–84: paratype) and A. gaoligongshanense (85–86: holotype): 83, 85—parameral view, 84, 86—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517634" httpUri="https://zenodo.org/record/3517634/files/figure.png" pageId="19" pageNumber="470">Fig. 85</figureCitation>
; parameres slightly asymmetrical. Habitus as in
<figureCitation id="27946A86FFB9FFE9FD8D638DFD11FD60" box="[577,648,663,685]" captionStart="FIGURES 73–77" captionStartId="20.[151,264,1834,1859]" captionTargetBox="[165,1424,451,1805]" captionTargetId="figure@20.[157,1431,445,1815]" captionTargetPageId="20" captionText="FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517628" httpUri="https://zenodo.org/record/3517628/files/figure.png" pageId="19" pageNumber="470">Fig. 75</figureCitation>
.
<collectingCountry id="C7B83693FFB9FFE9FD5D638DFD57FD60" box="[657,718,663,685]" name="China" pageId="19" pageNumber="470">China</collectingCountry>
:
<collectingRegion id="7D6BB8E1FFB9FFE9FD1A638DFC86FD61" box="[726,799,663,684]" country="China" name="Yunnan" pageId="19" pageNumber="470">Yunnan</collectingRegion>
...............................
<taxonomicName id="78AF0D80FFB9FFE9FB5E638DFAFFFD61" ID-CoL="8XTT5" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[1170,1382,663,684]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="gaoligongshanense" status="sp. nov.">
<emphasis id="8DDBAA11FFB9FFE9FB5E638DFAFFFD61" box="[1170,1382,663,684]" italics="true" pageId="19" pageNumber="470">A. gaoligongshanense</emphasis>
</taxonomicName>
<emphasis id="8DDBAA11FFB9FFE9FAA0638DFA04FD61" bold="true" box="[1388,1437,663,684]" pageId="19" pageNumber="470">
<taxonomicNameLabel id="96E8176AFFB9FFE9FAA0638DFA04FD61" box="[1388,1437,663,684]" pageId="19" pageNumber="470" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="BF107603FFB9FFE9FF5B63AFFA05FD2A" blockId="19.[151,1437,546,861]" pageId="19" pageNumber="470">
<keyLead id="045ED536FFB9FFE9FF5B63AFFA05FD2A" pageId="19" pageNumber="470">
- Elytra markedly more than twice as long as pronotum. Body yellow-brown to reddish-brown, with paler lateral portions of pronotum. Body larger. Aedeagus different; parameres symmetrical (
<figureCitation id="27946A86FFB9FFE9FCBA63C8FC5FFD2A" box="[886,966,722,744]" captionStart="FIGURES 78–81" captionStartId="22.[151,264,1975,2000]" captionTargetBox="[266,1322,176,1949]" captionTargetId="figure@22.[255,1333,166,1956]" captionTargetPageId="22" captionText="FIGURES 78–81. Aedeagus of Anthobium reflexum (78–79: Kazakhstan) and A. kashmiricum (80–81: India, Kashmir): 78, 80—parameral view, 79, 81—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517630" httpUri="https://zenodo.org/record/3517630/files/figure.png" pageId="19" pageNumber="470">Figs. 80</figureCitation>
,
<figureCitation id="27946A86FFB9FFE9FC1C63C8FC70FD25" box="[976,1001,722,744]" captionStart="FIGURES 83–86" captionStartId="28.[151,264,1973,1998]" captionTargetBox="[288,1302,176,1938]" captionTargetId="figure@28.[273,1314,166,1954]" captionTargetPageId="28" captionText="FIGURES 83–86. Aedeagus of Anthobium deplanatum (83–84: paratype) and A. gaoligongshanense (85–86: holotype): 83, 85—parameral view, 84, 86—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517634" httpUri="https://zenodo.org/record/3517634/files/figure.png" pageId="19" pageNumber="470">83</figureCitation>
).................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="045B6EA6FFB9FFE9FF5B63F5FA04FC91" pageId="19" pageNumber="470">
<paragraph id="BF107603FFB9FFE9FF5B63F5FA05FCEF" blockId="19.[151,1437,546,861]" pageId="19" pageNumber="470">
<keyLead id="045ED536FFB9FFE9FF5B63F5FA05FCEF" pageId="19" pageNumber="470">
3 Body paler, yellow-brown to brown. Body length:
<quantity id="7857DBE6FFB9FFE9FD0463F5FCCFFCC9" box="[712,854,751,773]" metricMagnitude="-3" metricUnit="m" metricValue="4.075" metricValueMax="4.45" metricValueMin="3.7" pageId="19" pageNumber="470" unit="mm" value="4.075" valueMax="4.45" valueMin="3.7">3.70–4.45 mm</quantity>
. Apical portion of parameres wider; internal sac simple (
<figureCitation id="27946A86FFB9FFE9FAB963F5FF64FCEF" captionStart="FIGURES 78–81" captionStartId="22.[151,264,1975,2000]" captionTargetBox="[266,1322,176,1949]" captionTargetId="figure@22.[255,1333,166,1956]" captionTargetPageId="22" captionText="FIGURES 78–81. Aedeagus of Anthobium reflexum (78–79: Kazakhstan) and A. kashmiricum (80–81: India, Kashmir): 78, 80—parameral view, 79, 81—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517630" httpUri="https://zenodo.org/record/3517630/files/figure.png" pageId="19" pageNumber="470">Fig. 80</figureCitation>
). Habitus as in
<figureCitation id="27946A86FFB9FFE9FE5D6216FE41FCEF" box="[401,472,780,802]" captionStart="FIGURES 73–77" captionStartId="20.[151,264,1834,1859]" captionTargetBox="[165,1424,451,1805]" captionTargetId="figure@20.[157,1431,445,1815]" captionTargetPageId="20" captionText="FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517628" httpUri="https://zenodo.org/record/3517628/files/figure.png" pageId="19" pageNumber="470">Fig. 76</figureCitation>
.
<collectingCountry id="C7B83693FFB9FFE9FE2D6216FD8FFCEF" box="[481,534,780,802]" name="India" pageId="19" pageNumber="470">India</collectingCountry>
: Kashmir.........................................
<taxonomicName id="78AF0D80FFB9FFE9FBAE6216FA05FCEF" ID-CoL="67CDF" authority="(Cameron, 1941)" baseAuthorityName="Cameron" baseAuthorityYear="1941" box="[1122,1436,780,802]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="kashmiricum">
<emphasis id="8DDBAA11FFB9FFE9FBAE6216FB6BFCEC" box="[1122,1266,780,801]" italics="true" pageId="19" pageNumber="470">A.kashmiricum</emphasis>
(
<bibRefCitation id="DB3E0BF2FFB9FFE9FB336216FA0CFCEF" author="Cameron, M." box="[1279,1429,780,802]" pageId="19" pageNumber="470" pagination="56 - 60" refId="ref19201" refString="Cameron, M. (1941) Descriptions of new Staphylinidae (Coleopt.). The Proceedings of the Royal Entomological Society of London (B), 10, 56 - 60. https: // doi. org / 10.1111 / j. 1365 - 3113.1941. tb 00693. x" type="journal article" year="1941">Cameron, 1941</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="BF107603FFB9FFE9FF5B6230FA04FC91" blockId="19.[151,1437,546,861]" pageId="19" pageNumber="470">
<keyLead id="045ED536FFB9FFE9FF5B6230FA04FC91" pageId="19" pageNumber="470">
- Body darker, reddish-brown. Body length:
<quantity id="7857DBE6FFB9FFE9FDB66233FC91FCF2" box="[634,776,809,831]" metricMagnitude="-3" metricUnit="m" metricValue="4.0649999999999995" metricValueMax="4.5" metricValueMin="3.63" pageId="19" pageNumber="470" unit="mm" value="4.0649999999999995" valueMax="4.5" valueMin="3.63">3.63–4.50 mm</quantity>
. Apical portion of parameres narrower; internal sac complicated (
<figureCitation id="27946A86FFB9FFE9FAB96230FF64FC90" captionStart="FIGURES 83–86" captionStartId="28.[151,264,1973,1998]" captionTargetBox="[288,1302,176,1938]" captionTargetId="figure@28.[273,1314,166,1954]" captionTargetPageId="28" captionText="FIGURES 83–86. Aedeagus of Anthobium deplanatum (83–84: paratype) and A. gaoligongshanense (85–86: holotype): 83, 85—parameral view, 84, 86—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3517634" httpUri="https://zenodo.org/record/3517634/files/figure.png" pageId="19" pageNumber="470">Fig. 83</figureCitation>
). Habitus as in
<figureCitation id="27946A86FFB9FFE9FE5D625DFE41FC91" box="[401,472,839,861]" captionStart="FIGURES 73–77" captionStartId="20.[151,264,1834,1859]" captionTargetBox="[165,1424,451,1805]" captionTargetId="figure@20.[157,1431,445,1815]" captionTargetPageId="20" captionText="FIGURES 73–77. Habitus of Anthobium: 73—A. reflexum (Kazakhstan) 74—A. deplanatum (holotype), 75—A. gaoligongshanense (holotype), 76—A. kashmiricum, 77—A. transversale (holotype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3517628" httpUri="https://zenodo.org/record/3517628/files/figure.png" pageId="19" pageNumber="470">Fig. 74</figureCitation>
.
<collectingCountry id="C7B83693FFB9FFE9FE2D625DFD87FC90" box="[481,542,839,861]" name="China" pageId="19" pageNumber="470">China</collectingCountry>
:
<collectingRegion id="7D6BB8E1FFB9FFE9FDEB625DFDEDFC90" box="[551,628,839,861]" country="China" name="Sichuan" pageId="19" pageNumber="470">Sichuan</collectingRegion>
...................................................
<taxonomicName id="78AF0D80FFB9FFE9FB16625DFAFFFC91" ID-CoL="8XTSY" authority="Shavrin &amp; Smetana, 2019" authorityName="Shavrin &amp; Smetana" authorityYear="2019" box="[1242,1382,839,860]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="deplanatum" status="sp. nov.">
<emphasis id="8DDBAA11FFB9FFE9FB16625DFAFFFC91" box="[1242,1382,839,860]" italics="true" pageId="19" pageNumber="470">A. deplanatum</emphasis>
</taxonomicName>
<emphasis id="8DDBAA11FFB9FFE9FAA0625DFA04FC91" bold="true" box="[1388,1437,839,860]" pageId="19" pageNumber="470">
<taxonomicNameLabel id="96E8176AFFB9FFE9FAA0625DFA04FC91" box="[1388,1437,839,860]" pageId="19" pageNumber="470" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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