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<document id="7FC83FC8F36818F8B6E14BE31794E702" ID-CLB-Dataset="2910" ID-DOI="10.11646/zootaxa.5082.3.2" ID-GBIF-Dataset="3997d831-ae29-42d0-ba99-cd20cc8b7fc3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5788224" ID-ZooBank="EFB0E4A5-6B60-4B39-A651-1C6470FD7222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639730480321" checkinUser="plazi" docAuthor="Li, Yan-Da, Tihelka, Erik, Dahan, Loïc, Huang, Di-Ying &amp; Cai, Chen-Yang" docDate="2021" docId="03D787A2FFE8EE7DFF52FA17891B6B59" docLanguage="en" docName="zootaxa.5082.3.2.pdf" docOrigin="Zootaxa 5082 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archaenosodendron remotidens Li &amp; Cai 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="233" masterDocId="FFEEFFDAFFE0EE77FFC5FFAA8B796C12" masterDocTitle="On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)" masterLastPageNumber="244" masterPageNumber="223" pageNumber="231" updateTime="1699472730423" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7A58C2C1E9E63D9E37322C59D60FFF89">On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)</mods:title>
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<mods:namePart id="EF5BC687599A2D7BC0B5E49E5CB6C7DC">Li, Yan-Da</mods:namePart>
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<mods:namePart id="755D4399EDAD2CE4620898CEABD9CCFA">Tihelka, Erik</mods:namePart>
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<mods:affiliation id="04BA05081841E1B3C2D9E0678B4C4B55">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK &amp; wn 20250 @ bristol. ac. uk; https: // orcid. org / 0000 - 0002 - 5048 - 5355</mods:affiliation>
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<mods:namePart id="41A2F7931B059258B34298EF221A917F">Dahan, Loïc</mods:namePart>
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<mods:affiliation id="AB9A289C5C50EAC48B11D5A8915FD98A">O. D. Taxonomy and Phylogeny - Entomology, Royal Belgian Institute of Natural Sciences, Vautierstreet 29, B- 1000 Brussels, Belgium ydli @ pku. edu. cn; https: // orcid. org / 0000 - 0002 - 9439 - 202 X &amp; loicdahan @ hotmail. com; https: // orcid. org / 0000 - 0001 - 5680 - 0304</mods:affiliation>
<mods:nameIdentifier id="2EF2254D9622903BBFB4BED2D0FC20E1" type="email">loicdahan@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="277F31D4B950B78153DDB3F90DED9FFF">Huang, Di-Ying</mods:namePart>
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<mods:affiliation id="73E830ADACAEDFD9AA93F063E47DE103">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; dyhuang @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 5637 - 4867</mods:affiliation>
<mods:nameIdentifier id="D8F528A70D25BB18CE35A793C937E492" type="email">dyhuang@nigpas.ac.cn</mods:nameIdentifier>
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<mods:namePart id="361D2BF1523152BF81A864B2C6B61986">Cai, Chen-Yang</mods:namePart>
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<mods:affiliation id="C834383A48C13DECECF2FC5CB6565879">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China &amp; School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol BS 8 1 TQ, UK &amp; cycai @ nigpas. ac. cn; https: // orcid. org / 0000 - 0002 - 9283 - 8323</mods:affiliation>
<mods:nameIdentifier id="51571071AA2F7BFB902EFC701045411A" type="email">cycai@nigpas.ac.cn</mods:nameIdentifier>
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<treatment id="03D787A2FFE8EE7DFF52FA17891B6B59" ID-DOI="http://doi.org/10.5281/zenodo.5788236" ID-GBIF-Taxon="191654783" ID-Zenodo-Dep="5788236" LSID="urn:lsid:plazi:treatment:03D787A2FFE8EE7DFF52FA17891B6B59" httpUri="http://treatment.plazi.org/id/03D787A2FFE8EE7DFF52FA17891B6B59" lastPageId="10" lastPageNumber="233" pageId="8" pageNumber="231">
<subSubSection id="C364653FFFE8EE7FFF52FA17898C69C5" box="[151,757,1469,1496]" pageId="8" pageNumber="231" type="nomenclature">
<paragraph id="8BC136B4FFE8EE7FFF52FA17898C69C5" blockId="8.[151,757,1469,1531]" box="[151,757,1469,1496]" pageId="8" pageNumber="231">
<heading id="D08981D8FFE8EE7FFF52FA17898C69C5" bold="true" box="[151,757,1469,1496]" fontSize="11" level="1" pageId="8" pageNumber="231" reason="1">
<emphasis id="B90AEAA6FFE8EE7FFF52FA17898C69C5" bold="true" box="[151,757,1469,1496]" pageId="8" pageNumber="231">
<taxonomicName id="4C7E4D37FFE8EE7FFF52FA1789E969C5" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2021" box="[151,656,1469,1496]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="231" phylum="Arthropoda" rank="species" species="remotidens" status="sp. nov.">
<emphasis id="B90AEAA6FFE8EE7FFF52FA17896269CA" bold="true" box="[151,539,1469,1496]" italics="true" pageId="8" pageNumber="231">Archaenosodendron remotidens</emphasis>
Li &amp; Cai
</taxonomicName>
<taxonomicNameLabel id="A23957DDFFE8EE7FFD5DFA17898C69C5" box="[664,757,1469,1495]" pageId="8" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFE8EE7FFF52FA4B8A6169E9" box="[151,280,1505,1531]" pageId="8" pageNumber="231" type="description">
<paragraph id="8BC136B4FFE8EE7FFF52FA4B8A6169E9" blockId="8.[151,757,1469,1531]" box="[151,280,1505,1531]" pageId="8" pageNumber="231">
(
<figureCitation id="13452A31FFE8EE7FFF5AFA4B8A6969E9" box="[159,272,1505,1531]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="10.[151,250,1447,1471]" captionStartId-1="11.[151,250,1447,1471]" captionStartId-2="12.[151,250,1503,1527]" captionTargetBox-0="[151,1436,181,1423]" captionTargetBox-1="[151,1436,181,1423]" captionTargetBox-2="[151,1436,181,1480]" captionTargetId-0="figure-22@10.[151,1436,181,1423]" captionTargetId-1="figure-18@11.[151,1436,181,1423]" captionTargetId-2="figure-22@12.[151,1436,181,1480]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 9. General habitus of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under incident light. A, Dorsal view. B, Ventral view. Scale bars: 1 mm." captionText-1="FIGURE 10. General habitus of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under widefield fluorescence. A, Dorsal view. B, Ventral view. Scale bars: 1 mm." captionText-2="FIGURE 11. Details of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi-0="http://doi.org/10.5281/zenodo.5788253" figureDoi-1="http://doi.org/10.5281/zenodo.5788255" figureDoi-2="http://doi.org/10.5281/zenodo.5788257" httpUri-0="https://zenodo.org/record/5788253/files/figure.png" httpUri-1="https://zenodo.org/record/5788255/files/figure.png" httpUri-2="https://zenodo.org/record/5788257/files/figure.png" pageId="8" pageNumber="231">Figs 9–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFE8EE7FFF52F98389586A51" box="[151,545,1577,1603]" pageId="8" pageNumber="231" type="materials_examined">
<paragraph id="8BC136B4FFE8EE7FFF52F98389586A51" blockId="8.[151,1437,1577,2036]" box="[151,545,1577,1603]" pageId="8" pageNumber="231">
<emphasis id="B90AEAA6FFE8EE7FFF52F9838A7F6A51" bold="true" box="[151,262,1577,1603]" pageId="8" pageNumber="231">Material.</emphasis>
<materialsCitation id="3B163CE9FFE8EE7FFEC8F98389586A51" ID-GBIF-Occurrence="3419363301" box="[269,545,1577,1603]" collectionCode="NIGP" pageId="8" pageNumber="231" specimenCode="NIGP177614" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C58816FFE8EE7FFEC8F9838A016A51" box="[269,376,1577,1603]" pageId="8" pageNumber="231" type="holotype">Holotype</typeStatus>
,
<specimenCode id="DBD89ECFFFE8EE7FFE41F98389646A51" box="[388,541,1577,1603]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="8" pageNumber="231">NIGP177614</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFE8EE7FFF02F9E78F9B6AC1" pageId="8" pageNumber="231" type="etymology">
<paragraph id="8BC136B4FFE8EE7FFF02F9E78A4F6A99" blockId="8.[151,1437,1577,2036]" pageId="8" pageNumber="231">
<emphasis id="B90AEAA6FFE8EE7FFF02F9E78A376A75" bold="true" box="[199,334,1613,1639]" pageId="8" pageNumber="231">Etymology.</emphasis>
The specific name refers to the relatively sparsely arranged denticles/stout setae on the outer edge of its protibia.
</paragraph>
<paragraph id="8BC136B4FFE8EE7FFF02F93F8F9B6AC1" blockId="8.[151,1437,1577,2036]" pageId="8" pageNumber="231">
<emphasis id="B90AEAA6FFE8EE7FFF02F93F8AB66ABD" bold="true" box="[199,463,1685,1711]" pageId="8" pageNumber="231">Locality and horizon.</emphasis>
Amber mine located near Noije Bum Village, Tanai Township, Myitkyina District,
<collectingRegion id="49BAF856FFE8EE7FFF52F9138A536AC1" box="[151,298,1721,1747]" country="Myanmar" name="Kachin" pageId="8" pageNumber="231">Kachin State</collectingRegion>
,
<collectingCountry id="F3697624FFE8EE7FFEF0F9108ADF6AC6" box="[309,422,1722,1748]" name="Myanmar" pageId="8" pageNumber="231">Myanmar</collectingCountry>
; unnamed horizon, mid-Cretaceous, Upper Albian to Lower Cenomanian.
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFE8EE7FFF02F9778E416B95" pageId="8" pageNumber="231" type="diagnosis">
<paragraph id="8BC136B4FFE8EE7FFF02F9778E416B95" blockId="8.[151,1437,1577,2036]" pageId="8" pageNumber="231">
<emphasis id="B90AEAA6FFE8EE7FFF02F9778A3B6AE5" bold="true" box="[199,322,1757,1783]" pageId="8" pageNumber="231">Diagnosis.</emphasis>
Medial antennomeres longer than wide (
<figureCitation id="13452A31FFE8EE7FFCDCF97788F96AE5" box="[793,896,1757,1783]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-22@12.[151,1436,181,1480]" captionTargetPageId="12" captionText="FIGURE 11. Details of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788257" httpUri="https://zenodo.org/record/5788257/files/figure.png" pageId="8" pageNumber="231">Fig. 11C</figureCitation>
). Lateral regions of mentum without longitudinally arranged larger punctures (
<figureCitation id="13452A31FFE8EE7FFDC5F8AB891C6B09" box="[512,613,1793,1819]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-22@12.[151,1436,181,1480]" captionTargetPageId="12" captionText="FIGURE 11. Details of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788257" httpUri="https://zenodo.org/record/5788257/files/figure.png" pageId="8" pageNumber="231">Fig. 11B</figureCitation>
). Anterior angles of mentum not obviously produced (
<figureCitation id="13452A31FFE8EE7FFB7AF8AB8E5D6B09" box="[1215,1316,1793,1819]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-22@12.[151,1436,181,1480]" captionTargetPageId="12" captionText="FIGURE 11. Details of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788257" httpUri="https://zenodo.org/record/5788257/files/figure.png" pageId="8" pageNumber="231">Fig. 11B</figureCitation>
). Antennal grooves on prosternum shorter than half-length of prosternum in front of coxae (
<figureCitation id="13452A31FFE8EE7FFBE0F88F8FF46B2D" box="[1061,1165,1829,1855]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-22@12.[151,1436,181,1480]" captionTargetPageId="12" captionText="FIGURE 11. Details of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788257" httpUri="https://zenodo.org/record/5788257/files/figure.png" pageId="8" pageNumber="231">Fig. 11D</figureCitation>
). Outer edge of protibia with about 14–16 relatively sparsely arranged denticles/stout setae, with the apical two distinctly stouter and more closely situated (
<figureCitation id="13452A31FFE8EE7FFE91F8C78AC26B95" box="[340,443,1901,1927]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-22@12.[151,1436,181,1480]" captionTargetPageId="12" captionText="FIGURE 11. Details of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788257" httpUri="https://zenodo.org/record/5788257/files/figure.png" pageId="8" pageNumber="231">Fig. 11G</figureCitation>
). Medial and lateral portions of prosternum not separated by a ridge (
<figureCitation id="13452A31FFE8EE7FFB01F8C78E526B95" box="[1220,1323,1901,1927]" captionStart="FIGURE 11" captionStartId="12.[151,250,1503,1527]" captionTargetBox="[151,1436,181,1480]" captionTargetId="figure-22@12.[151,1436,181,1480]" captionTargetPageId="12" captionText="FIGURE 11. Details of Archaenosodendron remotidens Li &amp; Cai sp. nov., holotype, NIGP177614, under confocal microscopy. A, Head, dorsal view. B, Mouthparts, ventral view. C, Antenna, ventral view. D, Prothorax, ventral view, showing the incomplete antennal groove on prosternum (arrowhead). E, Abdominal base, ventral view. F, Scutellar shield, dorsal view. G, Fore leg. H, Mesotibia and -tarsus. I, Metatibia and -tarsus. Abbreviations: an9–11, antennomeres 9–11; el, elytron; gl, galea; lbp, labial palp; md, mandible; mstb, mesotibia; msts, mesotarsus; msv, mesoventrite; mt, mentum; mtc, metacoxa; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; mxp, maxillary palp; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; pts, protarsus; sc, scutellum. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.5788257" httpUri="https://zenodo.org/record/5788257/files/figure.png" pageId="8" pageNumber="231">Fig. 11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C364653FFFE8EE7DFF02F83B891B6B59" lastPageId="10" lastPageNumber="233" pageId="8" pageNumber="231" type="description">
<paragraph id="8BC136B4FFE8EE7FFF02F83B88AB6BB9" blockId="8.[151,1437,1577,2036]" box="[199,978,1937,1964]" pageId="8" pageNumber="231">
<emphasis id="B90AEAA6FFE8EE7FFF02F83B8A206BB9" bold="true" box="[199,345,1937,1963]" pageId="8" pageNumber="231">Description.</emphasis>
Body elongated oval, about
<quantity id="4C869B51FFE8EE7FFD5EF83B89886BBE" box="[667,753,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="231" unit="mm" value="3.6">3.6 mm</quantity>
long,
<quantity id="4C869B51FFE8EE7FFCF2F83B88F46BBE" box="[823,909,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="231" unit="mm" value="1.8">1.8 mm</quantity>
wide.
</paragraph>
<paragraph id="8BC136B4FFE8EE7EFF02F81F8F2C6D53" blockId="8.[151,1437,1577,2036]" lastBlockId="9.[151,1437,151,501]" lastPageId="9" lastPageNumber="232" pageId="8" pageNumber="231">
Head prognathous. Eyes not strongly protuberant. Antennal insertions widely separated and concealed from above. Antennal grooves on head well developed. Frontoclypeal suture absent. Antennae 11-segmented; antennomere 2 about twice as long as wide; antennomere 3 about as long as but slender than 2; antennomeres 3–8 progressively shorter and wider; antennomeres 9–11 enlarged, forming a relatively loose club. Mandibles broad, unidentate. Maxilla with setose galea and lacinia; maxillary palps 4-segmented; palpomeres 1 and 4 longer than 2 and 3.
<collectingRegion id="49BAF856FFE9EE7EFF52FEA98B8F6D0F" box="[151,246,259,285]" country="Myanmar" name="Chin" pageId="9" pageNumber="232">Mentum</collectingRegion>
with lateral edges subparallel in the basal half, then converging anteriorly; surface without clearly visible punctures; anterior angles not produced. Apical labial palpomere seemingly enlarged.
</paragraph>
<paragraph id="8BC136B4FFE9EE7EFF02FEE18AC96DE7" blockId="9.[151,1437,151,501]" pageId="9" pageNumber="232">Pronotal disc subtrapezoidal, widest at base; anterior margin with shallow emarginations laterally, slightly protruding medially. Prosternum in front coxae transverse, not extremely shortened; medial and lateral portions of prosternum not separated by a ridge; antennal grooves on prosternum shorter than half-length of prosternum in front of coxae; prosternal process relatively slender. Procoxae strongly transverse, well separated. Procoxal cavities broadly open posteriorly.</paragraph>
<caption id="DF01663CFFE9EE7EFF52F89D88B86BCD" ID-DOI="http://doi.org/10.5281/zenodo.5788251" ID-Zenodo-Dep="5788251" httpUri="https://zenodo.org/record/5788251/files/figure.png" pageId="9" pageNumber="232" startId="9.[151,250,1847,1871]" targetBox="[151,1436,525,1824]" targetPageId="9">
<paragraph id="8BC136B4FFE9EE7EFF52F89D88B86BCD" blockId="9.[151,1437,1847,2015]" pageId="9" pageNumber="232">
<emphasis id="B90AEAA6FFE9EE7EFF52F89D8A6B6B5D" bold="true" box="[151,274,1847,1871]" pageId="9" pageNumber="232">FIGURE 8.</emphasis>
Details of
<taxonomicName id="4C7E4D37FFE9EE7EFE47F89D88506B5D" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2021" box="[386,809,1847,1871]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="232" phylum="Arthropoda" rank="species" species="explanatum" status="sp. nov.">
<emphasis id="B90AEAA6FFE9EE7EFE47F89D89B06B5D" box="[386,713,1847,1871]" italics="true" pageId="9" pageNumber="232">Archaenosodendron explanatum</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="B90AEAA6FFE9EE7EFCEAF89D88066B5D" bold="true" box="[815,895,1847,1871]" pageId="9" pageNumber="232">
<taxonomicNameLabel id="A23957DDFFE9EE7EFCEAF89D88066B5D" box="[815,895,1847,1871]" pageId="9" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP177613, under confocal microscopy.
<emphasis id="B90AEAA6FFE9EE7EFF52F8F18BD06B61" bold="true" box="[151,169,1883,1907]" pageId="9" pageNumber="232">A</emphasis>
, Head, dorsal view.
<emphasis id="B90AEAA6FFE9EE7EFEBFF8F18AF26B61" bold="true" box="[378,395,1883,1907]" pageId="9" pageNumber="232">B</emphasis>
, Head, ventral view.
<emphasis id="B90AEAA6FFE9EE7EFDA0F8F1890E6B61" bold="true" box="[613,631,1883,1907]" pageId="9" pageNumber="232">C</emphasis>
, Head, lateral view.
<emphasis id="B90AEAA6FFE9EE7EFC8CF8F188226B61" bold="true" box="[841,859,1883,1907]" pageId="9" pageNumber="232">D</emphasis>
, Prothorax, ventral view, showing the incomplete antennal groove on prosternum.
<emphasis id="B90AEAA6FFE9EE7EFE6AF8D58A996B85" bold="true" box="[431,480,1919,1943]" pageId="9" pageNumber="232">E–G</emphasis>
, Fore legs.
<emphasis id="B90AEAA6FFE9EE7EFD9EF8D589166B85" bold="true" box="[603,623,1919,1943]" pageId="9" pageNumber="232">H</emphasis>
, Mid leg.
<emphasis id="B90AEAA6FFE9EE7EFD19F8D5899F6B85" bold="true" box="[732,742,1919,1943]" pageId="9" pageNumber="232">I</emphasis>
, Hind leg. Abbreviations: an1, antennomere 1; md, mandible; msv, mesoventrite; mt, mentum; mtf, metafemur; mttb, metatibia; mtts, metatarsus; mtv, metaventrite; pc, procoxa; pf, profemur; pn, pronotum; ps, prosternum; ptb, protibia; v1,2, ventrites 1,2. Scale bars: 200 μm.
</paragraph>
</caption>
<caption id="DF01663CFFEAEE7DFF52FA0D89E569F0" ID-DOI="http://doi.org/10.5281/zenodo.5788253" ID-Zenodo-Dep="5788253" httpUri="https://zenodo.org/record/5788253/files/figure.png" pageId="10" pageNumber="233" startId="10.[151,250,1447,1471]" targetBox="[151,1436,181,1423]" targetPageId="10">
<paragraph id="8BC136B4FFEAEE7DFF52FA0D89E569F0" blockId="10.[151,1437,1447,1507]" pageId="10" pageNumber="233">
<emphasis id="B90AEAA6FFEAEE7DFF52FA0D8A6B69AD" bold="true" box="[151,274,1447,1471]" pageId="10" pageNumber="233">FIGURE 9.</emphasis>
General habitus of
<taxonomicName id="4C7E4D37FFEAEE7DFE1DFA0D880E69AD" authority="Li &amp; Cai" authorityName="Li &amp; Cai" authorityYear="2021" box="[472,887,1447,1471]" class="Insecta" family="Nosodendridae" genus="Archaenosodendron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="233" phylum="Arthropoda" rank="species" species="remotidens" status="sp. nov.">
<emphasis id="B90AEAA6FFEAEE7DFE1DFA0D886069AD" box="[472,793,1447,1471]" italics="true" pageId="10" pageNumber="233">Archaenosodendron remotidens</emphasis>
Li &amp; Cai
</taxonomicName>
<emphasis id="B90AEAA6FFEAEE7DFCB8FA0D88B569AD" bold="true" box="[893,972,1447,1471]" pageId="10" pageNumber="233">
<taxonomicNameLabel id="A23957DDFFEAEE7DFCB8FA0D88B569AD" box="[893,972,1447,1471]" pageId="10" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NIGP177614, under incident light.
<emphasis id="B90AEAA6FFEAEE7DFF52FA618BD069F1" bold="true" box="[151,169,1483,1507]" pageId="10" pageNumber="233">A</emphasis>
, Dorsal view.
<emphasis id="B90AEAA6FFEAEE7DFEF9FA618A3469F1" bold="true" box="[316,333,1483,1507]" pageId="10" pageNumber="233">B</emphasis>
, Ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BC136B4FFEAEE7DFF02F9BB8F686A61" blockId="10.[151,1437,1553,1867]" pageId="10" pageNumber="233">Scutellar shield triangular, posteriorly acute. Elytra widest at anterior third; anterior corners weakly obtuse. Mesoventrite short, anteriorly with well-developed mesoventral cavity and procoxal rests. Mesocoxal cavities widely separated. Metaventrite transverse, flattened. Metacoxae narrowly separated.</paragraph>
<paragraph id="8BC136B4FFEAEE7DFF02F9D48AA56B11" blockId="10.[151,1437,1553,1867]" pageId="10" pageNumber="233">Femora with groove for receiving tibia. Tibiae expanded distally, with two apical spurs; outer edge of protibia with relatively sparsely arranged denticles/stout setae, with the apical two distinctly stouter and more closely situated. Tarsi 5-5-5; tarsomeres slender, not distinctly lobed; tarsomeres 1–4 ventrally with a cluster of long hairs at apex. Pretarsal claws simple.</paragraph>
<paragraph id="8BC136B4FFEAEE7DFF02F8A7891B6B59" blockId="10.[151,1437,1553,1867]" pageId="10" pageNumber="233">Abdomen with five ventrites. Ventrites 1 and 2 with a pair of impressions for receiving hind legs. Anterior margin of ventrites smooth, without notches.</paragraph>
</subSubSection>
</treatment>
</document>
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