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<document id="0D0F2498524A5767DED8256D3D2778A9" ID-DOI="10.5281/zenodo.157462" ID-GBIF-Dataset="a9ccfc65-1459-414b-8a99-c99649f83d51" ID-ISSN="1175­5326" ID-Zenodo-Dep="157462" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461019330970" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2004" docId="160487E5FA6DFFBDFE95FEDEFC3DFB39" docLanguage="en" docName="zt00538.pdf" docOrigin="Zootaxa 538" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pneuminion nanum Perkins, 2004, new species" docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="EA3DFF9DFA66FFB1FF9DF83AFFBAFFCF" masterDocTitle="A revision of the South African endemic water beetle genus Pneuminion Perkins (Coleoptera: Hydraenidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="12" updateTime="1698569824382" updateUser="plazi">
<mods:mods id="1FF3240E15C36D49C4A50DC8ED055856" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="473D50C1DB5A91A54E9651599A1A250D">
<mods:title id="9DF4F807F91CE7E0962E578E6CE18F8B">A revision of the South African endemic water beetle genus Pneuminion Perkins (Coleoptera: Hydraenidae)</mods:title>
</mods:titleInfo>
<mods:name id="BDD5B9EDFE18D7BD35F6FDAF844ADB5E" type="personal">
<mods:role id="5DDF7FCB092D5E1551F0A60DC91DFD0B">
<mods:roleTerm id="3A7DEDA5A556703A4FF37EB83E691347">Author</mods:roleTerm>
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<mods:namePart id="0BBD2B74DEEB78F1A10A4D1000287F63">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="B140A5B17A962240540AB19D6A67912E">text</mods:typeOfResource>
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<mods:title id="C19BDE47B5CDA074EE8DEB0C87C54A59">Zootaxa</mods:title>
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<mods:part id="9EB8CF1C2CAEB670C9B0230C69A7275F">
<mods:date id="31052AECAE4858EB7A5DA3B84FBAABDF">2004</mods:date>
<mods:detail id="086DF65ED8EFB900970504CB2DB4CB1F" type="volume">
<mods:number id="8E10DF42EF0A3ABB943B7466A44DEB37">538</mods:number>
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<mods:classification id="87BBDE4E33EE848A37D5D5F5FBAD285A">journal article</mods:classification>
<mods:identifier id="611B88541DB8EB9D4F38A371C54C5CA9" type="DOI">10.5281/zenodo.157462</mods:identifier>
<mods:identifier id="7396F7E22F2F9B19C48282DCC2374B22" type="GBIF-Dataset">a9ccfc65-1459-414b-8a99-c99649f83d51</mods:identifier>
<mods:identifier id="3125359BB99346A581B6060FF9D49949" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="880262DC5980FD534AA361096E7D26FB" type="Zenodo-Dep">157462</mods:identifier>
<mods:identifier id="783EE6F847DB1EBB86FD9C06E8E5A9B1" type="ZooBank">3A93701A-E53F-4A89-B4C6-1F952ECC82BD</mods:identifier>
</mods:mods>
<treatment id="160487E5FA6DFFBDFE95FEDEFC3DFB39" ID-DOI="http://doi.org/10.5281/zenodo.6271133" ID-GBIF-Taxon="119602184" ID-Zenodo-Dep="6271133" LSID="urn:lsid:plazi:treatment:160487E5FA6DFFBDFE95FEDEFC3DFB39" httpUri="http://treatment.plazi.org/id/160487E5FA6DFFBDFE95FEDEFC3DFB39" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="D6B76578FA6DFFBAFE95FEDEFE4BF8EE" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="9E1236F3FA6DFFBAFE95FEDEFD29F931" blockId="11.[264,659,1764,1825]" box="[264,659,1764,1790]" pageId="11" pageNumber="12">
<heading id="C55A819FFA6DFFBAFE95FEDEFD29F931" bold="true" box="[264,659,1764,1790]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="ACD9EAE1FA6DFFBAFE95FEDEFD29F931" bold="true" box="[264,659,1764,1790]" pageId="11" pageNumber="12">
<taxonomicName id="59AD4D70FA6DFFBAFE95FEDEFE41F931" ID-CoL="783DH" box="[264,507,1764,1790]" class="Insecta" family="Hydraenidae" genus="Pneuminion" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nanum" status="sp. nov.">
<emphasis id="ACD9EAE1FA6DFFBAFE95FEDEFE41F931" bold="true" box="[264,507,1764,1790]" italics="true" pageId="11" pageNumber="12">Pneuminion nanum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7EA579AFA6DFFBAFD9EFEDEFD29F931" box="[515,659,1764,1790]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9E1236F3FA6DFFBAFE95FF3DFE4BF8EE" blockId="11.[264,659,1764,1825]" box="[264,497,1799,1825]" pageId="11" pageNumber="12">
(
<figureCitation id="06962A76FA6DFFBAFE8DFF3DFE2BF8EE" box="[272,401,1799,1825]" captionStart="FIGURE 10" captionStartId="23.[264,368,1558,1582]" captionTargetBox="[284,1309,329,1549]" captionTargetId="figure@23.[264,1323,309,1553]" captionTargetPageId="23" captionText="FIGURE 10. Pneuminion nanum holotype. Dorsal and lateral habitus." httpUri="https://zenodo.org/record/169056/files/figure.png" pageId="11" pageNumber="12">Figures 10</figureCitation>
,
<figureCitation id="06962A76FA6DFFBAFE00FF3DFE06F8EE" box="[413,444,1799,1825]" captionStart="FIGURE 19" captionStartId="32.[264,368,1711,1735]" captionTargetBox="[383,1194,354,1670]" captionTargetId="figure@32.[364,1222,326,1687]" captionTargetPageId="32" captionText="FIGURE 19. Pneuminion nanum. Aedeagus of holotype." httpUri="https://zenodo.org/record/157477/files/figure.png" pageId="11" pageNumber="12">19</figureCitation>
,
<figureCitation id="06962A76FA6DFFBAFE54FF3DFE52F8EE" box="[457,488,1799,1825]" captionStart="FIGURES 26 – 28" captionStartId="36.[264,383,1835,1859]" captionTargetBox="[275,1306,310,1827]" captionTargetId="figure@36.[275,1307,308,1830]" captionTargetPageId="36" captionText="FIGURES 26 – 28. Pneuminion distribution maps. — 26. P. balfourbrownei. — 27. P. impressum. — 28. P. nanum." httpUri="https://zenodo.org/record/169060/files/figure.png" pageId="11" pageNumber="12">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6DFFBDFE95FF6CFC4CFE91" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="9E1236F3FA6DFFBDFE95FF6CFC4CFE91" blockId="11.[264,1323,1878,1905]" lastBlockId="12.[264,1325,284,1270]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="ACD9EAE1FA6DFFBAFE95FF6CFE04F8BF" bold="true" box="[264,446,1878,1904]" pageId="11" pageNumber="12">
<typeStatus id="41168851FA6DFFBAFE95FF6CFEF9F8BF" box="[264,323,1878,1904]" pageId="11" pageNumber="12">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="41168851FA6DFFBAFE53FF6DFD81F8BE" box="[462,571,1879,1905]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="E6BA7663FA6DFFBAFD00FF6DFC8DF8BE" box="[669,823,1879,1905]" name="South Africa" pageId="11" pageNumber="12">South Africa</collectingCountry>
: Cape Mt., Helderfont. S,
<quantity id="59559B16FA6DFFBAFBF5FF6DFB06F8BE" box="[1128,1212,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="11" pageNumber="12" unit="m" value="1220.0">1220m</quantity>
,
<geoCoordinate id="FB995034FA6DFFBAFB55FF6DFAA0F8BE" box="[1224,1306,1879,1905]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="555" value="-33.56">33.56S</geoCoordinate>
­
<geoCoordinate id="FB995034FA6AFFBDFE95F926FEE4FEF9" box="[264,350,284,310]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="555" value="20.52">20.52E</geoCoordinate>
, sifted, alpine marsh,
<date id="EA131033FA6AFFBDFDE2F926FCBBFEF9" box="[639,769,284,310]" pageId="12" pageNumber="13" value="1979-03-10">10.iii.1979</date>
, Endrödy­Younga (#1566); deposited in the
<collectionCode id="F8BCAE36FA6AFFBDFE95F97EFEE7FE91" box="[264,349,324,350]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="12" pageNumber="13">TMSA</collectionCode>
.
<typeStatus id="41168851FA6AFFBDFEEDF97EFE5DFE91" box="[368,487,324,350]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
: Same data as
<typeStatus id="41168851FA6AFFBDFD04F97EFCB9FE91" box="[665,771,324,350]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
, (3
<collectionCode id="F8BCAE36FA6AFFBDFCB3F97EFC39FE91" box="[814,899,324,350]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="12" pageNumber="13">TMSA</collectionCode>
, 1
<collectionCode id="F8BCAE36FA6AFFBDFC38F97EFC50FE91" LSID="urn:lsid:biocol.org:col:33791" box="[933,1002,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="12" pageNumber="13">MCZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6AFFBDFEA5F951FC90FE31" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="9E1236F3FA6AFFBDFEA5F951FC90FE31" blockId="12.[264,1325,284,1270]" pageId="12" pageNumber="13">
<emphasis id="ACD9EAE1FA6AFFBDFEA5F951FE02FE4A" bold="true" box="[312,440,363,389]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
Recognized by the small size, ca.
<quantity id="59559B16FA6AFFBDFCC0F956FC7DFE49" box="[861,967,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="13" unit="mm" value="1.3">1.30 mm</quantity>
long (smallest known species in the genus), the subsulcate elytra with whitish setae forming uninterrupted unilinear row on intervals and interrupted unilinear row on series, and the small heart­shaped punctures on the frons disc and pronotal reliefs (
<figureCitation id="06962A76FA6AFFBDFD5BF9DEFCA7FE31" box="[710,797,484,510]" captionStart="FIGURE 10" captionStartId="23.[264,368,1558,1582]" captionTargetBox="[284,1309,329,1549]" captionTargetId="figure@23.[264,1323,309,1553]" captionTargetPageId="23" captionText="FIGURE 10. Pneuminion nanum holotype. Dorsal and lateral habitus." httpUri="https://zenodo.org/record/169056/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6AFFBDFEA5FA31FECFFB01" pageId="12" pageNumber="13" type="description">
<paragraph id="9E1236F3FA6AFFBDFEA5FA31FD42FD81" blockId="12.[264,1325,284,1270]" pageId="12" pageNumber="13">
<emphasis id="ACD9EAE1FA6AFFBDFEA5FA31FE6AFDEA" bold="true" box="[312,464,523,549]" pageId="12" pageNumber="13">Description.</emphasis>
Size (mm x 100; length/width): body (length to elytral apices) 130/57, head 22/39, pronotum 33/52, elytra 81/57.
</paragraph>
<paragraph id="9E1236F3FA6AFFBDFEA5FA66FC19FD21" blockId="12.[264,1325,284,1270]" pageId="12" pageNumber="13">
Dorsum dark brown to black, legs and maxillary palpi brown; dorsal setae moderately conspicuous, whitish. Labrum and clypeus dull, micropunctulate. Frons disc punctate, interpunctal surface smooth, shining, punctures separated by ca. 1.5xpd. Fovea in front of each ocellus large, ca.
<date id="EA131033FA6AFFBDFD8CFAEEFD94FD21" box="[529,558,724,750]" pageId="12" pageNumber="13">5x</date>
ocellus, deep, micropunctulate.
</paragraph>
<paragraph id="9E1236F3FA6AFFBDFEA5FAC6FB2AFC79" blockId="12.[264,1325,284,1270]" pageId="12" pageNumber="13">Pronotum weakly shining on disc, dull laterally, impressions deep; punctures on reliefs small, separated by ca. 1xpd, each with short whitish, granule­based seta; oblique anterior fovea on each side shallowly connecting to midlongitudinal fovea; posterior U­shaped fovea well­developed; setae at lateral margin flat on cuticle in dry specimens. Pronotal postocular emarginations deeper and less symmetrical in females than males.</paragraph>
<paragraph id="9E1236F3FA6AFFBDFEA5FBFEFBDDFB99" blockId="12.[264,1325,284,1270]" pageId="12" pageNumber="13">
Elytra subsulcate, intervals slightly raised, each with a unilinear row of slightly overlapping setae; serial punctures moderately large, 2–
<date id="EA131033FA6AFFBDFCF4FBD6FCCCFBC9" box="[873,886,1004,1030]" pageId="12" pageNumber="13">3</date>
x than those of pronotal reliefs, each with a short seta, setae not overlapping and not forming uninterrupted line like setae of intervals; each seta granule­based; explanate margin narrow in both sexes
</paragraph>
<paragraph id="9E1236F3FA6AFFBDFEA5FC5EFECFFB01" blockId="12.[264,1325,284,1270]" pageId="12" pageNumber="13">
Aedeagus: main­piece length 35, total length 70; duct very long, very slender over distal 3/4; paramere setae comparatively sparse; main­piece with ridge/groove to support duct (
<figureCitation id="06962A76FA6AFFBDFE8CFC8EFEDDFB01" box="[273,359,1204,1230]" captionStart="FIGURE 19" captionStartId="32.[264,368,1711,1735]" captionTargetBox="[383,1194,354,1670]" captionTargetId="figure@32.[364,1222,326,1687]" captionTargetPageId="32" captionText="FIGURE 19. Pneuminion nanum. Aedeagus of holotype." httpUri="https://zenodo.org/record/157477/files/figure.png" pageId="12" pageNumber="13">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D6B76578FA6AFFBDFEA5FCE6FC3DFB39" box="[312,903,1244,1270]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="9E1236F3FA6AFFBDFEA5FCE6FC3DFB39" blockId="12.[264,1325,284,1270]" box="[312,903,1244,1270]" pageId="12" pageNumber="13">
<emphasis id="ACD9EAE1FA6AFFBDFEA5FCE6FE7EFB39" bold="true" box="[312,452,1244,1270]" pageId="12" pageNumber="13">Etymology.</emphasis>
Named in reference to the small size.
</paragraph>
</subSubSection>
</treatment>
</document>
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