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<document id="0046D4867C93BFA16F718E994CA9BB6D" ID-CLB-Dataset="29986" ID-DOI="10.11646/zootaxa.4425.3.7" ID-GBIF-Dataset="4e5a28bc-9d7d-4b3f-b526-2fa812aae747" ID-ISSN="1175-5326" ID-Zenodo-Dep="1270435" ID-ZooBank="DE27A002-C0D2-4F96-851B-D0C1C51554B5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528144441950" checkinUser="plazi" docAuthor="Enghoff, Henrik &amp; Larsson, Tobias Berglund" docDate="2018" docId="03EC0551FFF9FFD186E264BC8667B3F6" docLanguage="en" docName="zootaxa.4425.3.7.pdf.imf" docOrigin="Zootaxa 4425 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudotibiozus Demange 1970" docType="treatment" docVersion="7" lastPageNumber="543" masterDocId="FFD57D29FFF8FFD3867560238326B072" masterDocTitle="Pseudotibiozus Demange, 1970 - millipedes of the Eastern Arc Mountains, Tanzania (Diplopoda, Spirostreptida, Spirostreptidae)" masterLastPageNumber="554" masterPageNumber="541" pageNumber="542" updateTime="1698546649654" updateUser="ExternalLinkService">
<mods:mods id="EEC79DC5EFEECE38FACA671EEBA09571" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0F377B13040FB652D9ABBB74EFFB7D2E">Pseudotibiozus Demange, 1970 - millipedes of the Eastern Arc Mountains, Tanzania (Diplopoda, Spirostreptida, Spirostreptidae)</mods:title>
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<mods:namePart id="3DBAF22BA55ED9EE04493A632E8E7BD2">Enghoff, Henrik</mods:namePart>
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<mods:name id="ADB8C9F49EF866A464B6EA56E11FCFF8" type="personal">
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<mods:namePart id="93734B43F9A7FFE72C79E592C51672FE">Larsson, Tobias Berglund</mods:namePart>
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<mods:typeOfResource id="91B77BA7E476CA79692AB93ACD9B289C">text</mods:typeOfResource>
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<mods:title id="C60675661A25B3C32EE16028CF18F6D1">Zootaxa</mods:title>
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<mods:date id="2CAA02797204DDB43F3116563BC83AF8">2018</mods:date>
<mods:detail id="7146DAD0505A6FA4AF076D2AB26EFDEC" type="volume">
<mods:number id="55FBD318B748EF79A189A06DCA0296D6">4425</mods:number>
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<mods:classification id="6B94D2EFB3609A428755B3F3315C4FD8">journal article</mods:classification>
<mods:identifier id="F3283235F9BAC778B318E3100FA71A51" type="CLB-Dataset">29986</mods:identifier>
<mods:identifier id="A5938DBFAABDE6C561644CE6F8F501A3" type="DOI">10.11646/zootaxa.4425.3.7</mods:identifier>
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<treatment id="03EC0551FFF9FFD186E264BC8667B3F6" ID-DOI="http://doi.org/10.5281/zenodo.5973512" ID-GBIF-Taxon="144235635" ID-Zenodo-Dep="5973512" LSID="urn:lsid:plazi:treatment:03EC0551FFF9FFD186E264BC8667B3F6" httpUri="http://treatment.plazi.org/id/03EC0551FFF9FFD186E264BC8667B3F6" lastPageId="2" lastPageNumber="543" pageId="1" pageNumber="542">
<subSubSection id="C35FE7CCFFF9FFD286E264BC8134B4C8" box="[151,530,1183,1210]" pageId="1" pageNumber="542" type="nomenclature">
<paragraph id="8BFAB447FFF9FFD286E264BC8134B4C8" blockId="1.[151,530,1183,1210]" box="[151,530,1183,1210]" pageId="1" pageNumber="542">
<heading id="D0B2032BFFF9FFD286E264BC8134B4C8" bold="true" box="[151,530,1183,1210]" fontSize="11" level="1" pageId="1" pageNumber="542" reason="1">
<taxonomicName id="4C45CFC4FFF9FFD286E264BC8134B4C8" ID-CoL="7PWS8" authority="Demange, 1970" authorityName="Demange" authorityYear="1970" box="[151,530,1183,1210]" class="Diplopoda" family="Spirostreptidae" genus="Pseudotibiozus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="B9316855FFF9FFD286E264BC8134B4C8" bold="true" box="[151,530,1183,1210]" pageId="1" pageNumber="542">
<emphasis id="B9316855FFF9FFD286E264BC826AB4CB" bold="true" box="[151,332,1183,1209]" italics="true" pageId="1" pageNumber="542">Pseudotibiozus</emphasis>
<bibRefCitation id="EFD4C9B6FFF9FFD2872664838134B4C8" box="[339,530,1184,1210]" pageId="1" pageNumber="542" refId="ref4935" refString="Demange, J. - M. (1970) Elements d'une revision des Spirostreptidae. I. Etude de quelques caracteres taxonomiques des Spirostreptinae. Bulletin d l'Institut fondamental de l'Afrique noire, 32 A (2), 366 - 411." type="journal article">Demange, 1970</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35FE7CCFFF9FFD286E264C580A3B551" pageId="1" pageNumber="542" type="materials_examined">
<paragraph id="8BFAB447FFF9FFD286E264C580C7B48D" blockId="1.[151,1437,1254,2035]" box="[151,993,1254,1279]" pageId="1" pageNumber="542">
<emphasis id="B9316855FFF9FFD286E264C5820DB48D" bold="true" box="[151,299,1254,1279]" pageId="1" pageNumber="542">Type species</emphasis>
.
<taxonomicName id="4C45CFC4FFF9FFD2874C64C581EAB48D" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[313,716,1254,1279]" class="Diplopoda" family="Spirostreptidae" genus="Mardonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="cerasopus">
<emphasis id="B9316855FFF9FFD2874C64C5810AB48C" box="[313,556,1254,1279]" italics="true" pageId="1" pageNumber="542">Mardonius cerasopus</emphasis>
<bibRefCitation id="EFD4C9B6FFF9FFD2844764C481EAB48D" box="[562,716,1255,1279]" pageId="1" pageNumber="542" refId="ref4634" refString="Attems, C. (1914) Afrikanische Spirostreptiden, nebst Uberblick uber die Spirostreptiden orbis terrarum. Zoologia, Stuttgart, 25 (65 / 66), 1 - 233." type="journal article">Attems, 1914</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8BFAB447FFF9FFD286B2652980A3B551" blockId="1.[151,1437,1254,2035]" box="[199,901,1290,1315]" pageId="1" pageNumber="542">
<emphasis id="B9316855FFF9FFD286B2652982FBB551" bold="true" box="[199,477,1290,1315]" pageId="1" pageNumber="542">Other included species.</emphasis>
<taxonomicName id="4C45CFC4FFF9FFD2879165298003B551" authority="Enghoff &amp; Larsson, 2018" authorityName="Enghoff &amp; Larsson" authorityYear="2018" box="[484,805,1290,1315]" class="Diplopoda" family="Spirostreptidae" genus="Pseudotibiozus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="zophoribates" status="sp. nov.">
<emphasis id="B9316855FFF9FFD2879165298003B551" box="[484,805,1290,1315]" italics="true" pageId="1" pageNumber="542">Pseudotibiozus zophoribates</emphasis>
</taxonomicName>
<emphasis id="B9316855FFF9FFD28559652980A3B551" bold="true" box="[812,901,1290,1315]" pageId="1" pageNumber="542">
<taxonomicNameLabel id="A202D52EFFF9FFD28559652980A3B551" box="[812,901,1290,1315]" pageId="1" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35FE7CCFFF9FFD286B2650D81ABB66D" pageId="1" pageNumber="542" type="diagnosis">
<paragraph id="8BFAB447FFF9FFD286B2650D8096B589" blockId="1.[151,1437,1254,2035]" pageId="1" pageNumber="542">
<emphasis id="B9316855FFF9FFD286B2650D8267B535" bold="true" box="[199,321,1326,1351]" pageId="1" pageNumber="542">Diagnosis.</emphasis>
A genus of
<taxonomicName id="4C45CFC4FFF9FFD287A1650D8158B535" authorityName="Brandt" authorityYear="1833" box="[468,638,1326,1351]" class="Diplopoda" family="Spirostreptidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="542" phylum="Arthropoda" rank="family">Spirostreptidae</taxonomicName>
characterized by the following combination of gonopodal characters: Apical lateral process of metaplica short, rounded. Antetorsal process of telopodite very short, branching off from main telopodite stem before the telopodite curves laterad and exits the coxa. Telopodite becoming thicker shortly after exiting coxa and flattening to an almost ribbonlike structure, telopodite bending mesad at mid-length and dividing into a spinelike branch and a thinner solenomere, solenomere twisting up and around the spinelike branch in an overall mesad direction. Tip of solenomere wrinkled, spongelike.
</paragraph>
<paragraph id="8BFAB447FFF9FFD286B2662581ABB66D" blockId="1.[151,1437,1254,2035]" box="[199,653,1542,1567]" pageId="1" pageNumber="542">
<emphasis id="B9316855FFF9FFD286B2662582F4B66D" bold="true" box="[199,466,1542,1567]" pageId="1" pageNumber="542">General description of</emphasis>
<taxonomicName id="4C45CFC4FFF9FFD287AC662581A3B66C" authorityName="Demange" authorityYear="1970" box="[473,645,1542,1566]" class="Diplopoda" family="Spirostreptidae" genus="Pseudotibiozus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="554" phylum="Arthropoda" rank="genus">Pseudotibiozus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35FE7CCFFF9FFD186B266098667B3F6" lastPageId="2" lastPageNumber="543" pageId="1" pageNumber="542" type="description">
<paragraph id="8BFAB447FFF9FFD286B2660983C4B615" blockId="1.[151,1437,1254,2035]" pageId="1" pageNumber="542">
<emphasis id="B9316855FFF9FFD286B266098230B631" bold="true" box="[199,278,1578,1603]" pageId="1" pageNumber="542">Males.</emphasis>
SIZE. Body length
<quantity id="4CBD19A2FFF9FFD2879A6609814AB630" box="[495,620,1578,1603]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.3" metricValueMin="5.2" pageId="1" pageNumber="542" unit="mm" value="62.5" valueMax="73.0" valueMin="52.0">52–73 mm</quantity>
; diameter
<quantity id="4CBD19A2FFF9FFD284946609804DB630" box="[737,875,1578,1603]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.6" metricValueMin="4.4" pageId="1" pageNumber="542" unit="mm" value="5.5" valueMax="6.6" valueMin="4.4">4.4–6.6 mm</quantity>
. 41–43 podous rings, no apodous rings in front of telson.
</paragraph>
<paragraph id="8BFAB447FFF9FFD286B2665087B4B769" blockId="1.[151,1437,1254,2035]" pageId="1" pageNumber="542">
COLOUR. Live colour contrastful (black body, red legs) in
<taxonomicName id="4C45CFC4FFF9FFD2852D665780C0B6F9" authorityName="Attems" authorityYear="1914" box="[856,998,1652,1675]" class="Diplopoda" family="Spirostreptidae" genus="Mardonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="cerasopus">
<emphasis id="B9316855FFF9FFD2852D665780C0B6F9" box="[856,998,1652,1675]" italics="true" pageId="1" pageNumber="542">P. cerasopus</emphasis>
</taxonomicName>
(
<figureCitation id="137EA8C2FFF9FFD285836651871FB6F9" box="[1014,1081,1650,1675]" captionStart="FIGURE 1" captionStartId="3.[151,250,1941,1963]" captionTargetBox="[151,1436,1060,1918]" captionTargetId="figure@3.[151,1436,1060,1918]" captionTargetPageId="3" captionText="FIGURE 1. Pseudotibiozus cerasopus (Attems, 1914), specimen from the Udzungwa Mts. Photo by N. Scharff." httpUri="https://zenodo.org/record/1270437/files/figure.png" pageId="1" pageNumber="542">Fig. 1</figureCitation>
), not known in
<taxonomicName id="4C45CFC4FFF9FFD2829A665786BAB6F9" authority="Enghoff &amp; Larsson, 2018" authorityName="Enghoff &amp; Larsson" authorityYear="2018" box="[1263,1436,1650,1675]" class="Diplopoda" family="Spirostreptidae" genus="Pseudotibiozus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="zophoribates" status="sp. nov.">
<emphasis id="B9316855FFF9FFD2829A665786BAB6F9" box="[1263,1436,1650,1675]" italics="true" pageId="1" pageNumber="542">P. zophoribates</emphasis>
</taxonomicName>
<emphasis id="B9316855FFF9FFD286E266B583D4B6DD" bold="true" box="[151,242,1686,1711]" pageId="1" pageNumber="542">
<taxonomicNameLabel id="A202D52EFFF9FFD286E266B583D4B6DD" box="[151,242,1686,1711]" pageId="1" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Metazona of preserved specimens with a light band posteriorly. Dark colouration anteriorly of the band extended as tiny dark 'spike' (
<emphasis id="B9316855FFF9FFD28794669982DFB6A1" box="[481,505,1722,1747]" italics="true" pageId="1" pageNumber="542">ds</emphasis>
) into light posterior band at level of ozopore (
<emphasis id="B9316855FFF9FFD2858A669F873EB6A1" box="[1023,1048,1724,1747]" italics="true" pageId="1" pageNumber="542">oz</emphasis>
) (even on ozopore-less rings), one or two additional 'spikes' may also be present; 'spikes' difficult to see from the outside (
<figureCitation id="137EA8C2FFF9FFD282F966FD87C9B685" box="[1164,1263,1758,1783]" captionStart="FIGURE 2" captionStartId="4.[151,250,1158,1180]" captionTargetBox="[181,1405,221,1112]" captionTargetId="figure@4.[151,1436,193,1136]" captionTargetPageId="4" captionText="FIGURE 2. Pseudotibiozus spp. A–B: midbody rings seen from the inside, showing sigilla (light spots) and dark ‘spikes’ projecting into light posterior part of metazona. A: P. cerasopus, specimen from East Usambara Mts, Amani, B: P. zophoribates sp. nov., paratype. C: P. cerasopus, specimen from East Usambara Mts, Amani, parts of three consecutive body rings, seen from the outside, showing dark ‘spikes’. Scales 1 mm. Abbreviations: ds = dark ‘spike’, oz = ozopore." httpUri="https://zenodo.org/record/1270441/files/figure.png" pageId="1" pageNumber="542">Fig. 2 C</figureCitation>
), especially on fresh specimens, but conspicuous when the body ring is seen from the inside (
<figureCitation id="137EA8C2FFF9FFD2858A672187A3B769" box="[1023,1157,1794,1819]" captionStart="FIGURE 2" captionStartId="4.[151,250,1158,1180]" captionTargetBox="[181,1405,221,1112]" captionTargetId="figure@4.[151,1436,193,1136]" captionTargetPageId="4" captionText="FIGURE 2. Pseudotibiozus spp. A–B: midbody rings seen from the inside, showing sigilla (light spots) and dark ‘spikes’ projecting into light posterior part of metazona. A: P. cerasopus, specimen from East Usambara Mts, Amani, B: P. zophoribates sp. nov., paratype. C: P. cerasopus, specimen from East Usambara Mts, Amani, parts of three consecutive body rings, seen from the outside, showing dark ‘spikes’. Scales 1 mm. Abbreviations: ds = dark ‘spike’, oz = ozopore." httpUri="https://zenodo.org/record/1270441/files/figure.png" pageId="1" pageNumber="542">Figs 2 A, B</figureCitation>
).
</paragraph>
<paragraph id="8BFAB447FFF9FFD286B26704879AB781" blockId="1.[151,1437,1254,2035]" pageId="1" pageNumber="542">
HEAD (
<figureCitation id="137EA8C2FFF9FFD2876E67058243B74D" box="[283,357,1830,1855]" captionStart="FIGURE 3" captionStartId="5.[151,250,1734,1756]" captionTargetBox="[199,1386,233,1673]" captionTargetId="figure@5.[151,1436,193,1712]" captionTargetPageId="5" captionText="FIGURE 3. Pseudotibiozus spp., gnathochilarium. A–D: P. cerasopus, specimen from Udzungwa Mts. B: close-up of circled area (ss) in A. C: close-up of circled area in B. D: close-up of squared area in B. E: P. zophoribates sp. nov., paratype. Scales 1 mm (E), 0.2 mm (A), 0.05 mm (B), 0.005 mm (C, D). Abbreviations: ll = lamella lingualis, mda = median depressed area of mentum, sbs = basal field of stipital setae, ss = soft area with stout seta, st = stipes." httpUri="https://zenodo.org/record/1270443/files/figure.png" pageId="1" pageNumber="542">Figs 3</figureCitation>
,
<figureCitation id="137EA8C2FFF9FFD28700670482E1B74D" box="[373,455,1831,1855]" captionStart="FIGURE 4" captionStartId="6.[151,250,1728,1750]" captionTargetBox="[227,1396,233,1664]" captionTargetId="figure@6.[151,1436,193,1707]" captionTargetPageId="6" captionText="FIGURE 4. Pseudotibiozus spp. A: P. cerasopus (Attems, 1914), specimen from Udzungwa Mts, anterior end. B, C: P. zophoribates sp. nov., holotype. B: anterior end, C: telson. Scales 1 mm." httpUri="https://zenodo.org/record/1270445/files/figure.png" pageId="1" pageNumber="542">4 A, B</figureCitation>
). Four supralabral setae. Eyes extending mesad just beyond antennal sockets. Ca. 11 vertical rows of ommatidia, ca. 48 ommatidia. Mandibular stipes with apicoventral lobelike extension. (Details of mandibular gnathal lobe are described under
<taxonomicName id="4C45CFC4FFF9FFD284C1674C806EB7F4" authorityName="Attems" authorityYear="1914" box="[692,840,1903,1926]" class="Diplopoda" family="Spirostreptidae" genus="Mardonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="1" pageNumber="542" phylum="Arthropoda" rank="species" species="cerasopus">
<emphasis id="B9316855FFF9FFD284C1674C806EB7F4" box="[692,840,1903,1926]" italics="true" pageId="1" pageNumber="542">P. cerasopus</emphasis>
</taxonomicName>
). Gnathochilarium (
<figureCitation id="137EA8C2FFF9FFD2824D674D87A5B7F5" box="[1080,1155,1902,1927]" captionStart="FIGURE 3" captionStartId="5.[151,250,1734,1756]" captionTargetBox="[199,1386,233,1673]" captionTargetId="figure@5.[151,1436,193,1712]" captionTargetPageId="5" captionText="FIGURE 3. Pseudotibiozus spp., gnathochilarium. A–D: P. cerasopus, specimen from Udzungwa Mts. B: close-up of circled area (ss) in A. C: close-up of circled area in B. D: close-up of squared area in B. E: P. zophoribates sp. nov., paratype. Scales 1 mm (E), 0.2 mm (A), 0.05 mm (B), 0.005 mm (C, D). Abbreviations: ll = lamella lingualis, mda = median depressed area of mentum, sbs = basal field of stipital setae, ss = soft area with stout seta, st = stipes." httpUri="https://zenodo.org/record/1270443/files/figure.png" pageId="1" pageNumber="542">Fig. 3</figureCitation>
): Mentum with median depressed area (
<emphasis id="B9316855FFF9FFD2873A67B18259B7D9" box="[335,383,1938,1963]" italics="true" pageId="1" pageNumber="542">mda</emphasis>
); lamellae linguales (
<emphasis id="B9316855FFF9FFD2840D67B181AEB7D9" box="[632,648,1938,1963]" italics="true" pageId="1" pageNumber="542">ll</emphasis>
) each with two strong apical setae, one similar-sized seta behind the two and two to three basal setae of varying size; stipites (
<emphasis id="B9316855FFF9FFD2855A67948064B7BC" box="[815,834,1975,1998]" italics="true" pageId="1" pageNumber="542">st</emphasis>
) with several setae along latero-apical edge, with or without a field of setae (
<emphasis id="B9316855FFF9FFD287DD67F982EAB781" box="[424,460,2010,2035]" italics="true" pageId="1" pageNumber="542">sbs</emphasis>
) basally, with a single stout setae placed in a soft area (
<emphasis id="B9316855FFF9FFD2824F67FF8776B781" box="[1082,1104,2012,2035]" italics="true" pageId="1" pageNumber="542">ss</emphasis>
) near tip.
</paragraph>
<paragraph id="8BFAB447FFFAFFD186B260BB8085B0C2" blockId="2.[151,1437,151,900]" box="[199,931,151,176]" pageId="2" pageNumber="543">
COLLUM (
<figureCitation id="137EA8C2FFFAFFD1874D60B4829BB0C2" box="[312,445,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1728,1750]" captionTargetBox="[227,1396,233,1664]" captionTargetId="figure@6.[151,1436,193,1707]" captionTargetPageId="6" captionText="FIGURE 4. Pseudotibiozus spp. A: P. cerasopus (Attems, 1914), specimen from Udzungwa Mts, anterior end. B, C: P. zophoribates sp. nov., holotype. B: anterior end, C: telson. Scales 1 mm." httpUri="https://zenodo.org/record/1270445/files/figure.png" pageId="2" pageNumber="543">Figs 4 A, B</figureCitation>
). Lateral lobes rectangular, with 4–5 striae.
</paragraph>
<paragraph id="8BFAB447FFFAFFD186B2609E8713B14D" blockId="2.[151,1437,151,900]" pageId="2" pageNumber="543">
BODY RINGS (
<figureCitation id="137EA8C2FFFAFFD18716609F828AB0A6" box="[355,428,188,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1158,1180]" captionTargetBox="[181,1405,221,1112]" captionTargetId="figure@4.[151,1436,193,1136]" captionTargetPageId="4" captionText="FIGURE 2. Pseudotibiozus spp. A–B: midbody rings seen from the inside, showing sigilla (light spots) and dark ‘spikes’ projecting into light posterior part of metazona. A: P. cerasopus, specimen from East Usambara Mts, Amani, B: P. zophoribates sp. nov., paratype. C: P. cerasopus, specimen from East Usambara Mts, Amani, parts of three consecutive body rings, seen from the outside, showing dark ‘spikes’. Scales 1 mm. Abbreviations: ds = dark ‘spike’, oz = ozopore." httpUri="https://zenodo.org/record/1270441/files/figure.png" pageId="2" pageNumber="543">Figs 2</figureCitation>
,
<figureCitation id="137EA8C2FFFAFFD187CD609F82EFB0A6" box="[440,457,188,212]" captionStart="FIGURE 4" captionStartId="6.[151,250,1728,1750]" captionTargetBox="[227,1396,233,1664]" captionTargetId="figure@6.[151,1436,193,1707]" captionTargetPageId="6" captionText="FIGURE 4. Pseudotibiozus spp. A: P. cerasopus (Attems, 1914), specimen from Udzungwa Mts, anterior end. B, C: P. zophoribates sp. nov., holotype. B: anterior end, C: telson. Scales 1 mm." httpUri="https://zenodo.org/record/1270445/files/figure.png" pageId="2" pageNumber="543">4</figureCitation>
). Prozonal ring furrows very weak. Metazona vaulted and with longitudinal striae from below ozopores and downward, striae in lover part of striate area more ridgelike and ending in denticles at metazontal hind edge. Ozopore situated ca. 1/3 metazona length behind suture, a weak horizontal suture from ozopore to metazonal hind margin. Sigilla (
<figureCitation id="137EA8C2FFFAFFD18408610481E6B132" box="[637,704,295,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1158,1180]" captionTargetBox="[181,1405,221,1112]" captionTargetId="figure@4.[151,1436,193,1136]" captionTargetPageId="4" captionText="FIGURE 2. Pseudotibiozus spp. A–B: midbody rings seen from the inside, showing sigilla (light spots) and dark ‘spikes’ projecting into light posterior part of metazona. A: P. cerasopus, specimen from East Usambara Mts, Amani, B: P. zophoribates sp. nov., paratype. C: P. cerasopus, specimen from East Usambara Mts, Amani, parts of three consecutive body rings, seen from the outside, showing dark ‘spikes’. Scales 1 mm. Abbreviations: ds = dark ‘spike’, oz = ozopore." httpUri="https://zenodo.org/record/1270441/files/figure.png" pageId="2" pageNumber="543">Fig. 2</figureCitation>
) obvious, in one or several rows.
</paragraph>
<paragraph id="8BFAB447FFFAFFD186B2616E825DB1DF" blockId="2.[151,1437,151,900]" pageId="2" pageNumber="543">
TELSON (
<figureCitation id="137EA8C2FFFAFFD18745616F82B3B116" box="[304,405,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1728,1750]" captionTargetBox="[227,1396,233,1664]" captionTargetId="figure@6.[151,1436,193,1707]" captionTargetPageId="6" captionText="FIGURE 4. Pseudotibiozus spp. A: P. cerasopus (Attems, 1914), specimen from Udzungwa Mts, anterior end. B, C: P. zophoribates sp. nov., holotype. B: anterior end, C: telson. Scales 1 mm." httpUri="https://zenodo.org/record/1270445/files/figure.png" pageId="2" pageNumber="543">Fig. 4 C</figureCitation>
). Preanal ring with slightly curved dorsal margin, posteriorly with a more or less straight edge, i.e., no proper epiproct; hypoproct a narrow band; anal valves with a submarginal depression, margins forming blunt ‘lips’.
</paragraph>
<paragraph id="8BFAB447FFFAFFD186B2619B8142B26A" blockId="2.[151,1437,151,900]" pageId="2" pageNumber="543">
LEGS. As long as body diameter or slightly shorter. Tibia with small apical ventral pad extending under basis of tarsus; no pads on other podomeres. First pair (
<figureCitation id="137EA8C2FFFAFFD184C661FF81D1B187" box="[691,759,476,501]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[183,1405,380,1884]" captionTargetId="figure@7.[151,1436,349,1922]" captionTargetPageId="7" captionText="FIGURE 5. Pseudotibiozus spp., A, B: first pair of male legs. C, D: left gonopod telopodite. A, C: P. cerasopus (Attems, 1914) specimen from Udzungwa Mts. B, D: P. zophoribates sp. nov., paratype. Scales 0.2 mm (A), 0.5 mm (B—D). Abbreviations: atp = antetorsal process, cxs = field of coxosternal setae, pfl = prefemoral lobe." httpUri="https://zenodo.org/record/1270447/files/figure.png" pageId="2" pageNumber="543">Fig. 5</figureCitation>
) with short, triangular basal prefemoral lobes (
<emphasis id="B9316855FFFAFFD1836161FF8617B187" box="[1300,1329,476,501]" italics="true" pageId="2" pageNumber="543">pfl</emphasis>
), with or without a field of coxosternal setae (
<emphasis id="B9316855FFFAFFD1844562228172B26A" box="[560,596,513,536]" italics="true" pageId="2" pageNumber="543">cxs</emphasis>
).
</paragraph>
<paragraph id="8BFAB447FFFAFFD186B262078007B34A" blockId="2.[151,1437,151,900]" pageId="2" pageNumber="543">
GONOPODS (
<figureCitation id="137EA8C2FFFAFFD18723620782C7B24E" box="[342,481,548,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[183,1405,380,1884]" captionTargetId="figure@7.[151,1436,349,1922]" captionTargetPageId="7" captionText="FIGURE 5. Pseudotibiozus spp., A, B: first pair of male legs. C, D: left gonopod telopodite. A, C: P. cerasopus (Attems, 1914) specimen from Udzungwa Mts. B, D: P. zophoribates sp. nov., paratype. Scales 0.2 mm (A), 0.5 mm (B—D). Abbreviations: atp = antetorsal process, cxs = field of coxosternal setae, pfl = prefemoral lobe." httpUri="https://zenodo.org/record/1270447/files/figure.png" pageId="2" pageNumber="543">Figs 5 C, D</figureCitation>
,
<figureCitation id="137EA8C2FFFAFFD1879B620682D8B24E" box="[494,510,549,572]" captionStart="FIGURE 7" captionStartId="9.[151,250,1866,1888]" captionTargetBox="[190,1402,189,1822]" captionTargetId="figure@9.[151,1436,157,1885]" captionTargetPageId="9" captionText="FIGURE 7. Pseudotibiozus cerasopus (Attems, 1914), specimen from Udzungwa Mts, gonopods. A: anterior view. B: posterior view. C: left telopodite. D: proplical setae (circled area in A). E: tip of solenomere. Scales 0.2 mm (A, B), 0.1 mm (C), 0.02 mm (D), 0.01 mm (E). Abbreviations: aml = apical metaplical lobe, apl = apical proplical lobe, map = mesapical process of proplica, mp = metaplica, pp= proplica, px = paracoxite, sb = spinelike branch, slm = solenomere, st = sternum, tlp = telopodite." httpUri="https://zenodo.org/record/1270451/files/figure.png" pageId="2" pageNumber="543">7</figureCitation>
,
<figureCitation id="137EA8C2FFFAFFD184796207813AB24E" box="[524,540,548,572]" captionStart="FIGURE 8" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[224,1374,194,1848]" captionTargetId="figure@10.[152,1435,101,1915]" captionTargetPageId="10" captionText="FIGURE 8. Pseudotibiozus zophoribates sp. nov., holotype, gonopods (sternum missing). A: anterior view. B: posterior view. C: Tip of left pro- and metaplica. D: close-up of squared area in C, outline of antetorsal process traced in black. Scales 0.5 mm (A, B), 0.2 mm (C, D). Abbreviations: aml = apical metaplical lobe, apl = apical proplical lobe, atp = antetorsal process." httpUri="https://zenodo.org/record/1270453/files/figure.png" pageId="2" pageNumber="543">8</figureCitation>
). Sternum (
<emphasis id="B9316855FFFAFFD184D162068191B24E" box="[676,695,549,572]" italics="true" pageId="2" pageNumber="543">st</emphasis>
) blunt triangular to subsemicircular, not reaching as far distal as paracoxites (
<emphasis id="B9316855FFFAFFD18752626A8267B212" box="[295,321,585,608]" italics="true" pageId="2" pageNumber="543">px</emphasis>
). Proplica (
<emphasis id="B9316855FFFAFFD187BD626A82C2B212" box="[456,484,585,608]" italics="true" pageId="2" pageNumber="543">pp</emphasis>
) straight, ending in a simple apical lobe (
<emphasis id="B9316855FFFAFFD185C9626480C6B212" box="[956,992,583,608]" italics="true" pageId="2" pageNumber="543">apl</emphasis>
) with a small patch of short setae at its base. Metaplica (
<emphasis id="B9316855FFFAFFD1872D624E825CB2F6" box="[344,378,621,644]" italics="true" pageId="2" pageNumber="543">mp</emphasis>
) longer than proplica, apically with a triangular-rounded or ovoid lobe (
<emphasis id="B9316855FFFAFFD182DE624F87F3B2F7" box="[1195,1237,620,645]" italics="true" pageId="2" pageNumber="543">aml</emphasis>
). Telopodite (
<emphasis id="B9316855FFFAFFD18300624F86B4B2F7" box="[1397,1426,620,645]" italics="true" pageId="2" pageNumber="543">tlp</emphasis>
) (
<figureCitation id="137EA8C2FFFAFFD186EA62AC820EB2DA" box="[159,296,655,680]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1969]" captionTargetBox="[183,1405,380,1884]" captionTargetId="figure@7.[151,1436,349,1922]" captionTargetPageId="7" captionText="FIGURE 5. Pseudotibiozus spp., A, B: first pair of male legs. C, D: left gonopod telopodite. A, C: P. cerasopus (Attems, 1914) specimen from Udzungwa Mts. B, D: P. zophoribates sp. nov., paratype. Scales 0.2 mm (A), 0.5 mm (B—D). Abbreviations: atp = antetorsal process, cxs = field of coxosternal setae, pfl = prefemoral lobe." httpUri="https://zenodo.org/record/1270447/files/figure.png" pageId="2" pageNumber="543">Figs 5 C, D</figureCitation>
) with a very small antetorsal process (
<emphasis id="B9316855FFFAFFD1849262B2802DB2DA" box="[743,779,657,680]" italics="true" pageId="2" pageNumber="543">atp</emphasis>
) originating before the telopodite bends laterad and exits from the coxa,
<emphasis id="B9316855FFFAFFD1873662968241B2BE" box="[323,359,693,716]" italics="true" pageId="2" pageNumber="543">atp</emphasis>
hence either completely hidden between pro- and metaplica or barely projecting (
<figureCitation id="137EA8C2FFFAFFD18373629786B6B2BE" box="[1286,1424,692,717]" captionStart="FIGURE 8" captionStartId="10.[151,250,1937,1959]" captionTargetBox="[224,1374,194,1848]" captionTargetId="figure@10.[152,1435,101,1915]" captionTargetPageId="10" captionText="FIGURE 8. Pseudotibiozus zophoribates sp. nov., holotype, gonopods (sternum missing). A: anterior view. B: posterior view. C: Tip of left pro- and metaplica. D: close-up of squared area in C, outline of antetorsal process traced in black. Scales 0.5 mm (A, B), 0.2 mm (C, D). Abbreviations: aml = apical metaplical lobe, apl = apical proplical lobe, atp = antetorsal process." httpUri="https://zenodo.org/record/1270453/files/figure.png" pageId="2" pageNumber="543">Figs 8 C, D</figureCitation>
). Telopodite ribbonlike from emergence from coxa, ca. at midlength bending mesad and dividing into a mesad spinelike branch (
<emphasis id="B9316855FFFAFFD1872A62DF825EB367" box="[351,376,764,789]" italics="true" pageId="2" pageNumber="543">sb</emphasis>
) and a thinner solenomere (
<emphasis id="B9316855FFFAFFD184C662DF81FCB367" box="[691,730,764,789]" italics="true" pageId="2" pageNumber="543">slm</emphasis>
); solenomere twisting around
<emphasis id="B9316855FFFAFFD1824562DF876FB367" box="[1072,1097,764,789]" italics="true" pageId="2" pageNumber="543">sb</emphasis>
in an overall mesad direction. Tip of telopodite strongly wrinkled, spongelike (
<figureCitation id="137EA8C2FFFAFFD184C3633C8035B34A" box="[694,787,799,824]" captionStart="FIGURE 7" captionStartId="9.[151,250,1866,1888]" captionTargetBox="[190,1402,189,1822]" captionTargetId="figure@9.[151,1436,157,1885]" captionTargetPageId="9" captionText="FIGURE 7. Pseudotibiozus cerasopus (Attems, 1914), specimen from Udzungwa Mts, gonopods. A: anterior view. B: posterior view. C: left telopodite. D: proplical setae (circled area in A). E: tip of solenomere. Scales 0.2 mm (A, B), 0.1 mm (C), 0.02 mm (D), 0.01 mm (E). Abbreviations: aml = apical metaplical lobe, apl = apical proplical lobe, map = mesapical process of proplica, mp = metaplica, pp= proplica, px = paracoxite, sb = spinelike branch, slm = solenomere, st = sternum, tlp = telopodite." httpUri="https://zenodo.org/record/1270451/files/figure.png" pageId="2" pageNumber="543">Fig. 7 E</figureCitation>
).
</paragraph>
<paragraph id="8BFAB447FFFAFFD186B263648667B3F6" blockId="2.[151,1437,151,900]" pageId="2" pageNumber="543">
<emphasis id="B9316855FFFAFFD186B263648136B312" bold="true" box="[199,528,839,864]" pageId="2" pageNumber="543">Females (known only for</emphasis>
<taxonomicName id="4C45CFC4FFFAFFD18457636B819AB32D" authorityName="Attems" authorityYear="1914" box="[546,700,840,863]" class="Diplopoda" family="Spirostreptidae" genus="Mardonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirostreptida" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="cerasopus">P. cerasopus</taxonomicName>
<emphasis id="B9316855FFFAFFD184C9636481EAB312" bold="true" box="[700,716,839,864]" pageId="2" pageNumber="543">).</emphasis>
Thicker than males, with relatively shorter legs, lacking modifications of mandibular stipes and legs. 40–44 podous body rings, no apodous rings in front of telson.
</paragraph>
</subSubSection>
</treatment>
</document>
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