treatments-xml/data/03/D7/87/03D78797B34AFFD4FF3F40FAFEA24296.xml

<document id="92E5A9184961AB2259A6EF4E93F335EC" ID-DOI="10.5281/zenodo.196937" ID-GBIF-Dataset="10fa8557-a024-4949-ab0f-28706b1e6736" ID-ISSN="1175-5326" ID-Zenodo-Dep="196937" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460258187564" checkinUser="plazi" docAuthor="Hadiaty, Renny Kurnia &amp; Kottelat, Maurice" docDate="2010" docId="03D78797B34AFFD4FF3F40FAFEA24296" docLanguage="en" docName="zt02557p048.pdf" docOrigin="Zootaxa 2557" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nemacheilus marang Hadiaty &amp; Kottelat, 2010, new species" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FFEEFFEFB348FFDCFFA8406DFE514548" masterDocTitle="Nemacheilus marang, a new loach (Teleostei: Nemacheilidae) from Sangkulirang karst, eastern Borneo" masterLastPageNumber="48" masterPageNumber="39" pageNumber="41" updateTime="1698243760367" updateUser="plazi">
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<mods:title id="79754F009337BE1DE6B787D7C6EC542B">Nemacheilus marang, a new loach (Teleostei: Nemacheilidae) from Sangkulirang karst, eastern Borneo</mods:title>
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<mods:namePart id="12007C37924BD6B8E745719400D9035F">Hadiaty, Renny Kurnia</mods:namePart>
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<mods:namePart id="DD04E7E7EA155EDC668739F8508DCB85">Kottelat, Maurice</mods:namePart>
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<mods:date id="D84CE431F1E45C93AC414EC03D73B69A">2010</mods:date>
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<treatment id="03D78797B34AFFD4FF3F40FAFEA24296" ID-DOI="http://doi.org/10.5281/zenodo.6200620" ID-GBIF-Taxon="119392733" ID-Zenodo-Dep="6200620" LSID="urn:lsid:plazi:treatment:03D78797B34AFFD4FF3F40FAFEA24296" httpUri="http://treatment.plazi.org/id/03D78797B34AFFD4FF3F40FAFEA24296" lastPageId="8" lastPageNumber="47" pageId="2" pageNumber="41">
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<paragraph id="8BC13681B34AFFDEFF3F40FAFC6845F9" blockId="2.[151,569,151,212]" box="[151,569,151,177]" pageId="2" pageNumber="41">
<heading id="D08981EDB34AFFDEFF3F40FAFC6845F9" bold="true" box="[151,569,151,177]" fontSize="11" level="1" pageId="2" pageNumber="41" reason="1">
<emphasis id="B90AEA93B34AFFDEFF3F40FAFC6845F9" bold="true" box="[151,569,151,177]" pageId="2" pageNumber="41">
<taxonomicName id="4C7E4D02B34AFFDEFF3F40FAFFCB45F9" ID-CoL="465VJ" box="[151,410,151,177]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="marang" status="sp. nov.">
<emphasis id="B90AEA93B34AFFDEFF3F40FAFFCB45F9" bold="true" box="[151,410,151,177]" italics="true" pageId="2" pageNumber="41">Nemacheilus marang</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A23957E8B34AFFDEFE0140FAFC6845F9" box="[425,569,151,177]" pageId="2" pageNumber="41" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC13681B34AFFDEFF3F40D7FF49459C" blockId="2.[151,569,151,212]" box="[151,280,186,212]" pageId="2" pageNumber="41">
(
<figureCitation id="13452A04B34AFFDEFF3740D7FF5E459C" box="[159,271,186,212]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="2.[151,255,1380,1404]" captionStartId-1="2.[151,255,2009,2033]" captionTargetBox-0="[222,1348,1075,1343]" captionTargetBox-1="[200,1386,1476,1971]" captionTargetId-0="figure@2.[202,1385,1050,1359]" captionTargetId-1="figure@2.[179,1409,1455,1987]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURE 2. Nemacheilus marang, MZB 13301, holotype, 38.9 mm SL; Indonesia: Kalimantan Timur: Sungai Marang." captionText-1="FIGURE 3. Nemacheilus marang, CMK 18908, paratype, 49.6 mm SL; Indonesia: Kalimantan Timur: Sungai Marang." httpUri-0="https://zenodo.org/record/196939/files/figure.png" httpUri-1="https://zenodo.org/record/196940/files/figure.png" pageId="2" pageNumber="41">Figs. 2–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C364650AB34AFFDEFF3F4168FF9146AC" pageId="2" pageNumber="41" type="materials_examined">
<paragraph id="8BC13681B34AFFDEFF3F4168FC2F4422" blockId="2.[151,1437,261,996]" pageId="2" pageNumber="41">
<emphasis id="B90AEA93B34AFFDEFF3F4168FF414457" bold="true" box="[151,272,261,287]" pageId="2" pageNumber="41">
<typeStatus id="54C58823B34AFFDEFF3F4168FF5D4457" box="[151,268,261,287]" pageId="2" pageNumber="41" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED6FAE44B34AFFDEFEB14168FF084457" box="[281,345,261,287]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="41">MZB</collectionCode>
13301, 
<quantity id="4C869B64B34AFFDEFE124168FC724457" box="[442,547,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="3.8899999999999997" pageId="2" pageNumber="41" unit="mm" value="38.9">38.9 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFDF54168FC984457" box="[605,713,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="4.9399999999999995" pageId="2" pageNumber="41" unit="mm" value="49.4">49.4 mm</quantity>
TL); 
<collectingCountry id="F3697611B34AFFDEFCA74168FDD74457" box="[783,902,261,287]" name="Indonesia" pageId="2" pageNumber="41">Indonesia</collectingCountry>
: Kalimantan Timur: Kabupaten Kutai Timur: Kecamatan Kelai: Tepian Langsat village, Sungai 
<taxonomicName id="4C7E4D02B34AFFDEFD494147FD6D440C" box="[737,828,298,324]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
(
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<geoCoordinate id="EE4A5046B34AFFDEFC624147FA0F440C" box="[970,1118,298,324]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="9" value="117.75916">117°45.55'E</geoCoordinate>
), Bengalon drainage; R. K. Hadiaty, Sugeng &amp; July, 
<date id="FFC01041B34AFFDEFE13413DFC2B4422" box="[443,634,336,362]" pageId="2" pageNumber="41" value="2004-08-25">25 August 2004</date>
.
</paragraph>
<paragraph id="8BC13681B34AFFDEFF6E4118FC444727" blockId="2.[151,1437,261,996]" pageId="2" pageNumber="41">
<emphasis id="B90AEA93B34AFFDEFF6E4118FF1A44C7" bold="true" box="[198,331,373,399]" pageId="2" pageNumber="41">
<typeStatus id="54C58823B34AFFDEFF6E4118FF1744C7" box="[198,326,373,399]" pageId="2" pageNumber="41" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED6FAE44B34AFFDEFEFE4118FFF044C7" LSID="urn:lsid:biocol.org:col:34784" box="[342,417,373,399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="2" pageNumber="41">ANSP</collectionCode>
187007, 1, 43.0 mm SL (
<quantity id="4C869B64B34AFFDEFD4D4118FD0244C7" box="[741,851,373,399]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" pageId="2" pageNumber="41" unit="mm" value="53.6">53.6 mm</quantity>
TL); 
<collectionCode id="ED6FAE44B34AFFDEFC364118FDA744C7" box="[926,1014,373,399]" pageId="2" pageNumber="41">BMNH</collectionCode>
2006.3.20.2, 2, 
<quantity id="4C869B64B34AFFDEFB684118FB3E44C7" box="[1216,1391,373,399]" metricMagnitude="-2" metricUnit="m" metricValue="3.96" metricValueMax="4.07" metricValueMin="3.85" pageId="2" pageNumber="41" unit="mm" value="39.6" valueMax="40.7" valueMin="38.5">38.5–40.7 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFF3741F7FF1A44FC" box="[159,331,410,436]" metricMagnitude="-2" metricUnit="m" metricValue="4.9799999999999995" metricValueMax="5.08" metricValueMin="4.88" pageId="2" pageNumber="41" unit="mm" value="49.8" valueMax="50.8" valueMin="48.8">48.8–50.8 mm</quantity>
TL); UF 165708, 2, 
<quantity id="4C869B64B34AFFDEFD9741F7FCBB44FC" box="[575,746,410,436]" metricMagnitude="-2" metricUnit="m" metricValue="4.17" metricValueMax="4.38" metricValueMin="3.96" pageId="2" pageNumber="41" unit="mm" value="41.7" valueMax="43.8" valueMin="39.6">39.6–43.8 mm</quantity>
SL (50.6–55.0 mm TL); 
<collectionCode id="ED6FAE44B34AFFDEFBBA41F7FA3644FC" LSID="urn:lsid:biocol.org:col:34871" box="[1042,1127,410,436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="41">USNM</collectionCode>
388745, 2, 
<quantity id="4C869B64B34AFFDEFB5A41F7FBCD44FC" box="[1266,1436,410,436]" metricMagnitude="-2" metricUnit="m" metricValue="3.9799999999999995" metricValueMax="4.09" metricValueMin="3.87" pageId="2" pageNumber="41" unit="mm" value="39.8" valueMax="40.9" valueMin="38.7">38.7–40.9 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFF6241ADFF2B4492" box="[202,378,448,474]" metricMagnitude="-2" metricUnit="m" metricValue="5.034999999999999" metricValueMax="5.18" metricValueMin="4.89" pageId="2" pageNumber="41" unit="mm" value="50.349999999999994" valueMax="51.8" valueMin="48.9">48.9–51.8 mm</quantity>
TL); 
<collectionCode id="ED6FAE44B34AFFDEFE6A41ADFFAD4492" box="[450,508,448,474]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="41">ZRC</collectionCode>
50729, 2, 
<quantity id="4C869B64B34AFFDEFDD641ADFD7A4492" box="[638,811,448,474]" metricMagnitude="-2" metricUnit="m" metricValue="3.565" metricValueMax="3.6399999999999997" metricValueMin="3.4899999999999998" pageId="2" pageNumber="41" unit="mm" value="35.65" valueMax="36.4" valueMin="34.9">34.9–36.4 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFCCF41ADFA464492" box="[871,1047,448,474]" metricMagnitude="-2" metricUnit="m" metricValue="4.529999999999999" metricValueMax="4.58" metricValueMin="4.4799999999999995" pageId="2" pageNumber="41" unit="mm" value="45.3" valueMax="45.8" valueMin="44.8">44.8–45.8 mm</quantity>
TL); CMK 20314, 2, 
<quantity id="4C869B64B34AFFDEFA8D41ADFE9444B7" metricMagnitude="-2" metricUnit="m" metricValue="4.215" metricValueMax="4.42" metricValueMin="4.01" pageId="2" pageNumber="41" unit="mm" value="42.15" valueMax="44.2" valueMin="40.1">40.1–44.2 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFF574188FFFC44B7" box="[255,429,485,511]" metricMagnitude="-2" metricUnit="m" metricValue="5.3549999999999995" metricValueMax="5.62" metricValueMin="5.09" pageId="2" pageNumber="41" unit="mm" value="53.55" valueMax="56.2" valueMin="50.9">50.9–56.2 mm</quantity>
TL); CMK 18908, 2, 
<quantity id="4C869B64B34AFFDEFD1D4188FD3044B7" box="[693,865,485,511]" metricMagnitude="-2" metricUnit="m" metricValue="4.575" metricValueMax="4.96" metricValueMin="4.1899999999999995" pageId="2" pageNumber="41" unit="mm" value="45.75" valueMax="49.6" valueMin="41.9">41.9–49.6 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFC334188FA1844B7" box="[923,1097,485,511]" metricMagnitude="-2" metricUnit="m" metricValue="5.845" metricValueMax="6.26" metricValueMin="5.43" pageId="2" pageNumber="41" unit="mm" value="58.45" valueMax="62.6" valueMin="54.3">54.3–62.6 mm</quantity>
TL); same data as 
<typeStatus id="54C58823B34AFFDEFA854188FBC944B7" box="[1325,1432,485,511]" pageId="2" pageNumber="41" type="holotype">holotype</typeStatus>
. — 
<collectionCode id="ED6FAE44B34AFFDEFF154267FEAF476C" box="[189,254,522,548]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="41">MZB</collectionCode>
13321, 4, 
<quantity id="4C869B64B34AFFDEFE2A4267FC4D476C" box="[386,540,522,548]" metricMagnitude="-2" metricUnit="m" metricValue="4.43" metricValueMax="5.16" metricValueMin="3.7" pageId="2" pageNumber="41" unit="mm" value="44.3" valueMax="51.6" valueMin="37.0">37–51.6 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFDF04267FD5A476C" box="[600,779,522,548]" metricMagnitude="-2" metricUnit="m" metricValue="5.625" metricValueMax="6.57" metricValueMin="4.68" pageId="2" pageNumber="41" unit="mm" value="56.25" valueMax="65.7" valueMin="46.8">46.8–65.7 mm</quantity>
TL); 
<collectingCountry id="F3697611B34AFFDEFCFC4267FD9E476C" box="[852,975,522,548]" name="Indonesia" pageId="2" pageNumber="41">Indonesia</collectingCountry>
: Kalimantan Timur: Kabupaten Kutai Timur: Kecamatan Kelai: Tepian Langsat village, Sungai Jelai (
<geoCoordinate id="EE4A5046B34AFFDEFC07425DFA7F4702" box="[943,1070,560,586]" direction="north" orientation="latitude" pageId="2" pageNumber="41" precision="9" value="1.6766666">1°40.60'N</geoCoordinate>
<geoCoordinate id="EE4A5046B34AFFDEFB9E425DFA834702" box="[1078,1234,560,586]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="9" value="117.27917">117°16.75'E</geoCoordinate>
); R. K. Hadiaty, Sugeng &amp; July, 
<date id="FFC01041B34AFFDEFEFA4238FC414727" box="[338,528,597,623]" pageId="2" pageNumber="41" value="2004-08-26">26 August 2004</date>
.
</paragraph>
<paragraph id="8BC13681B34AFFDEFF6E4217FF9146AC" blockId="2.[151,1437,261,996]" pageId="2" pageNumber="41">
<emphasis id="B90AEA93B34AFFDEFF6E4217FC0E47DC" bold="true" box="[198,607,634,660]" pageId="2" pageNumber="41">
Additional material (non 
<typeStatus id="54C58823B34AFFDEFDA54217FC0347DC" box="[525,594,634,660]" pageId="2" pageNumber="41">types</typeStatus>
).
</emphasis>
<collectionCode id="ED6FAE44B34AFFDEFDC24217FCFB47DC" box="[618,682,634,660]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="41">MZB</collectionCode>
13295, 1, 
<quantity id="4C869B64B34AFFDEFC864217FDCB47DC" box="[814,922,634,660]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" pageId="2" pageNumber="41" unit="mm" value="41.5">41.5 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFC714217FA1447DC" box="[985,1093,634,660]" metricMagnitude="-2" metricUnit="m" metricValue="5.41" pageId="2" pageNumber="41" unit="mm" value="54.1">54.1 mm</quantity>
TL); 
<collectingCountry id="F3697611B34AFFDEFB274217FB5447DC" box="[1167,1285,634,660]" name="Indonesia" pageId="2" pageNumber="41">Indonesia</collectingCountry>
: Kalimantan Timur: Kabupaten Kutai Timur: Kecamatan Kelai: Tepian Langsat village, Sungai 
<taxonomicName id="4C7E4D02B34AFFDEFB0242CDFB5B47F2" box="[1194,1290,672,698]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
(
<geoCoordinate id="EE4A5046B34AFFDEFAB542CDFBCD47F2" box="[1309,1436,672,698]" direction="north" orientation="latitude" pageId="2" pageNumber="41" precision="9" value="1.7078333">1°42.47'N</geoCoordinate>
<geoCoordinate id="EE4A5046B34AFFDEFF3F42A8FF7A4797" box="[151,299,709,735]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="9" value="117.75916">117°45.55'E</geoCoordinate>
); Daeng, 
<date id="FFC01041B34AFFDEFE3242A8FC094797" box="[410,600,709,735]" pageId="2" pageNumber="41" value="2004-08-21">21 August 2004</date>
. — 
<collectionCode id="ED6FAE44B34AFFDEFD2042A8FC994797" box="[648,712,709,735]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="41">MZB</collectionCode>
13302, 1, 
<quantity id="4C869B64B34AFFDEFCEC42A8FDFD4797" box="[836,940,709,735]" metricMagnitude="-2" metricUnit="m" metricValue="3.96" pageId="2" pageNumber="41" unit="mm" value="39.6">39.6 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFC4C42A8FA1F4797" box="[996,1102,709,735]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" pageId="2" pageNumber="41" unit="mm" value="50.5">50.5 mm</quantity>
TL); 
<collectingCountry id="F3697611B34AFFDEFB3A42A8FB594797" box="[1170,1288,709,735]" name="Indonesia" pageId="2" pageNumber="41">Indonesia</collectingCountry>
: Kalimantan Timur: Kabupaten Kutai Timur: Kecamatan Kelai: Tepian Langsat village, Sungai 
<taxonomicName id="4C7E4D02B34AFFDEFB024287FB5B464C" box="[1194,1290,746,772]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
(
<geoCoordinate id="EE4A5046B34AFFDEFAB54287FBCD464C" box="[1309,1436,746,772]" direction="north" orientation="latitude" pageId="2" pageNumber="41" precision="9" value="1.7078333">1°42.47'N</geoCoordinate>
<geoCoordinate id="EE4A5046B34AFFDEFF3F437DFF7A4662" box="[151,299,784,810]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="9" value="117.75916">117°45.55'E</geoCoordinate>
); Daeng, 
<date id="FFC01041B34AFFDEFE34437DFC0D4662" box="[412,604,784,810]" pageId="2" pageNumber="41" value="2004-08-23">23 August 2004</date>
. — 
<collectionCode id="ED6FAE44B34AFFDEFD25437DFC9C4662" box="[653,717,784,810]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="41">MZB</collectionCode>
13304, 3, 
<quantity id="4C869B64B34AFFDEFCE3437DFDA74662" box="[843,1014,784,810]" metricMagnitude="-2" metricUnit="m" metricValue="4.375" metricValueMax="4.58" metricValueMin="4.17" pageId="2" pageNumber="41" unit="mm" value="43.75" valueMax="45.8" valueMin="41.7">41.7–45.8 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFB98437DFA8C4662" box="[1072,1245,784,810]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" metricValueMax="5.88" metricValueMin="5.32" pageId="2" pageNumber="41" unit="mm" value="56.0" valueMax="58.8" valueMin="53.2">53.2–58.8 mm</quantity>
TL); 
<collectingCountry id="F3697611B34AFFDEFA8A437DFBC94662" box="[1314,1432,784,810]" name="Indonesia" pageId="2" pageNumber="41">Indonesia</collectingCountry>
: Kalimantan Timur: Kabupaten Kutai Timur: Kecamatan Kelai: Tepian Langsat village, mouth of Sungai 
<taxonomicName id="4C7E4D02B34AFFDEFF3F4337FEA4463C" box="[151,245,858,884]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
(
<geoCoordinate id="EE4A5046B34AFFDEFEAF4337FFD2463C" box="[263,387,858,884]" direction="north" orientation="latitude" pageId="2" pageNumber="41" precision="9" value="1.6766666">1°40.60'N</geoCoordinate>
<geoCoordinate id="EE4A5046B34AFFDEFE244337FC76463C" box="[396,551,858,884]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="9" value="117.27917">117°16.75'E</geoCoordinate>
); R. K. Hadiaty, 
<collectingCountry id="F3697611B34AFFDEFD5F4337FD10463C" box="[759,833,858,884]" name="Oman" pageId="2" pageNumber="41">Oman</collectingCountry>
&amp; Sugeng, 
<date id="FFC01041B34AFFDEFC7D4337FACC463C" box="[981,1181,858,884]" pageId="2" pageNumber="41" value="2004-08-24">24 August 2004</date>
. — 
<collectionCode id="ED6FAE44B34AFFDEFB784337FB40463C" box="[1232,1297,858,884]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="41">MZB</collectionCode>
13306, 10, 
<quantity id="4C869B64B34AFFDEFF3F43EDFF1546D2" box="[151,324,896,922]" metricMagnitude="-2" metricUnit="m" metricValue="4.61" metricValueMax="5.47" metricValueMin="3.75" pageId="2" pageNumber="41" unit="mm" value="46.1" valueMax="54.7" valueMin="37.5">37.5–54.7 mm</quantity>
SL (
<quantity id="4C869B64B34AFFDEFE2843EDFC7E46D2" box="[384,559,896,922]" metricMagnitude="-2" metricUnit="m" metricValue="5.865" metricValueMax="6.92" metricValueMin="4.81" pageId="2" pageNumber="41" unit="mm" value="58.65" valueMax="69.2" valueMin="48.1">48.1–69.2 mm</quantity>
TL); 
<collectingCountry id="F3697611B34AFFDEFDDF43EDFCA146D2" box="[631,752,896,922]" name="Indonesia" pageId="2" pageNumber="41">Indonesia</collectingCountry>
: Kalimantan Timur: Kabupaten Kutai Timur: Kecamatan Kelai: Tepian Langsat village, mouth of Sungai 
<taxonomicName id="4C7E4D02B34AFFDEFD7543C8FD6B46F7" box="[733,826,933,959]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
(
<geoCoordinate id="EE4A5046B34AFFDEFCE443C8FD9646F7" box="[844,967,933,959]" direction="north" orientation="latitude" pageId="2" pageNumber="41" precision="9" value="1.6766666">1°40.60'N</geoCoordinate>
<geoCoordinate id="EE4A5046B34AFFDEFC6743C8FA3746F7" box="[975,1126,933,959]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="9" value="117.27917">117°16.75'E</geoCoordinate>
); R. K. Hadiaty, 
<collectingCountry id="F3697611B34AFFDEFA9C43C8FB2C46F7" box="[1332,1405,933,959]" name="Oman" pageId="2" pageNumber="41">Oman</collectingCountry>
&amp; Sugeng, 
<date id="FFC01041B34AFFDEFF5543A7FFEC46AC" box="[253,445,970,996]" pageId="2" pageNumber="41" value="2004-08-25">25 August 2004</date>
.
</paragraph>
</subSubSection>
<caption id="DF016609B34AFFDEFF3F4509FBC04034" box="[151,1425,1380,1404]" httpUri="https://zenodo.org/record/196939/files/figure.png" pageId="2" pageNumber="41" targetBox="[222,1348,1075,1343]" targetPageId="2">
<paragraph id="8BC13681B34AFFDEFF3F4509FBC04034" blockId="2.[151,1425,1380,1404]" box="[151,1425,1380,1404]" pageId="2" pageNumber="41">
<emphasis id="B90AEA93B34AFFDEFF3F4509FF484034" bold="true" box="[151,281,1380,1404]" pageId="2" pageNumber="41">FIGURE 2.</emphasis>
<taxonomicName id="4C7E4D02B34AFFDEFE884508FC524034" box="[288,515,1381,1404]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34AFFDEFE884508FC524034" box="[288,515,1381,1404]" italics="true" pageId="2" pageNumber="41">Nemacheilus marang</emphasis>
</taxonomicName>
, MZB 13301, holotype, 38.9 mm SL; Indonesia: Kalimantan Timur: Sungai 
<taxonomicName id="4C7E4D02B34AFFDEFA9F4508FBDF4034" box="[1335,1422,1381,1404]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF016609B34AFFDEFF3F47B4FBC242B8" box="[151,1427,2009,2033]" httpUri="https://zenodo.org/record/196940/files/figure.png" pageId="2" pageNumber="41" targetBox="[200,1386,1476,1971]" targetPageId="2">
<paragraph id="8BC13681B34AFFDEFF3F47B4FBC242B8" blockId="2.[151,1427,2009,2033]" box="[151,1427,2009,2033]" pageId="2" pageNumber="41">
<emphasis id="B90AEA93B34AFFDEFF3F47B4FF4842B9" bold="true" box="[151,281,2009,2033]" pageId="2" pageNumber="41">FIGURE 3.</emphasis>
<taxonomicName id="4C7E4D02B34AFFDEFE8847B7FC5242B9" box="[288,515,2010,2033]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34AFFDEFE8847B7FC5242B9" box="[288,515,2010,2033]" italics="true" pageId="2" pageNumber="41">Nemacheilus marang</emphasis>
</taxonomicName>
, CMK 18908, paratype, 49.6 mm SL; Indonesia: Kalimantan Timur: Sungai 
<taxonomicName id="4C7E4D02B34AFFDEFA9147B4FBDE42B8" box="[1337,1423,2009,2032]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C364650AB34BFFDFFF6E40FAFDB2461A" pageId="3" pageNumber="42" type="diagnosis">
<paragraph id="8BC13681B34BFFDFFF6E40FAFF6644FF" blockId="3.[151,1437,151,999]" pageId="3" pageNumber="42">
<emphasis id="B90AEA93B34BFFDFFF6E40FAFF1745F9" bold="true" box="[198,326,151,177]" pageId="3" pageNumber="42">Diagnosis.</emphasis>
<taxonomicName id="4C7E4D02B34BFFDFFEFA40F5FC0145F9" box="[338,592,152,177]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34BFFDFFEFA40F5FC0145F9" box="[338,592,152,177]" italics="true" pageId="3" pageNumber="42">Nemacheilus marang</emphasis>
</taxonomicName>
is distinguished from the other species of 
<taxonomicName id="4C7E4D02B34BFFDFFBCC40F5FAAD45F9" box="[1124,1276,152,177]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B90AEA93B34BFFDFFBCC40F5FAAD45F9" box="[1124,1276,152,177]" italics="true" pageId="3" pageNumber="42">Nemacheilus</emphasis>
</taxonomicName>
known from Borneo by its unique colour pattern, which is made of 10–18 dark brown bars on the flank, extending from the dorsal midline to just below the lateral line, continuous across the back, not or only slightly wider on the back and where crossing the midlateral stripe; the bars usually similarly-shaped, straight, and of width less than half that of interspaces. In the other Bornean species (except 
<taxonomicName id="4C7E4D02B34BFFDFFCD84140FA53440E" box="[880,1026,301,326]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B34BFFDFFCD84140FA53440E" box="[880,1026,301,326]" italics="true" pageId="3" pageNumber="42">N. olivaceus</emphasis>
</taxonomicName>
) the bars are even less regular (
<taxonomicName id="4C7E4D02B34BFFDFFA294140FF7F4423" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B34BFFDFFA294140FF7F4423" italics="true" pageId="3" pageNumber="42">N. saravacensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34BFFDFFE95413FFC414423" box="[317,528,338,363]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B34BFFDFFE95413FFC414423" box="[317,528,338,363]" italics="true" pageId="3" pageNumber="42">N. longipectoralis</emphasis>
</taxonomicName>
), or dissociated into a midlateral series of blotches and a mid-dorsal series of saddles (
<taxonomicName id="4C7E4D02B34BFFDFFEA84115FFE544D9" box="[256,436,376,401]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="kapuasensis">
<emphasis id="B90AEA93B34BFFDFFEA84115FFE544D9" box="[256,436,376,401]" italics="true" pageId="3" pageNumber="42">N. kapuasensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34BFFDFFE6B4115FCC744D9" box="[451,662,376,401]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="elegantissimus">
<emphasis id="B90AEA93B34BFFDFFE6B4115FCC744D9" box="[451,662,376,401]" italics="true" pageId="3" pageNumber="42">N. elegantissimus</emphasis>
</taxonomicName>
); see Remarks for more information on the colour pattern of the other species.
</paragraph>
<paragraph id="8BC13681B34BFFDFFF6E41AFFDB2461A" blockId="3.[151,1437,151,999]" pageId="3" pageNumber="42">
Other diagnostic characters (none unique to 
<taxonomicName id="4C7E4D02B34BFFDFFD6441AFFD1B4493" box="[716,842,450,475]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34BFFDFFD6441AFFD1B4493" box="[716,842,450,475]" italics="true" pageId="3" pageNumber="42">N. marang</emphasis>
</taxonomicName>
) are: scales of the row above and below the lateral line on caudal peduncle not elongate and without a tubercle at their posterior extremity (vs. scales elongate and with a tubercle in 
<taxonomicName id="4C7E4D02B34BFFDFFE094260FC69476E" box="[417,568,525,550]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="spiniferus">
<emphasis id="B90AEA93B34BFFDFFE094260FC69476E" box="[417,568,525,550]" italics="true" pageId="3" pageNumber="42">N. spiniferus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34BFFDFFDEF4260FD50476E" box="[583,769,525,550]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B34BFFDFFDEF4260FD50476E" box="[583,769,525,550]" italics="true" pageId="3" pageNumber="42">N. selangoricus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7E4D02B34BFFDFFC944260FDC0476E" box="[828,913,525,550]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="tebo">
<emphasis id="B90AEA93B34BFFDFFC944260FDC0476E" box="[828,913,525,550]" italics="true" pageId="3" pageNumber="42">N. tebo</emphasis>
</taxonomicName>
); pectoral fin reaching only slightly beyond half of distance to pelvic-fin base (vs. to pelvic-fin origin or beyond in 
<taxonomicName id="4C7E4D02B34BFFDFFC76425FFAE04703" box="[990,1201,562,587]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B34BFFDFFC76425FFAE04703" box="[990,1201,562,587]" italics="true" pageId="3" pageNumber="42">N. longipectoralis</emphasis>
</taxonomicName>
); caudal fin forked, its upper lobe 1.0–1.5 times longer than its median rays (vs. deeply emarginate, the upper lobe 1.1–1.3 times longer than the median rays in 
<taxonomicName id="4C7E4D02B34BFFDFFDAD4210FCC647DE" box="[517,663,637,662]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B34BFFDFFDAD4210FCC647DE" box="[517,663,637,662]" italics="true" pageId="3" pageNumber="42">N. olivaceus</emphasis>
</taxonomicName>
); anus located behind middle of distance between pelvic-fin base and anal-fin origin (vs. about at midlength in 
<taxonomicName id="4C7E4D02B34BFFDFFD1B42CFFD1447F3" box="[691,837,674,699]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B34BFFDFFD1B42CFFD1447F3" box="[691,837,674,699]" italics="true" pageId="3" pageNumber="42">N. olivaceus</emphasis>
</taxonomicName>
); anterior nostril at the front side of a pointed flap (vs. at the extremity of an obliquely cut tube in 
<taxonomicName id="4C7E4D02B34BFFDFFD7D42A5FDC247A9" box="[725,915,712,737]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B34BFFDFFD7D42A5FDC247A9" box="[725,915,712,737]" italics="true" pageId="3" pageNumber="42">N. saravacensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34BFFDFFC0A42A5FA2B47A9" box="[930,1146,712,737]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B34BFFDFFC0A42A5FA2B47A9" box="[930,1146,712,737]" italics="true" pageId="3" pageNumber="42">N. longipectoralis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34BFFDFFB2242A5FB1747A9" box="[1162,1350,712,737]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B34BFFDFFB2242A5FB1747A9" box="[1162,1350,712,737]" italics="true" pageId="3" pageNumber="42">N. selangoricus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7E4D02B34BFFDFFA2942A5FF5A464E" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="spiniferus">
<emphasis id="B90AEA93B34BFFDFFA2942A5FF5A464E" italics="true" pageId="3" pageNumber="42">N. spiniferus</emphasis>
</taxonomicName>
); 17 branched caudal-fin rays (vs. 
<quantity id="4C869B64B34BFFDFFD034280FCB6464F" box="[683,743,749,775]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="3" pageNumber="42" unit="in" value="16.0">16 in</quantity>
most specimens of 
<taxonomicName id="4C7E4D02B34BFFDFFC7C4280FADE464E" box="[980,1167,749,774]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B34BFFDFFC7C4280FADE464E" box="[980,1167,749,774]" italics="true" pageId="3" pageNumber="42">N. saravacensis</emphasis>
</taxonomicName>
); 8½ branched dorsalfin rays (vs. 9–10½ in 
<taxonomicName id="4C7E4D02B34BFFDFFE06437FFC214663" box="[430,624,786,811]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B34BFFDFFE06437FFC214663" box="[430,624,786,811]" italics="true" pageId="3" pageNumber="42">N. saravacensis</emphasis>
</taxonomicName>
, 9½ in 
<taxonomicName id="4C7E4D02B34BFFDFFD66437FFDFD4663" box="[718,940,786,811]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B34BFFDFFD66437FFDFD4663" box="[718,940,786,811]" italics="true" pageId="3" pageNumber="42">N. longipectoralis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34BFFDFFC14437FFA224663" box="[956,1139,786,811]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="kapuasensis">
<emphasis id="B90AEA93B34BFFDFFC14437FFA224663" box="[956,1139,786,811]" italics="true" pageId="3" pageNumber="42">N. kapuasensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34BFFDFFB2C437FFB124663" box="[1156,1347,786,811]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B34BFFDFFB2C437FFB124663" box="[1156,1347,786,811]" italics="true" pageId="3" pageNumber="42">N. selangoricus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7E4D02B34BFFDFFA28437FFF5A4619" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="spiniferus">
<emphasis id="B90AEA93B34BFFDFFA28437FFF5A4619" italics="true" pageId="3" pageNumber="42">N. spiniferus</emphasis>
</taxonomicName>
); and 84–92 lateral-line pores (vs. 
<quantity id="4C869B64B34BFFDFFD0D4355FD4A461A" box="[677,795,824,850]" metricMagnitude="0" metricUnit="m" metricValue="2.4765" metricValueMax="2.5907999999999998" metricValueMin="2.3622" pageId="3" pageNumber="42" unit="in" value="97.5" valueMax="102.0" valueMin="93.0">93–102 in</quantity>
<taxonomicName id="4C7E4D02B34BFFDFFC8B4355FD834619" box="[803,978,824,849]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="kapuasensis">
<emphasis id="B90AEA93B34BFFDFFC8B4355FD834619" box="[803,978,824,849]" italics="true" pageId="3" pageNumber="42">N. kapuasensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C364650AB34BFFDAFF6E4330FB51471D" lastPageId="6" lastPageNumber="45" pageId="3" pageNumber="42" type="description">
<paragraph id="8BC13681B34BFFDFFF6E4330FC8746AF" blockId="3.[151,1437,151,999]" pageId="3" pageNumber="42">
<emphasis id="B90AEA93B34BFFDFFF6E4330FF0E463F" bold="true" box="[198,351,861,887]" pageId="3" pageNumber="42">Description.</emphasis>
See 
<figureCitation id="13452A04B34BFFDFFE314330FC77463F" box="[409,550,861,887]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="2.[151,255,1380,1404]" captionStartId-1="2.[151,255,2009,2033]" captionTargetBox-0="[222,1348,1075,1343]" captionTargetBox-1="[200,1386,1476,1971]" captionTargetId-0="figure@2.[202,1385,1050,1359]" captionTargetId-1="figure@2.[179,1409,1455,1987]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="FIGURE 2. Nemacheilus marang, MZB 13301, holotype, 38.9 mm SL; Indonesia: Kalimantan Timur: Sungai Marang." captionText-1="FIGURE 3. Nemacheilus marang, CMK 18908, paratype, 49.6 mm SL; Indonesia: Kalimantan Timur: Sungai Marang." httpUri-0="https://zenodo.org/record/196939/files/figure.png" httpUri-1="https://zenodo.org/record/196940/files/figure.png" pageId="3" pageNumber="42">Figures 2–3</figureCitation>
for general appearance and 
<tableCitation id="C6FC033AB34BFFDFFCDE4330FD9F463F" box="[886,974,861,887]" captionStart="TABLE 1" captionStartId="3.[151,241,1047,1071]" captionText="TABLE 1. Morphometric data of holotype, 17 paratypes and 12 other specimens of Nemacheilus marang. Mean and" pageId="3" pageNumber="42">Table 1</tableCitation>
for morphometric data of 
<typeStatus id="54C58823B34BFFDFFAA04330FB23463F" box="[1288,1394,861,887]" pageId="3" pageNumber="42" type="holotype">holotype</typeStatus>
, 17 
<typeStatus id="54C58823B34BFFDFFF3F43EFFF5646D4" box="[151,263,898,924]" pageId="3" pageNumber="42" type="paratype">paratypes</typeStatus>
and 12 non-type specimens. Body elongate, depth slowly increasing up to dorsal-fin origin. Behind dorsal fin, body depth decreasing slowly to caudal-fin base. Head slightly depressed; body slightly compressed anteriorly to compressed posteriorly.
</paragraph>
<caption id="DF016609B34BFFDFFF3F447AFBCD4167" box="[151,1436,1047,1071]" pageId="3" pageNumber="42">
<paragraph id="8BC13681B34BFFDFFF3F447AFBCD4167" blockId="3.[151,1436,1047,2007]" box="[151,1436,1047,1071]" pageId="3" pageNumber="42">
<emphasis id="B90AEA93B34BFFDFFF3F447AFF5D4167" bold="true" box="[151,268,1047,1071]" pageId="3" pageNumber="42">TABLE 1.</emphasis>
Morphometric data of holotype, 17 paratypes and 12 other specimens of 
<taxonomicName id="4C7E4D02B34BFFDFFB904475FB704167" box="[1080,1313,1048,1071]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34BFFDFFB904475FB704167" box="[1080,1313,1048,1071]" italics="true" pageId="3" pageNumber="42">Nemacheilus marang</emphasis>
</taxonomicName>
. Mean and
</paragraph>
</caption>
<paragraph id="8BC13681B34BFFDFFF3F4455FCF84107" blockId="3.[151,1436,1047,2007]" box="[151,681,1080,1103]" pageId="3" pageNumber="42">
standard deviation (s.d.) include data of 
<typeStatus id="54C58823B34BFFDFFDEC4455FCF44107" box="[580,677,1080,1103]" pageId="3" pageNumber="42" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BC13681B34BFFDFFD5C4407FB1041C9" blockId="3.[151,1436,1047,2007]" box="[756,1345,1130,1153]" pageId="3" pageNumber="42">
<typeStatus id="54C58823B34BFFDFFD5C4407FD0041C9" box="[756,849,1130,1153]" pageId="3" pageNumber="42" type="holotype">holotype</typeStatus>
range mean s.d.
</paragraph>
<paragraph id="8BC13681B34BFFDFFF3744F8FA5541E4" blockId="3.[151,1436,1047,2007]" box="[159,1028,1173,1196]" pageId="3" pageNumber="42">Standard length 38.9 34.9–54.1</paragraph>
<paragraph id="8BC13681B34BFFDFFF3744ADFA55419F" blockId="3.[151,1436,1047,2007]" box="[159,1028,1216,1239]" pageId="3" pageNumber="42">Total length 49.4 44.8–69.6</paragraph>
<paragraph id="8BC13681B34BFFDFFF374487FF804049" blockId="3.[151,1436,1047,2007]" box="[159,465,1258,1281]" pageId="3" pageNumber="42">In percent of standard length</paragraph>
<paragraph id="8BC13681B34BFFDFFF044578FB6F42E4" blockId="3.[151,1436,1047,2007]" pageId="3" pageNumber="42">Total length 127.0 123.6–130.5 126.3 1.6 Lateral length of head 23.9 22.0–26.5 23.9 1.1 Dorsal length of head 18.8 18.8–22.1 20.2 0.9 Predorsal length 48.9 46.7–52.5 48.9 1.3 Prepelvic length 51.0 49.3–54.3 51.0 1.5 Preanal length 76.6 75.1–79.8 77.0 1.3 Preanus length 67.6 63.3–72.4 68.5 2.4 Head depth at eye 9.5 8.3–12.2 10.2 1.0 Head depth at nape 12.6 11.7–14.2 12.9 0.7 Body depth at dorsal-fin origin 14.9 14.0–20.7 16.9 1.7 Depth of caudal peduncle 11.3 10.7–13.0 11.5 0.5 Length of caudal peduncle 15.2 13.5–16.0 15.0 0.7 Snout length 9.0 8.0–10.8 9.1 0.9 Head width at nares 8.2 7.3–9.1 7.3 0.5 Maximal headwidth 12.6 12.0–15.0 13.4 0.8 Body depth at dorsal-fin origin 9.8 9.7–15.7 11.8 1.7</paragraph>
<paragraph id="8BC13681B34BFFD8FB6947ADFD124354" blockId="3.[151,1436,1047,2007]" lastBlockId="4.[151,1437,1314,2012]" lastPageId="4" lastPageNumber="43" pageId="3" pageNumber="42">
<emphasis id="B90AEA93B34BFFDFFB6947ADFBC9429F" box="[1217,1432,1984,2007]" italics="true" pageId="3" pageNumber="42">continued next page</emphasis>
Dorsal fin with 4 simple and 8½ branched rays. Anal fin with 3 simple and 5½ branched rays. Pectoral fin with 11–13 rays, reaching ½ to 2/3 of distance to pelvic-fin base. Pelvic fin with 8 rays, origin below base of branched dorsal-fin ray 1 or 2, reaching about midway of distance to anal-fin origin; just reaching anus, which is about 1.5–2 eye diameter in front of anal fin. Axillary pelvic lobe present. Caudal fin with 8+9 branched rays, forked, lobes subequal, upper lobe 1.0–1.5 times longer than median rays. Caudal peduncle 1.1–1.5 times longer than deep, with low dorsal and ventral adipose crests on posterior half. Distal margin of dorsal fin straight. Largest recorded size 
<quantity id="4C869B64B34CFFD8FE56466FFC364354" box="[510,615,1538,1564]" metricMagnitude="-2" metricUnit="m" metricValue="5.47" pageId="4" pageNumber="43" unit="mm" value="54.7">54.7 mm</quantity>
SL (MZB 13306).
</paragraph>
<paragraph id="8BC13681B34CFFD8FF3740F2FB6F41A9" pageId="4" pageNumber="43">
<table id="F97EC421B34C0023FF3740F2FB1041A9" box="[159,1345,159,1249]" gridcols="5" gridrows="26" pageId="4" pageNumber="43">
<tr id="354E34C3B34C0023FF3740F2FB1045FF" box="[159,1345,159,183]" gridrow="0" pageId="4" pageNumber="43" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="769F5DBFB34C0023FF3740F2FC5A45FF" box="[159,523,159,183]" gridcol="0" gridrow="0" pageId="4" pageNumber="43">
<emphasis id="B90AEA93B34CFFD8FF3740F2FF4145FF" bold="true" box="[159,272,159,183]" pageId="4" pageNumber="43">TABLE 1.</emphasis>
(continued)
</th>
</tr>
<tr id="354E34C3B34C0023FF3740A7FB1045A9" box="[159,1345,202,225]" gridrow="1" pageId="4" pageNumber="43" rowspan-0="1">
<td id="769F5DBFB34C0023FD5C40A7FD0045A9" box="[756,849,202,225]" gridcol="1" gridrow="1" pageId="4" pageNumber="43">holotype</td>
<td id="769F5DBFB34C0023FC3240A7FA4F45A9" box="[922,1054,202,225]" gridcol="2" gridrow="1" pageId="4" pageNumber="43">range</td>
<td id="769F5DBFB34C0023FB2B40A7FAEC45A9" box="[1155,1213,202,225]" gridcol="3" gridrow="1" pageId="4" pageNumber="43">mean</td>
<td id="769F5DBFB34C0023FAB440A7FB1045A9" box="[1308,1345,202,225]" gridcol="4" gridrow="1" pageId="4" pageNumber="43">s.d.</td>
</tr>
<tr id="354E34C3B34C0023FF374098FB104444" box="[159,1345,245,268]" gridrow="2" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374098FC5A4444" box="[159,523,245,268]" gridcol="0" gridrow="2" pageId="4" pageNumber="43">Body depth at anal-fin origin</th>
<td id="769F5DBFB34C0023FD5C4098FD004444" box="[756,849,245,268]" gridcol="1" gridrow="2" pageId="4" pageNumber="43">6.2</td>
<td id="769F5DBFB34C0023FC324098FA4F4444" box="[922,1054,245,268]" gridcol="2" gridrow="2" pageId="4" pageNumber="43">5.7–8.1</td>
<td id="769F5DBFB34C0023FB2B4098FAEC4444" box="[1155,1213,245,268]" gridcol="3" gridrow="2" pageId="4" pageNumber="43">6.9</td>
<td id="769F5DBFB34C0023FAB44098FB104444" box="[1308,1345,245,268]" gridcol="4" gridrow="2" pageId="4" pageNumber="43">0.6</td>
</tr>
<tr id="354E34C3B34C0023FF37414DFB10447F" box="[159,1345,288,311]" gridrow="3" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF37414DFC5A447F" box="[159,523,288,311]" gridcol="0" gridrow="3" pageId="4" pageNumber="43">Eye diameter</th>
<td id="769F5DBFB34C0023FD5C414DFD00447F" box="[756,849,288,311]" gridcol="1" gridrow="3" pageId="4" pageNumber="43">6.2</td>
<td id="769F5DBFB34C0023FC32414DFA4F447F" box="[922,1054,288,311]" gridcol="2" gridrow="3" pageId="4" pageNumber="43">5.2–6.5</td>
<td id="769F5DBFB34C0023FB2B414DFAEC447F" box="[1155,1213,288,311]" gridcol="3" gridrow="3" pageId="4" pageNumber="43">5.8</td>
<td id="769F5DBFB34C0023FAB4414DFB10447F" box="[1308,1345,288,311]" gridcol="4" gridrow="3" pageId="4" pageNumber="43">0.3</td>
</tr>
<tr id="354E34C3B34C0023FF374127FB104429" box="[159,1345,330,353]" gridrow="4" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374127FC5A4429" box="[159,523,330,353]" gridcol="0" gridrow="4" pageId="4" pageNumber="43">Interorbital width</th>
<td id="769F5DBFB34C0023FD5C4127FD004429" box="[756,849,330,353]" gridcol="1" gridrow="4" pageId="4" pageNumber="43">5.9</td>
<td id="769F5DBFB34C0023FC324127FA4F4429" box="[922,1054,330,353]" gridcol="2" gridrow="4" pageId="4" pageNumber="43">5.4–7.6</td>
<td id="769F5DBFB34C0023FB2B4127FAEC4429" box="[1155,1213,330,353]" gridcol="3" gridrow="4" pageId="4" pageNumber="43">6.3</td>
<td id="769F5DBFB34C0023FAB44127FB104429" box="[1308,1345,330,353]" gridcol="4" gridrow="4" pageId="4" pageNumber="43">0.5</td>
</tr>
<tr id="354E34C3B34C0023FF374118FB1044C4" box="[159,1345,373,396]" gridrow="5" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374118FC5A44C4" box="[159,523,373,396]" gridcol="0" gridrow="5" pageId="4" pageNumber="43">Height of dorsal fin</th>
<td id="769F5DBFB34C0023FD5C4118FD0044C4" box="[756,849,373,396]" gridcol="1" gridrow="5" pageId="4" pageNumber="43">22.4</td>
<td id="769F5DBFB34C0023FC324118FA4F44C4" box="[922,1054,373,396]" gridcol="2" gridrow="5" pageId="4" pageNumber="43">18.1–24.6</td>
<td id="769F5DBFB34C0023FB2B4118FAEC44C4" box="[1155,1213,373,396]" gridcol="3" gridrow="5" pageId="4" pageNumber="43">22.8</td>
<td id="769F5DBFB34C0023FAB44118FB1044C4" box="[1308,1345,373,396]" gridcol="4" gridrow="5" pageId="4" pageNumber="43">1.3</td>
</tr>
<tr id="354E34C3B34C0023FF3741CDFB1044FF" box="[159,1345,416,439]" gridrow="6" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF3741CDFC5A44FF" box="[159,523,416,439]" gridcol="0" gridrow="6" pageId="4" pageNumber="43">Length of upper caudal-fin lobe</th>
<td id="769F5DBFB34C0023FD5C41CDFD0044FF" box="[756,849,416,439]" gridcol="1" gridrow="6" pageId="4" pageNumber="43">28.0</td>
<td id="769F5DBFB34C0023FC3241CDFA4F44FF" box="[922,1054,416,439]" gridcol="2" gridrow="6" pageId="4" pageNumber="43">24.2–30.9</td>
<td id="769F5DBFB34C0023FB2B41CDFAEC44FF" box="[1155,1213,416,439]" gridcol="3" gridrow="6" pageId="4" pageNumber="43">26.9</td>
<td id="769F5DBFB34C0023FAB441CDFB1044FF" box="[1308,1345,416,439]" gridcol="4" gridrow="6" pageId="4" pageNumber="43">1.5</td>
</tr>
<tr id="354E34C3B34C0023FF3741A7FB1044A9" box="[159,1345,458,481]" gridrow="7" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF3741A7FC5A44A9" box="[159,523,458,481]" gridcol="0" gridrow="7" pageId="4" pageNumber="43">Length of lower caudal-fin lobe</th>
<td id="769F5DBFB34C0023FD5C41A7FD0044A9" box="[756,849,458,481]" gridcol="1" gridrow="7" pageId="4" pageNumber="43">27.2</td>
<td id="769F5DBFB34C0023FC3241A7FA4F44A9" box="[922,1054,458,481]" gridcol="2" gridrow="7" pageId="4" pageNumber="43">22.6–29.3</td>
<td id="769F5DBFB34C0023FB2B41A7FAEC44A9" box="[1155,1213,458,481]" gridcol="3" gridrow="7" pageId="4" pageNumber="43">26.8</td>
<td id="769F5DBFB34C0023FAB441A7FB1044A9" box="[1308,1345,458,481]" gridcol="4" gridrow="7" pageId="4" pageNumber="43">1.7</td>
</tr>
<tr id="354E34C3B34C0023FF374198FB104744" box="[159,1345,501,524]" gridrow="8" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374198FC5A4744" box="[159,523,501,524]" gridcol="0" gridrow="8" pageId="4" pageNumber="43">Length of middle caudal-fin rays</th>
<td id="769F5DBFB34C0023FD5C4198FD004744" box="[756,849,501,524]" gridcol="1" gridrow="8" pageId="4" pageNumber="43">19.8</td>
<td id="769F5DBFB34C0023FC324198FA4F4744" box="[922,1054,501,524]" gridcol="2" gridrow="8" pageId="4" pageNumber="43">17.0–28.3</td>
<td id="769F5DBFB34C0023FB2B4198FAEC4744" box="[1155,1213,501,524]" gridcol="3" gridrow="8" pageId="4" pageNumber="43">19.7</td>
<td id="769F5DBFB34C0023FAB44198FB104744" box="[1308,1345,501,524]" gridcol="4" gridrow="8" pageId="4" pageNumber="43">2.4</td>
</tr>
<tr id="354E34C3B34C0023FF37424DFB10477F" box="[159,1345,544,567]" gridrow="9" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF37424DFC5A477F" box="[159,523,544,567]" gridcol="0" gridrow="9" pageId="4" pageNumber="43">Length of anal fin</th>
<td id="769F5DBFB34C0023FD5C424DFD00477F" box="[756,849,544,567]" gridcol="1" gridrow="9" pageId="4" pageNumber="43">17.5</td>
<td id="769F5DBFB34C0023FC32424DFA4F477F" box="[922,1054,544,567]" gridcol="2" gridrow="9" pageId="4" pageNumber="43">16.5–20.6</td>
<td id="769F5DBFB34C0023FB2B424DFAEC477F" box="[1155,1213,544,567]" gridcol="3" gridrow="9" pageId="4" pageNumber="43">18.3</td>
<td id="769F5DBFB34C0023FAB4424DFB10477F" box="[1308,1345,544,567]" gridcol="4" gridrow="9" pageId="4" pageNumber="43">0.9</td>
</tr>
<tr id="354E34C3B34C0023FF374227FB104729" box="[159,1345,586,609]" gridrow="10" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374227FC5A4729" box="[159,523,586,609]" gridcol="0" gridrow="10" pageId="4" pageNumber="43">Length of pelvic fin</th>
<td id="769F5DBFB34C0023FD5C4227FD004729" box="[756,849,586,609]" gridcol="1" gridrow="10" pageId="4" pageNumber="43">17.7</td>
<td id="769F5DBFB34C0023FC324227FA4F4729" box="[922,1054,586,609]" gridcol="2" gridrow="10" pageId="4" pageNumber="43">16.2–22.6</td>
<td id="769F5DBFB34C0023FB2B4227FAEC4729" box="[1155,1213,586,609]" gridcol="3" gridrow="10" pageId="4" pageNumber="43">17.7</td>
<td id="769F5DBFB34C0023FAB44227FB104729" box="[1308,1345,586,609]" gridcol="4" gridrow="10" pageId="4" pageNumber="43">1.2</td>
</tr>
<tr id="354E34C3B34C0023FF374218FB1047C4" box="[159,1345,629,652]" gridrow="11" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374218FC5A47C4" box="[159,523,629,652]" gridcol="0" gridrow="11" pageId="4" pageNumber="43">Length of pectoral fin</th>
<td id="769F5DBFB34C0023FD5C4218FD0047C4" box="[756,849,629,652]" gridcol="1" gridrow="11" pageId="4" pageNumber="43">19.0</td>
<td id="769F5DBFB34C0023FC324218FA4F47C4" box="[922,1054,629,652]" gridcol="2" gridrow="11" pageId="4" pageNumber="43">19.0–23.8</td>
<td id="769F5DBFB34C0023FB2B4218FAEC47C4" box="[1155,1213,629,652]" gridcol="3" gridrow="11" pageId="4" pageNumber="43">21.0</td>
<td id="769F5DBFB34C0023FAB44218FB1047C4" box="[1308,1345,629,652]" gridcol="4" gridrow="11" pageId="4" pageNumber="43">1.2</td>
</tr>
<tr id="354E34C3B34C0023FF3742CDFB1047FF" box="[159,1345,672,695]" gridrow="12" pageId="4" pageNumber="43" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="769F5DBFB34C0023FF3742CDFC5A47FF" box="[159,523,672,695]" gridcol="0" gridrow="12" pageId="4" pageNumber="43">In percent of dorsal head length</th>
</tr>
<tr id="354E34C3B34C0023FF3742A7FB1047A9" box="[159,1345,714,737]" gridrow="13" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF3742A7FC5A47A9" box="[159,523,714,737]" gridcol="0" gridrow="13" pageId="4" pageNumber="43">Lateral length of head</th>
<td id="769F5DBFB34C0023FD5C42A7FD0047A9" box="[756,849,714,737]" gridcol="1" gridrow="13" pageId="4" pageNumber="43">127.4</td>
<td id="769F5DBFB34C0023FC3242A7FA4F47A9" box="[922,1054,714,737]" gridcol="2" gridrow="13" pageId="4" pageNumber="43">111.5–129.7</td>
<td id="769F5DBFB34C0023FB2B42A7FAEC47A9" box="[1155,1213,714,737]" gridcol="3" gridrow="13" pageId="4" pageNumber="43">118.7</td>
<td id="769F5DBFB34C0023FAB442A7FB1047A9" box="[1308,1345,714,737]" gridcol="4" gridrow="13" pageId="4" pageNumber="43">4.3</td>
</tr>
<tr id="354E34C3B34C0023FF374298FB104644" box="[159,1345,757,780]" gridrow="14" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374298FC5A4644" box="[159,523,757,780]" gridcol="0" gridrow="14" pageId="4" pageNumber="43">Head depth at eye</th>
<td id="769F5DBFB34C0023FD5C4298FD004644" box="[756,849,757,780]" gridcol="1" gridrow="14" pageId="4" pageNumber="43">57.0</td>
<td id="769F5DBFB34C0023FC324298FA4F4644" box="[922,1054,757,780]" gridcol="2" gridrow="14" pageId="4" pageNumber="43">40.2–57.6</td>
<td id="769F5DBFB34C0023FB2B4298FAEC4644" box="[1155,1213,757,780]" gridcol="3" gridrow="14" pageId="4" pageNumber="43">50.8</td>
<td id="769F5DBFB34C0023FAB44298FB104644" box="[1308,1345,757,780]" gridcol="4" gridrow="14" pageId="4" pageNumber="43">3.8</td>
</tr>
<tr id="354E34C3B34C0023FF37434DFB10467F" box="[159,1345,800,823]" gridrow="15" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF37434DFC5A467F" box="[159,523,800,823]" gridcol="0" gridrow="15" pageId="4" pageNumber="43">Head depth at nape</th>
<td id="769F5DBFB34C0023FD5C434DFD00467F" box="[756,849,800,823]" gridcol="1" gridrow="15" pageId="4" pageNumber="43">67.1</td>
<td id="769F5DBFB34C0023FC32434DFA4F467F" box="[922,1054,800,823]" gridcol="2" gridrow="15" pageId="4" pageNumber="43">52.9–69.4</td>
<td id="769F5DBFB34C0023FB2B434DFAEC467F" box="[1155,1213,800,823]" gridcol="3" gridrow="15" pageId="4" pageNumber="43">63.9</td>
<td id="769F5DBFB34C0023FAB4434DFB10467F" box="[1308,1345,800,823]" gridcol="4" gridrow="15" pageId="4" pageNumber="43">3.8</td>
</tr>
<tr id="354E34C3B34C0023FF374327FB104629" box="[159,1345,842,865]" gridrow="16" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374327FC5A4629" box="[159,523,842,865]" gridcol="0" gridrow="16" pageId="4" pageNumber="43">Body depth at dorsal-fin origin</th>
<td id="769F5DBFB34C0023FD5C4327FD004629" box="[756,849,842,865]" gridcol="1" gridrow="16" pageId="4" pageNumber="43">79.5</td>
<td id="769F5DBFB34C0023FC324327FA4F4629" box="[922,1054,842,865]" gridcol="2" gridrow="16" pageId="4" pageNumber="43">63.5–97.2</td>
<td id="769F5DBFB34C0023FB2B4327FAEC4629" box="[1155,1213,842,865]" gridcol="3" gridrow="16" pageId="4" pageNumber="43">83.8</td>
<td id="769F5DBFB34C0023FAB44327FB104629" box="[1308,1345,842,865]" gridcol="4" gridrow="16" pageId="4" pageNumber="43">8.3</td>
</tr>
<tr id="354E34C3B34C0023FF374318FB1046C4" box="[159,1345,885,908]" gridrow="17" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374318FC5A46C4" box="[159,523,885,908]" gridcol="0" gridrow="17" pageId="4" pageNumber="43">Depth of caudal peduncle</th>
<td id="769F5DBFB34C0023FD5C4318FD0046C4" box="[756,849,885,908]" gridcol="1" gridrow="17" pageId="4" pageNumber="43">60.3</td>
<td id="769F5DBFB34C0023FC324318FA4F46C4" box="[922,1054,885,908]" gridcol="2" gridrow="17" pageId="4" pageNumber="43">51.8–63.1</td>
<td id="769F5DBFB34C0023FB2B4318FAEC46C4" box="[1155,1213,885,908]" gridcol="3" gridrow="17" pageId="4" pageNumber="43">57.3</td>
<td id="769F5DBFB34C0023FAB44318FB1046C4" box="[1308,1345,885,908]" gridcol="4" gridrow="17" pageId="4" pageNumber="43">2.6</td>
</tr>
<tr id="354E34C3B34C0023FF3743CDFB1046FF" box="[159,1345,928,951]" gridrow="18" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF3743CDFC5A46FF" box="[159,523,928,951]" gridcol="0" gridrow="18" pageId="4" pageNumber="43">Length of caudal peduncle</th>
<td id="769F5DBFB34C0023FD5C43CDFD0046FF" box="[756,849,928,951]" gridcol="1" gridrow="18" pageId="4" pageNumber="43">80.8</td>
<td id="769F5DBFB34C0023FC3243CDFA4F46FF" box="[922,1054,928,951]" gridcol="2" gridrow="18" pageId="4" pageNumber="43">63.0–83.8</td>
<td id="769F5DBFB34C0023FB2B43CDFAEC46FF" box="[1155,1213,928,951]" gridcol="3" gridrow="18" pageId="4" pageNumber="43">74.8</td>
<td id="769F5DBFB34C0023FAB443CDFB1046FF" box="[1308,1345,928,951]" gridcol="4" gridrow="18" pageId="4" pageNumber="43">5.4</td>
</tr>
<tr id="354E34C3B34C0023FF3743A7FB1046A9" box="[159,1345,970,993]" gridrow="19" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF3743A7FC5A46A9" box="[159,523,970,993]" gridcol="0" gridrow="19" pageId="4" pageNumber="43">Snout length</th>
<td id="769F5DBFB34C0023FD5C43A7FD0046A9" box="[756,849,970,993]" gridcol="1" gridrow="19" pageId="4" pageNumber="43">48.0</td>
<td id="769F5DBFB34C0023FC3243A7FA4F46A9" box="[922,1054,970,993]" gridcol="2" gridrow="19" pageId="4" pageNumber="43">39.5–54.9</td>
<td id="769F5DBFB34C0023FB2B43A7FAEC46A9" box="[1155,1213,970,993]" gridcol="3" gridrow="19" pageId="4" pageNumber="43">45.0</td>
<td id="769F5DBFB34C0023FAB443A7FB1046A9" box="[1308,1345,970,993]" gridcol="4" gridrow="19" pageId="4" pageNumber="43">3.5</td>
</tr>
<tr id="354E34C3B34C0023FF374398FB104144" box="[159,1345,1013,1036]" gridrow="20" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374398FC5A4144" box="[159,523,1013,1036]" gridcol="0" gridrow="20" pageId="4" pageNumber="43">Head width at nares</th>
<td id="769F5DBFB34C0023FD5C4398FD004144" box="[756,849,1013,1036]" gridcol="1" gridrow="20" pageId="4" pageNumber="43">45.2</td>
<td id="769F5DBFB34C0023FC324398FA4F4144" box="[922,1054,1013,1036]" gridcol="2" gridrow="20" pageId="4" pageNumber="43">37.0–45.9</td>
<td id="769F5DBFB34C0023FB2B4398FAEC4144" box="[1155,1213,1013,1036]" gridcol="3" gridrow="20" pageId="4" pageNumber="43">40.7</td>
<td id="769F5DBFB34C0023FAB44398FB104144" box="[1308,1345,1013,1036]" gridcol="4" gridrow="20" pageId="4" pageNumber="43">2.2</td>
</tr>
<tr id="354E34C3B34C0023FF37444DFB10417F" box="[159,1345,1056,1079]" gridrow="21" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF37444DFC5A417F" box="[159,523,1056,1079]" gridcol="0" gridrow="21" pageId="4" pageNumber="43">Maximal headwidth</th>
<td id="769F5DBFB34C0023FD5C444DFD00417F" box="[756,849,1056,1079]" gridcol="1" gridrow="21" pageId="4" pageNumber="43">67.1</td>
<td id="769F5DBFB34C0023FC32444DFA4F417F" box="[922,1054,1056,1079]" gridcol="2" gridrow="21" pageId="4" pageNumber="43">60.0–71.4</td>
<td id="769F5DBFB34C0023FB2B444DFAEC417F" box="[1155,1213,1056,1079]" gridcol="3" gridrow="21" pageId="4" pageNumber="43">65.0</td>
<td id="769F5DBFB34C0023FAB4444DFB10417F" box="[1308,1345,1056,1079]" gridcol="4" gridrow="21" pageId="4" pageNumber="43">2.9</td>
</tr>
<tr id="354E34C3B34C0023FF374427FB104129" box="[159,1345,1098,1121]" gridrow="22" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374427FC5A4129" box="[159,523,1098,1121]" gridcol="0" gridrow="22" pageId="4" pageNumber="43">Body width at dorsal-fin origin</th>
<td id="769F5DBFB34C0023FD5C4427FD004129" box="[756,849,1098,1121]" gridcol="1" gridrow="22" pageId="4" pageNumber="43">52.1</td>
<td id="769F5DBFB34C0023FC324427FA4F4129" box="[922,1054,1098,1121]" gridcol="2" gridrow="22" pageId="4" pageNumber="43">45.9–75.7</td>
<td id="769F5DBFB34C0023FB2B4427FAEC4129" box="[1155,1213,1098,1121]" gridcol="3" gridrow="22" pageId="4" pageNumber="43">58.7</td>
<td id="769F5DBFB34C0023FAB44427FB104129" box="[1308,1345,1098,1121]" gridcol="4" gridrow="22" pageId="4" pageNumber="43">7.6</td>
</tr>
<tr id="354E34C3B34C0023FF374418FB1041C4" box="[159,1345,1141,1164]" gridrow="23" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF374418FC5A41C4" box="[159,523,1141,1164]" gridcol="0" gridrow="23" pageId="4" pageNumber="43">Body width at anal-fin origin</th>
<td id="769F5DBFB34C0023FD5C4418FD0041C4" box="[756,849,1141,1164]" gridcol="1" gridrow="23" pageId="4" pageNumber="43">32.9</td>
<td id="769F5DBFB34C0023FC324418FA4F41C4" box="[922,1054,1141,1164]" gridcol="2" gridrow="23" pageId="4" pageNumber="43">30.0–39.1</td>
<td id="769F5DBFB34C0023FB2B4418FAEC41C4" box="[1155,1213,1141,1164]" gridcol="3" gridrow="23" pageId="4" pageNumber="43">34.1</td>
<td id="769F5DBFB34C0023FAB44418FB1041C4" box="[1308,1345,1141,1164]" gridcol="4" gridrow="23" pageId="4" pageNumber="43">2.7</td>
</tr>
<tr id="354E34C3B34C0023FF3744CDFB1041FF" box="[159,1345,1184,1207]" gridrow="24" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF3744CDFC5A41FF" box="[159,523,1184,1207]" gridcol="0" gridrow="24" pageId="4" pageNumber="43">Eye diameter</th>
<td id="769F5DBFB34C0023FD5C44CDFD0041FF" box="[756,849,1184,1207]" gridcol="1" gridrow="24" pageId="4" pageNumber="43">32.9</td>
<td id="769F5DBFB34C0023FC3244CDFA4F41FF" box="[922,1054,1184,1207]" gridcol="2" gridrow="24" pageId="4" pageNumber="43">25.0–32.9</td>
<td id="769F5DBFB34C0023FB2B44CDFAEC41FF" box="[1155,1213,1184,1207]" gridcol="3" gridrow="24" pageId="4" pageNumber="43">28.7</td>
<td id="769F5DBFB34C0023FAB444CDFB1041FF" box="[1308,1345,1184,1207]" gridcol="4" gridrow="24" pageId="4" pageNumber="43">2.1</td>
</tr>
<tr id="354E34C3B34C0023FF3744A7FB1041A9" box="[159,1345,1226,1249]" gridrow="25" pageId="4" pageNumber="43">
<th id="769F5DBFB34C0023FF3744A7FC5A41A9" box="[159,523,1226,1249]" gridcol="0" gridrow="25" pageId="4" pageNumber="43">Interorbital width</th>
<td id="769F5DBFB34C0023FD5C44A7FD0041A9" box="[756,849,1226,1249]" gridcol="1" gridrow="25" pageId="4" pageNumber="43">32.9</td>
<td id="769F5DBFB34C0023FC3244A7FA4F41A9" box="[922,1054,1226,1249]" gridcol="2" gridrow="25" pageId="4" pageNumber="43">25.0–32.9</td>
<td id="769F5DBFB34C0023FB2B44A7FAEC41A9" box="[1155,1213,1226,1249]" gridcol="3" gridrow="25" pageId="4" pageNumber="43">28.6</td>
<td id="769F5DBFB34C0023FAB444A7FB1041A9" box="[1308,1345,1226,1249]" gridcol="4" gridrow="25" pageId="4" pageNumber="43">2.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC13681B34CFFD8FF6E4645FCA643FA" blockId="4.[151,1437,1314,2012]" pageId="4" pageNumber="43">Body entirely covered by deeply embedded scales. No scales on belly in front of pelvic-fin base (one specimen with scales extending forward until almost between pectoral-fin base). Lateral line complete, straight, with 84–92 pores. Cephalic lateral line system with 5 supraorbital, 4+11–12 infraorbital, 9–10 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
<paragraph id="8BC13681B34CFFD8FF6E46D0FD6F4224" blockId="4.[151,1437,1314,2012]" pageId="4" pageNumber="43">
Anterior nostril pierced in front side of a pointed flap. Mouth gape about 2 times wider than long (
<figureCitation id="13452A04B34CFFD8FAC546D0FEF943B4" captionStart="FIGURE 4" captionStartId="5.[151,255,1033,1057]" captionTargetBox="[151,1434,530,1012]" captionTargetId="figure@5.[151,1436,530,1012]" captionTargetPageId="5" captionText="FIGURE 4. Nemacheilus marang; a, holotype, MZB 13301, 38.9 mm SL, head, lateral view; b, MZB 13306, 44.6 mm SL; mouth, ventral view." httpUri="https://zenodo.org/record/196941/files/figure.png" pageId="4" pageNumber="43">Fig. 4</figureCitation>
b). Lips thin but fleshy, lower lip in some specimens with a few shallow grooves in median area. No median incision in upper lip. No median interruption in lower lip. Processus dentiformis present. No median notch in lower jaw. Inner rostral barbel reaching below anterior half of eye; outer one reaching about vertical of middle of eye. Maxillary barbel reaching posterior margin of eye.
</paragraph>
<paragraph id="8BC13681B34CFFD8FF6E471AFD2142DA" blockId="4.[151,1437,1314,2012]" box="[198,880,1911,1938]" pageId="4" pageNumber="43">
<emphasis id="B90AEA93B34CFFD8FF6E471AFF9042D9" bold="true" box="[198,449,1911,1937]" pageId="4" pageNumber="43">Sexual dimorphism.</emphasis>
Males with suborbital flap (
<figureCitation id="13452A04B34CFFD8FCA44715FD0542DA" box="[780,852,1912,1938]" captionStart="FIGURE 4" captionStartId="5.[151,255,1033,1057]" captionTargetBox="[151,1434,530,1012]" captionTargetId="figure@5.[151,1436,530,1012]" captionTargetPageId="5" captionText="FIGURE 4. Nemacheilus marang; a, holotype, MZB 13301, 38.9 mm SL, head, lateral view; b, MZB 13306, 44.6 mm SL; mouth, ventral view." httpUri="https://zenodo.org/record/196941/files/figure.png" pageId="4" pageNumber="43">Fig. 4</figureCitation>
a).
</paragraph>
<paragraph id="8BC13681B34CFFD9FF6E47F0FD134494" blockId="4.[151,1437,1314,2012]" lastBlockId="5.[151,1437,152,476]" lastPageId="5" lastPageNumber="44" pageId="4" pageNumber="43">
<emphasis id="B90AEA93B34CFFD8FF6E47F0FF0442FF" bold="true" box="[198,341,1949,1975]" pageId="4" pageNumber="43">Coloration.</emphasis>
Body background colour pale yellowish brown, somewhat darker on back, paler on belly. A deep layer of pigment making a dark greyish axial stripe along lateral line. 10–18 dark brown bars on flank, extending from dorsal midline to just below mid-lateral stripe, continuous across back, not or only slightly wider on back and where crossing midlateral stripe. Bars usually all similarly shaped (
<figureCitation id="13452A04B34DFFD9FB2140D0FA9E459F" box="[1161,1231,189,215]" captionStart="FIGURE 3" captionStartId="2.[151,255,2009,2033]" captionTargetBox="[200,1386,1476,1971]" captionTargetId="figure@2.[179,1409,1455,1987]" captionTargetPageId="2" captionText="FIGURE 3. Nemacheilus marang, CMK 18908, paratype, 49.6 mm SL; Indonesia: Kalimantan Timur: Sungai Marang." httpUri="https://zenodo.org/record/196940/files/figure.png" pageId="5" pageNumber="44">Fig. 3</figureCitation>
shows one of the less regularly patterned specimen), straight, their width less than half that of interspaces. Bars on caudal peduncle (and sometimes on posterior half of body) usually dissociated into a small saddle on dorsal midline and a blotch on midlateral stripe. A conspicuous black blotch on upper part of lower half of caudal-fin base, slightly vertically elongated, not reaching ventral midline; a fainter spot at upper part of caudal-fin base, close to dorsal midline; space between these two marks pale yellow; another small pale yellow patch in front of lower spot. Top of head dark grey, a darker triangular blotch on nape, a darker band between eyes, a pair of longitudinal darker lines between nostrils and tip of snout.
</paragraph>
<caption id="DF016609B34DFFD9FF3F4464FFF14109" httpUri="https://zenodo.org/record/196941/files/figure.png" pageId="5" pageNumber="44" targetBox="[151,1434,530,1012]" targetPageId="5">
<paragraph id="8BC13681B34DFFD9FF3F4464FFF14109" blockId="5.[151,1436,1033,1089]" pageId="5" pageNumber="44">
<emphasis id="B90AEA93B34DFFD9FF3F4464FF4B4169" bold="true" box="[151,282,1033,1057]" pageId="5" pageNumber="44">FIGURE 4.</emphasis>
<taxonomicName id="4C7E4D02B34DFFD9FE894467FC544169" box="[289,517,1034,1057]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="44" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34DFFD9FE894467FC544169" box="[289,517,1034,1057]" italics="true" pageId="5" pageNumber="44">Nemacheilus marang</emphasis>
</taxonomicName>
; 
<emphasis id="B90AEA93B34DFFD9FDBC4464FC794169" bold="true" box="[532,552,1033,1057]" pageId="5" pageNumber="44">a,</emphasis>
holotype, MZB 13301, 38.9 mm SL, head, lateral view; 
<emphasis id="B90AEA93B34DFFD9FB264464FAF24169" bold="true" box="[1166,1187,1033,1057]" pageId="5" pageNumber="44">b,</emphasis>
MZB 13306, 44.6 mm SL; mouth, ventral view.
</paragraph>
</caption>
<caption id="DF016609B34DFFD9FF3F4585FE9E4357" httpUri="https://zenodo.org/record/196942/files/figure.png" pageId="5" pageNumber="44" targetBox="[168,1418,1140,1489]" targetPageId="5">
<paragraph id="8BC13681B34DFFD9FF3F4585FE9E4357" blockId="5.[151,1436,1512,1567]" pageId="5" pageNumber="44">
<emphasis id="B90AEA93B34DFFD9FF3F4585FF484348" bold="true" box="[151,281,1512,1536]" pageId="5" pageNumber="44">FIGURE 5.</emphasis>
<taxonomicName id="4C7E4D02B34DFFD9FE884584FC454348" box="[288,532,1513,1536]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="44" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B34DFFD9FE884584FC454348" box="[288,532,1513,1536]" italics="true" pageId="5" pageNumber="44">Nemacheilus olivaceus</emphasis>
</taxonomicName>
, ZRC 45453, 55.4 mm SL; Borneo: Sabah: Danum Valley (photograph by Tan Heok Hui).
</paragraph>
</caption>
<caption id="DF016609B34DFFD9FF3F47B4FB6942B8" box="[151,1336,2009,2033]" httpUri="https://zenodo.org/record/196943/files/figure.png" pageId="5" pageNumber="44" targetBox="[202,1387,1642,1955]" targetPageId="5">
<paragraph id="8BC13681B34DFFD9FF3F47B4FB6942B8" blockId="5.[151,1336,2009,2033]" box="[151,1336,2009,2033]" pageId="5" pageNumber="44">
<emphasis id="B90AEA93B34DFFD9FF3F47B4FF4B42B9" bold="true" box="[151,282,2009,2033]" pageId="5" pageNumber="44">FIGURE 6.</emphasis>
<taxonomicName id="4C7E4D02B34DFFD9FE8847B4FC6042B8" box="[288,561,2009,2032]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="44" phylum="Chordata" rank="species" species="kapuasensis">
<emphasis id="B90AEA93B34DFFD9FE8847B4FC6042B8" box="[288,561,2009,2032]" italics="true" pageId="5" pageNumber="44">Nemacheilus kapuasensis</emphasis>
</taxonomicName>
, CMK 3187, 49.7 mm SL; Borneo: Kalimantan Barat: Kapuas drainage.
</paragraph>
</caption>
<paragraph id="8BC13681B34EFFDAFF6E40F5FB224590" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">Dorsal fin hyaline, with two or three rows of faint brown spots on rays. Caudal fin hyaline with two or three irregular vertical rows of faint brown spots on median rays, other rays unmarked. Other fins hyaline.</paragraph>
<paragraph id="8BC13681B34EFFDAFF6E4089FDAF45B6" blockId="6.[151,1437,152,2008]" box="[198,1022,228,254]" pageId="6" pageNumber="45">In life: background pale greyish yellow, markings as above, dark grey.</paragraph>
<paragraph id="8BC13681B34EFFDAFF6E4167FC8E44DE" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">
<emphasis id="B90AEA93B34EFFDAFF6E4167FC6F446C" bold="true" box="[198,574,266,292]" pageId="6" pageNumber="45">Notes on habitat and biology.</emphasis>
At the 
<typeStatus id="54C58823B34EFFDAFD364167FC81446C" box="[670,720,266,292]" pageId="6" pageNumber="45">type</typeStatus>
locality, Sungai 
<taxonomicName id="4C7E4D02B34EFFDAFC364167FDA8446C" box="[926,1017,266,292]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
was about 
<quantity id="4C869B64B34EFFDAFB2F4167FA94446C" box="[1159,1221,266,292]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="45" unit="m" value="10.0">10 m</quantity>
wide (dry season conditions) and about 
<quantity id="4C869B64B34EFFDAFE37415DFFB94402" box="[415,488,304,330]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="45" unit="cm" value="50.0">50 cm</quantity>
deep with the bottom made of a smooth rock layer, without any relief, with fast current and clear water. 
<taxonomicName id="4C7E4D02B34EFFDAFE19413AFCFA4438" box="[433,683,343,368]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFE19413AFCFA4438" box="[433,683,343,368]" italics="true" pageId="6" pageNumber="45">Nemacheilus marang</emphasis>
</taxonomicName>
was collected in submerged debris and leaf litter along the edge but was totally absent in the middle of the stream.
</paragraph>
<paragraph id="8BC13681B34EFFDAFF6E41CFFA714741" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">
In Sungai Jelai, the bottom is made of large stones and mud. 
<taxonomicName id="4C7E4D02B34EFFDAFC2641CEFAD944F4" box="[910,1160,419,444]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFC2641CEFAD944F4" box="[910,1160,419,444]" italics="true" pageId="6" pageNumber="45">Nemacheilus marang</emphasis>
</taxonomicName>
was less abundant than at the 
<typeStatus id="54C58823B34EFFDAFF4941A4FF4544AB" box="[225,276,457,483]" pageId="6" pageNumber="45">type</typeStatus>
locality. It was obtained from submerged debris and leaf litter that accumulates around the large stones. The samples from the 
<taxonomicName id="4C7E4D02B34EFFDAFE5D4182FC014741" box="[501,592,495,521]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
area include only 15 other fish species.
</paragraph>
<paragraph id="8BC13681B34EFFDAFF6E4278FB51471D" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">
One female (MZB 13306, 
<quantity id="4C869B64B34EFFDAFDAC4278FC3F4767" box="[516,622,533,559]" metricMagnitude="-2" metricUnit="m" metricValue="5.21" pageId="6" pageNumber="45" unit="mm" value="52.1">52.1 mm</quantity>
SL) was dissected and its ovaries contained 3942 ripe eggs, 
<quantity id="4C869B64B34EFFDAFAEC4278FE94471D" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="6" pageNumber="45" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.4–0.6 mm</quantity>
diameter. The size of all eggs is almost similar, indicating that 
<taxonomicName id="4C7E4D02B34EFFDAFC014256FA76471C" box="[937,1063,571,596]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFC014256FA76471C" box="[937,1063,571,596]" italics="true" pageId="6" pageNumber="45">N. marang</emphasis>
</taxonomicName>
is a total spawner.
</paragraph>
</subSubSection>
<subSubSection id="C364650AB34EFFDAFF6E420CFF58478F" pageId="6" pageNumber="45" type="distribution">
<paragraph id="8BC13681B34EFFDAFF6E420CFF58478F" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">
<emphasis id="B90AEA93B34EFFDAFF6E420CFF364733" bold="true" box="[198,359,609,635]" pageId="6" pageNumber="45">Distribution.</emphasis>
<taxonomicName id="4C7E4D02B34EFFDAFED9420CFC3C4732" box="[369,621,609,634]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFED9420CFC3C4732" box="[369,621,609,634]" italics="true" pageId="6" pageNumber="45">Nemacheilus marang</emphasis>
</taxonomicName>
is presently known only from the 
<taxonomicName id="4C7E4D02B34EFFDAFBBC420CFA3E4733" box="[1044,1135,609,635]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
and Jelai Rivers in Jelai drainage, East Kalimantan (
<figureCitation id="13452A04B34EFFDAFE7442EAFC7047E9" box="[476,545,647,673]" captionStart="FIGURE 1" captionStartId="1.[151,255,1932,1956]" captionTargetBox="[313,1268,197,1905]" captionTargetId="figure@1.[313,1269,197,1927]" captionTargetPageId="1" captionText="FIGURE 1. Sangkulirang Peninsula, Kalimantan Timur, Borneo, showing location of Sungai (river) Marang, in Bengalon drainage (above) and collecting sites of Nemacheilus marang in Sungai Marang (below). 1, Tepian Langsat village (type locality); 2, mouth of Sungai Marang; and 3, Sungai Jelai." httpUri="https://zenodo.org/record/196938/files/figure.png" pageId="6" pageNumber="45">Fig. 1</figureCitation>
). The Jelai is a tributary of Sungai Bengalon, which flows into the Straits of Sulawesi.
</paragraph>
</subSubSection>
<subSubSection id="C364650AB34EFFDAFF6E42BEFA2947A6" box="[198,1144,723,750]" pageId="6" pageNumber="45" type="etymology">
<paragraph id="8BC13681B34EFFDAFF6E42BEFA2947A6" blockId="6.[151,1437,152,2008]" box="[198,1144,723,750]" pageId="6" pageNumber="45">
<emphasis id="B90AEA93B34EFFDAFF6E42BEFF0247A5" bold="true" box="[198,339,723,749]" pageId="6" pageNumber="45">Etymology.</emphasis>
Named after the 
<typeStatus id="54C58823B34EFFDAFDB642B9FC0147A6" box="[542,592,724,750]" pageId="6" pageNumber="45">type</typeStatus>
locality Sungai 
<taxonomicName id="4C7E4D02B34EFFDAFCB842B9FD3E47A6" box="[784,879,724,750]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="marang">Marang</taxonomicName>
. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C364650AB34EFFDAFF6E4294FBC543BB" pageId="6" pageNumber="45" type="discussion">
<paragraph id="8BC13681B34EFFDAFF6E4294FFC84195" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">
<emphasis id="B90AEA93B34EFFDAFF6E4294FF6E465B" bold="true" box="[198,319,761,787]" pageId="6" pageNumber="45">Remarks.</emphasis>
Seven species of the genus 
<taxonomicName id="4C7E4D02B34EFFDAFD0A4297FD6B465B" box="[674,826,762,787]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B90AEA93B34EFFDAFD0A4297FD6B465B" box="[674,826,762,787]" italics="true" pageId="6" pageNumber="45">Nemacheilus</emphasis>
</taxonomicName>
are known from Borneo (
<bibRefCitation id="EFEF4B70B34EFFDAFBD74297FB61465C" author="Kottelat" box="[1151,1328,762,788]" pageId="6" pageNumber="45" refString="Kottelat, M. (1984) Revision of the Indonesian and Malaysian loaches of the subfamily Noemacheilinae. Japanese Journal of Ichthyology, 31, 225 - 260." type="journal article" year="1984">Kottelat, 1984</bibRefCitation>
; 
<bibRefCitation id="EFEF4B70B34EFFDAFAE84297FF7C4672" author="Chin" pageId="6" pageNumber="45" refString="Chin, P. K. &amp; Samat, A. (1992) A new species of loach, Nemacheilus elegantissimus, (family Balitoridae, subfamily Nemacheilinae), from Danum Valley, Sabah, Malaysia. Malayan Nature Journal, 46, 25 - 33." type="journal article" year="1992">
<collectingCountry id="F3697611B34EFFDAFAE84297FB28465C" box="[1344,1401,762,788]" name="China" pageId="6" pageNumber="45">Chin</collectingCountry>
&amp; Samat, 1992
</bibRefCitation>
; 
<bibRefCitation id="EFEF4B70B34EFFDAFE92434DFC3D4672" author="Hadiaty" box="[314,620,800,826]" pageId="6" pageNumber="45" refString="Hadiaty, R. K. &amp; Kottelat, M. (2009) Nemacheilus tebo, a new nemacheiline loach from Sangkulirang Karst, East Kalimantan, Indonesia (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 57, 119 - 125." type="journal article" year="2009">Hadiaty &amp; Kottelat, 2009</bibRefCitation>
): 
<taxonomicName id="4C7E4D02B34EFFDAFD28434DFD1E4671" box="[640,847,800,825]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="elegantissimus">
<emphasis id="B90AEA93B34EFFDAFD28434DFD1E4671" box="[640,847,800,825]" italics="true" pageId="6" pageNumber="45">N. elegantissimus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34EFFDAFCF7434DFA5E4671" box="[863,1039,800,825]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="kapuasensis">
<emphasis id="B90AEA93B34EFFDAFCF7434DFA5E4671" box="[863,1039,800,825]" italics="true" pageId="6" pageNumber="45">N. kapuasensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34EFFDAFBB7434DFAA24671" box="[1055,1267,800,825]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B34EFFDAFBB7434DFAA24671" box="[1055,1267,800,825]" italics="true" pageId="6" pageNumber="45">N. longipectoralis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34EFFDAFAAB434DFBC44671" box="[1283,1429,800,825]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B34EFFDAFAAB434DFBC44671" box="[1283,1429,800,825]" italics="true" pageId="6" pageNumber="45">N. olivaceus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34EFFDAFF3F432BFF0F4617" box="[151,350,838,863]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B34EFFDAFF3F432BFF0F4617" box="[151,350,838,863]" italics="true" pageId="6" pageNumber="45">N. saravacensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34EFFDAFEC7432BFC414617" box="[367,528,838,863]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="spiniferus">
<emphasis id="B90AEA93B34EFFDAFEC7432BFC414617" box="[367,528,838,863]" italics="true" pageId="6" pageNumber="45">N. spiniferus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7E4D02B34EFFDAFDE7432BFCFA4617" box="[591,683,838,863]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="tebo">
<emphasis id="B90AEA93B34EFFDAFDE7432BFCFA4617" box="[591,683,838,863]" italics="true" pageId="6" pageNumber="45">N. tebo</emphasis>
</taxonomicName>
. Three additional species have been at some time placed in 
<taxonomicName id="4C7E4D02B34EFFDAFF3F4301FF6B46CD" box="[151,314,876,901]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B90AEA93B34EFFDAFF3F4301FF6B46CD" box="[151,314,876,901]" italics="true" pageId="6" pageNumber="45">Nemacheilus</emphasis>
</taxonomicName>
but are now in the genera 
<taxonomicName id="4C7E4D02B34EFFDAFD3A4301FD0846CD" box="[658,857,876,901]" class="Actinopterygii" family="Nemacheilidae" genus="Sundoreonectes" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B90AEA93B34EFFDAFD3A4301FD0846CD" box="[658,857,876,901]" italics="true" pageId="6" pageNumber="45">Sundoreonectes</emphasis>
</taxonomicName>
(
<taxonomicName id="4C7E4D02B34EFFDAFCC44301FDB846CD" box="[876,1001,876,901]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="obesus">
<emphasis id="B90AEA93B34EFFDAFCC44301FDB846CD" box="[876,1001,876,901]" italics="true" pageId="6" pageNumber="45">N. obesus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C7E4D02B34EFFDAFC524301FAFB46CD" box="[1018,1194,876,901]" family="Nemacheilidae" genus="Elxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="sabanus">
<emphasis id="B90AEA93B34EFFDAFC524301FAFB46CD" box="[1018,1194,876,901]" italics="true" pageId="6" pageNumber="45">Elxis sabanus</emphasis>
</taxonomicName>
) and 
<taxonomicName id="4C7E4D02B34EFFDAFB5C4301FB3D46CD" box="[1268,1388,876,901]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B90AEA93B34EFFDAFB5C4301FB3D46CD" box="[1268,1388,876,901]" italics="true" pageId="6" pageNumber="45">Schistura</emphasis>
</taxonomicName>
(
<taxonomicName id="4C7E4D02B34EFFDAFA284301FF4A46E3" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="maculiceps">
<emphasis id="B90AEA93B34EFFDAFA284301FF4A46E3" italics="true" pageId="6" pageNumber="45">N. maculiceps</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFEF4B70B34EFFDAFE9C43FFFFB046E4" author="Kottelat" box="[308,481,914,940]" pageId="6" pageNumber="45" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat, 1990</bibRefCitation>
; 
<bibRefCitation id="EFEF4B70B34EFFDAFE4643FFFC2546E4" author="Chin" box="[494,628,914,940]" pageId="6" pageNumber="45" refString="Chin, P. K. (1990) A new Bornean species of loach, Elxis sabanus (Cobitidae) from Mendolong, Sabah. Malayan Nature Journal, 43 (1989 [1990]), 72 - 76." type="journal article" year="1990">
<collectingCountry id="F3697611B34EFFDAFE4643FFFC7B46E4" box="[494,554,914,940]" name="China" pageId="6" pageNumber="45">Chin</collectingCountry>
, 1990
</bibRefCitation>
; 
<bibRefCitation id="EFEF4B70B34EFFDAFD2943FFFD3E46E4" author="Kottelat" box="[641,879,914,940]" pageId="6" pageNumber="45" refString="Kottelat, M., Whitten, A. J., Kartikasari, S. N. &amp; Wirjoatmodjo, S. (1993) Freshwater fishes of Western Indonesia and Sulawesi. Periplus, Hong Kong, 259 pp., 84 pls." type="book" year="1993">Kottelat et al., 1993</bibRefCitation>
). Among the Bornean 
<taxonomicName id="4C7E4D02B34EFFDAFBD243FFFF05469B" authority="Hadiaty &amp; Kottelat (2009)" authorityName="Hadiaty &amp; Kottelat" authorityYear="2009" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B90AEA93B34EFFDAFBD243FFFB4046E3" box="[1146,1297,914,939]" italics="true" pageId="6" pageNumber="45">Nemacheilus</emphasis>
, 
<bibRefCitation id="EFEF4B70B34EFFDAFA8843FFFF05469B" author="Hadiaty" pageId="6" pageNumber="45" refString="Hadiaty, R. K. &amp; Kottelat, M. (2009) Nemacheilus tebo, a new nemacheiline loach from Sangkulirang Karst, East Kalimantan, Indonesia (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 57, 119 - 125." type="journal article" year="2009">Hadiaty &amp; Kottelat (2009)</bibRefCitation>
</taxonomicName>
recognised a 
<taxonomicName id="4C7E4D02B34EFFDAFDA843D4FC90469A" box="[512,705,953,978]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B34EFFDAFDA843D4FC90469A" box="[512,705,953,978]" italics="true" pageId="6" pageNumber="45">N. selangoricus</emphasis>
</taxonomicName>
group diagnosed by the presence of two rows of modified, elongated scales on the caudal peduncle, with a tubercle on their posterior extremity, often at the tip of an acuminate process. In Borneo, this group includes 
<taxonomicName id="4C7E4D02B34EFFDAFCAC4468FDF14156" box="[772,928,1029,1054]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="spiniferus">
<emphasis id="B90AEA93B34EFFDAFCAC4468FDF14156" box="[772,928,1029,1054]" italics="true" pageId="6" pageNumber="45">N. spiniferus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C7E4D02B34EFFDAFC754468FA644156" box="[989,1077,1029,1054]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="tebo">
<emphasis id="B90AEA93B34EFFDAFC754468FA644156" box="[989,1077,1029,1054]" italics="true" pageId="6" pageNumber="45">N. tebo</emphasis>
</taxonomicName>
. In addition, there is a single record of 
<taxonomicName id="4C7E4D02B34EFFDAFEA14446FF90410C" box="[265,449,1067,1092]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B34EFFDAFEA14446FF90410C" box="[265,449,1067,1092]" italics="true" pageId="6" pageNumber="45">N. selangoricus</emphasis>
</taxonomicName>
from Sabah based on specimens collected in 1956 (
<bibRefCitation id="EFEF4B70B34EFFDAFB8A4446FA81410D" author="Kottelat" box="[1058,1232,1067,1093]" pageId="6" pageNumber="45" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat, 1990</bibRefCitation>
). The species has not been collected again in Sabah and its presence needs confirmation; there are indications that specimens from different lots and their respective labels may have been mixed by an earlier researcher. 
<taxonomicName id="4C7E4D02B34EFFDAFAA94415FF7C41FF" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B34EFFDAFAA94415FF7C41FF" italics="true" pageId="6" pageNumber="45">Nemacheilus selangoricus</emphasis>
</taxonomicName>
is known from Sumatra and the southern Malay Peninsula. The remaining species have not been classified into groups.
</paragraph>
<paragraph id="8BC13681B34EFFDAFF6E4487FD574062" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">
The colour pattern of 
<taxonomicName id="4C7E4D02B34EFFDAFE784487FC1E404B" box="[464,591,1258,1283]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFE784487FC1E404B" box="[464,591,1258,1283]" italics="true" pageId="6" pageNumber="45">N. marang</emphasis>
</taxonomicName>
distinguishes it from the other species of 
<taxonomicName id="4C7E4D02B34EFFDAFBE64487FAB4404B" box="[1102,1253,1258,1283]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B90AEA93B34EFFDAFBE64487FAB4404B" box="[1102,1253,1258,1283]" italics="true" pageId="6" pageNumber="45">Nemacheilus</emphasis>
</taxonomicName>
of Borneo and only shares gross similarity with that of 
<taxonomicName id="4C7E4D02B34EFFDAFDC6457DFCAF4061" box="[622,766,1296,1321]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B34EFFDAFDC6457DFCAF4061" box="[622,766,1296,1321]" italics="true" pageId="6" pageNumber="45">N. olivaceus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC13681B34EFFDAFF6E455BFDEB4313" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">
<taxonomicName id="4C7E4D02B34EFFDAFF6E455BFF8A4007" box="[198,475,1334,1359]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B34EFFDAFF6E455BFF8A4007" box="[198,475,1334,1359]" italics="true" pageId="6" pageNumber="45">Nemacheilus olivaceus</emphasis>
</taxonomicName>
(from Sabah; 
<figureCitation id="13452A04B34EFFDAFD3F455BFCB34018" box="[663,738,1334,1360]" captionStart="FIGURE 5" captionStartId="5.[151,255,1512,1536]" captionTargetBox="[168,1418,1140,1489]" captionTargetId="figure@5.[168,1419,1140,1491]" captionTargetPageId="5" captionText="FIGURE 5. Nemacheilus olivaceus, ZRC 45453, 55.4 mm SL; Borneo: Sabah: Danum Valley (photograph by Tan Heok Hui)." httpUri="https://zenodo.org/record/196942/files/figure.png" pageId="6" pageNumber="45">Fig. 5</figureCitation>
) has a colour pattern made of 12–15 dark-brown bars, regularly shaped except for the last few ones, slightly narrower or equal to interspaces; without conspicuous axial stripe; no patterning on the dorsal fin, except in very few specimens, which have a faint subdistal row of brown spots on rays; and the blackish brown bar at caudal-fin base made of a lower elongate, straight vertical part on lower 2/3 of caudal, and an upper slanted blackish mark, more or less reaching lower one. Further 
<taxonomicName id="4C7E4D02B34EFFDAFA2945A2FEA54346" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFA2945A2FEA54346" italics="true" pageId="6" pageNumber="45">N. marang</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C7E4D02B34EFFDAFDAD4598FCCA4346" box="[517,667,1525,1550]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B34EFFDAFDAD4598FCCA4346" box="[517,667,1525,1550]" italics="true" pageId="6" pageNumber="45">N. olivaceus</emphasis>
</taxonomicName>
by the absence of a median notch in lower lip (vs. presence), a forked caudal fin (vs. deeply emarginate), and the anus located more posteriorly (behind middle of distance between pelvic-fin base and anal-fin origin, vs. about in the middle).
</paragraph>
<paragraph id="8BC13681B34EFFDAFF6E460AFBC543BB" blockId="6.[151,1437,152,2008]" pageId="6" pageNumber="45">
<taxonomicName id="4C7E4D02B34EFFDAFF6E460AFFA243C8" box="[198,499,1639,1664]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="kapuasensis">
<emphasis id="B90AEA93B34EFFDAFF6E460AFFA243C8" box="[198,499,1639,1664]" italics="true" pageId="6" pageNumber="45">Nemacheilus kapuasensis</emphasis>
</taxonomicName>
(from Kapuas drainage, Kalimantan Barat; 
<figureCitation id="13452A04B34EFFDAFC50460AFA6C43C9" box="[1016,1085,1639,1665]" captionStart="FIGURE 6" captionStartId="5.[151,255,2009,2033]" captionTargetBox="[202,1387,1642,1955]" captionTargetId="figure@5.[175,1411,1619,1988]" captionTargetPageId="5" captionText="FIGURE 6. Nemacheilus kapuasensis, CMK 3187, 49.7 mm SL; Borneo: Kalimantan Barat: Kapuas drainage." httpUri="https://zenodo.org/record/196943/files/figure.png" pageId="6" pageNumber="45">Fig. 6</figureCitation>
) has a colour pattern made of 10–12 broad dark-brown bars (width equal to interspaces), in large specimens broken up into a series of saddles alternating with a series of midlateral blotches. 
<taxonomicName id="4C7E4D02B34EFFDAFC8E46DEFA714384" box="[806,1056,1715,1740]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFC8E46DEFA714384" box="[806,1056,1715,1740]" italics="true" pageId="6" pageNumber="45">Nemacheilus marang</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName id="4C7E4D02B34EFFDAFA2946DEFF7443BA" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="kapuasensis">
<emphasis id="B90AEA93B34EFFDAFA2946DEFF7443BA" italics="true" pageId="6" pageNumber="45">N. kapuasensis</emphasis>
</taxonomicName>
in having 8½ branched dorsal-fin rays (vs. 9½) and fewer lateral-line pores (84–92, vs. 93–102).
</paragraph>
</subSubSection>
<subSubSection id="C364650AB34EFFD4FF6E476DFEA24296" lastPageId="8" lastPageNumber="47" pageId="6" pageNumber="45" type="materials_examined">
<paragraph id="8BC13681B34EFFDBFF6E476DFF154590" blockId="6.[151,1437,152,2008]" lastBlockId="7.[151,1437,152,553]" lastPageId="7" lastPageNumber="46" pageId="6" pageNumber="45">
<taxonomicName id="4C7E4D02B34EFFDAFF6E476DFC484251" box="[198,537,1792,1817]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="elegantissimus">
<emphasis id="B90AEA93B34EFFDAFF6E476DFC484251" box="[198,537,1792,1817]" italics="true" pageId="6" pageNumber="45">Nemacheilus elegantissimus</emphasis>
</taxonomicName>
(from Sabah; 
<figureCitation id="13452A04B34EFFDAFD724692FD764251" box="[730,807,1791,1817]" captionStart="FIGURE 7" captionStartId="7.[151,255,1035,1059]" captionTargetBox="[161,1412,635,978]" captionTargetId="figure@7.[151,1436,606,1014]" captionTargetPageId="7" captionText="FIGURE 7. Nemacheilus elegantissimus, CMK 7937, paratype, 46.0 mm SL; Borneo: Sabah: Danum Valley." httpUri="https://zenodo.org/record/196944/files/figure.png" pageId="6" pageNumber="45">Fig. 7</figureCitation>
) has a midlateral row of 17–21 small, vertically elongated black blotches, about 17–21 saddles along the dorsal mid-line, a vertically elongated black spot occupying most of the lower half of the caudal-fin base, circled by a whitish area. Further, in 
<taxonomicName id="4C7E4D02B34EFFDAFB594721FB3E422D" box="[1265,1391,1868,1893]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFB594721FB3E422D" box="[1265,1391,1868,1893]" italics="true" pageId="6" pageNumber="45">N. marang</emphasis>
</taxonomicName>
the dorsal fin pattern is restricted to 2–3 rows of faint brown spots on rays, while in 
<taxonomicName id="4C7E4D02B34EFFDAFBE4471FFB4A42C3" box="[1100,1307,1906,1931]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="elegantissimus">
<emphasis id="B90AEA93B34EFFDAFBE4471FFB4A42C3" box="[1100,1307,1906,1931]" italics="true" pageId="6" pageNumber="45">N. elegantissimus</emphasis>
</taxonomicName>
part of the anterior edge of the fin is black, there are 2 rows of black spots on rays, one subdistal and one subproximal; and 2–3 black blotches extending on membranes over the subproximal row. 
<taxonomicName id="4C7E4D02B34EFFDAFB8B47D3FB71429F" box="[1059,1312,1982,2007]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="45" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34EFFDAFB8B47D3FB71429F" box="[1059,1312,1982,2007]" italics="true" pageId="6" pageNumber="45">Nemacheilus marang</emphasis>
</taxonomicName>
also has a somewhat stouter body (body depth 14.0–20.7 % SL, vs. 10.7–18.9 [examined material and from 
<bibRefCitation id="EFEF4B70B34FFFDBFAEA40F5FF614590" author="Chin" pageId="7" pageNumber="46" refString="Chin, P. K. &amp; Samat, A. (1992) A new species of loach, Nemacheilus elegantissimus, (family Balitoridae, subfamily Nemacheilinae), from Danum Valley, Sabah, Malaysia. Malayan Nature Journal, 46, 25 - 33." type="journal article" year="1992">
<collectingCountry id="F3697611B34FFFDBFAEA40F5FB2C45FA" box="[1346,1405,152,178]" name="China" pageId="7" pageNumber="46">Chin</collectingCountry>
&amp; Samat, 1992
</bibRefCitation>
]).
</paragraph>
<paragraph id="8BC13681B34FFFDBFF6E4089FD434761" blockId="7.[151,1437,152,553]" pageId="7" pageNumber="46">
<taxonomicName id="4C7E4D02B34FFFDBFF6E4089FC4745B5" box="[198,534,228,253]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="46" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B34FFFDBFF6E4089FC4745B5" box="[198,534,228,253]" italics="true" pageId="7" pageNumber="46">Nemacheilus longipectoralis</emphasis>
</taxonomicName>
(from Sabah and Kalimantan Timur; 
<figureCitation id="13452A04B34FFFDBFC644089FA4145B6" box="[972,1040,228,254]" captionStart="FIGURE 8" captionStartId="7.[151,256,1488,1512]" captionTargetBox="[202,1372,1144,1427]" captionTargetId="figure@7.[181,1405,1110,1466]" captionTargetPageId="7" captionText="FIGURE 8. Nemacheilus longipectoralis, MZB uncat., 35.0 mm SL; Borneo: Kalimantan Timur: Mahakam drainage; pectoral fin oriented perpendicular to fish, appearing shorter." httpUri="https://zenodo.org/record/196945/files/figure.png" pageId="7" pageNumber="46">Fig. 8</figureCitation>
) has a colour pattern made of 16– 20 long, narrow dark brown bars, irregularly shaped, some connected, some branched, continuous or not across the back, often appearing as pairs of bars connected to a single saddle in front of dorsal fin. In 
<taxonomicName id="4C7E4D02B34FFFDBFA284142FEA34425" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="46" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34FFFDBFA284142FEA34425" italics="true" pageId="7" pageNumber="46">N. marang</emphasis>
</taxonomicName>
the dorsal-fin pattern is restricted to 2–3 rows of faint brown spots on rays, while in 
<taxonomicName id="4C7E4D02B34FFFDBFB614139FBCD4425" box="[1225,1436,340,365]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="46" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B34FFFDBFB614139FBCD4425" box="[1225,1436,340,365]" italics="true" pageId="7" pageNumber="46">N. longipectoralis</emphasis>
</taxonomicName>
part of the anterior edge of the fin is black; there are 2 rows of black spots on rays, one subdistal and one at about 1/3 the length of the rays; the anterior spot of the lower row is enlarged and extends over the first two branched rays and the membrane inbetween. Further, in 
<taxonomicName id="4C7E4D02B34FFFDBFCE141A9FD9A4495" box="[841,971,452,477]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="46" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B34FFFDBFCE141A9FD9A4495" box="[841,971,452,477]" italics="true" pageId="7" pageNumber="46">N. marang</emphasis>
</taxonomicName>
, the pectoral fin reaches only slightly beyond half of distance to pelvic-fin base (vs. almost to pelvic-fin origin in material from Sabah and beyond in material from the Mahakam drainage [
<typeStatus id="54C58823B34FFFDBFDDF4262FCFD4761" box="[631,684,527,553]" pageId="7" pageNumber="46">type</typeStatus>
series]).
</paragraph>
<caption id="DF016609B34FFFDBFF3F4466FB76416B" box="[151,1319,1035,1059]" httpUri="https://zenodo.org/record/196944/files/figure.png" pageId="7" pageNumber="46" targetBox="[161,1412,635,978]" targetPageId="7">
<paragraph id="8BC13681B34FFFDBFF3F4466FB76416B" blockId="7.[151,1319,1035,1059]" box="[151,1319,1035,1059]" pageId="7" pageNumber="46">
<emphasis id="B90AEA93B34FFFDBFF3F4466FF48416B" bold="true" box="[151,281,1035,1059]" pageId="7" pageNumber="46">FIGURE 7.</emphasis>
<taxonomicName id="4C7E4D02B34FFFDBFE884461FC1C416B" box="[288,589,1036,1059]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="46" phylum="Chordata" rank="species" species="elegantissimus">
<emphasis id="B90AEA93B34FFFDBFE884461FC1C416B" box="[288,589,1036,1059]" italics="true" pageId="7" pageNumber="46">Nemacheilus elegantissimus</emphasis>
</taxonomicName>
, CMK 7937, paratype, 46.0 mm SL; Borneo: Sabah: Danum Valley.
</paragraph>
</caption>
<caption id="DF016609B34FFFDBFF3F45BDFD4E434F" httpUri="https://zenodo.org/record/196945/files/figure.png" pageId="7" pageNumber="46" targetBox="[202,1372,1144,1427]" targetPageId="7">
<paragraph id="8BC13681B34FFFDBFF3F45BDFD4E434F" blockId="7.[151,1436,1488,1543]" pageId="7" pageNumber="46">
<emphasis id="B90AEA93B34FFFDBFF3F45BDFF4D40A0" bold="true" box="[151,284,1488,1512]" pageId="7" pageNumber="46">FIGURE 8.</emphasis>
<taxonomicName id="4C7E4D02B34FFFDBFE8B45BDFC0840AF" box="[291,601,1488,1511]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="46" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B34FFFDBFE8B45BDFC0840AF" box="[291,601,1488,1511]" italics="true" pageId="7" pageNumber="46">Nemacheilus longipectoralis</emphasis>
</taxonomicName>
, MZB uncat., 35.0 mm SL; Borneo: Kalimantan Timur: Mahakam drainage; pectoral fin oriented perpendicular to fish, appearing shorter.
</paragraph>
</caption>
<caption id="DF016609B34FFFDBFF3F47F1FF6F429C" httpUri="https://zenodo.org/record/196946/files/figure.png" pageId="7" pageNumber="46" targetBox="[185,1401,1595,1927]" targetPageId="7">
<paragraph id="8BC13681B34FFFDBFF3F47F1FF6F429C" blockId="7.[151,1436,1948,2004]" pageId="7" pageNumber="46">
<emphasis id="B90AEA93B34FFFDBFF3F47F1FF4A42FC" bold="true" box="[151,283,1948,1972]" pageId="7" pageNumber="46">FIGURE 9.</emphasis>
<taxonomicName id="4C7E4D02B34FFFDBFE8A47F0FC6E42FC" box="[290,575,1949,1972]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="46" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B34FFFDBFE8A47F0FC6E42FC" box="[290,575,1949,1972]" italics="true" pageId="7" pageNumber="46">Nemacheilus saravacensis</emphasis>
</taxonomicName>
, ZRC 39433, 38.8 mm SL; Borneo: Sarawak: Sadong drainage (photograph by Tan Heok Hui).
</paragraph>
</caption>
<caption id="DF016609B340FFD4FF3F42E4FA6B47E8" box="[151,1082,649,673]" httpUri="https://zenodo.org/record/196947/files/figure.png" pageId="8" pageNumber="47" targetBox="[190,1396,194,624]" targetPageId="8">
<paragraph id="8BC13681B340FFD4FF3F42E4FA6B47E8" blockId="8.[151,1082,649,673]" box="[151,1082,649,673]" pageId="8" pageNumber="47">
<emphasis id="B90AEA93B340FFD4FF3F42E4FF7747E9" bold="true" box="[151,294,649,673]" pageId="8" pageNumber="47">FIGURE 10.</emphasis>
<taxonomicName id="4C7E4D02B340FFD4FE8542E4FC1447E8" box="[301,581,649,672]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B340FFD4FE8542E4FC1447E8" box="[301,581,649,672]" italics="true" pageId="8" pageNumber="47">Nemacheilus selangoricus</emphasis>
</taxonomicName>
, CMK 9636, 48.0 mm SL; Indonesia: Bangka.
</paragraph>
</caption>
<paragraph id="8BC13681B340FFD4FF6E42A2FA1546EB" blockId="8.[151,1437,718,2014]" pageId="8" pageNumber="47">
<taxonomicName id="4C7E4D02B340FFD4FF6E42A2FFAC47A0" box="[198,509,719,744]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B340FFD4FF6E42A2FFAC47A0" box="[198,509,719,744]" italics="true" pageId="8" pageNumber="47">Nemacheilus saravacensis</emphasis>
</taxonomicName>
(from Kalimantan Barat and 
<collectingCountry id="F3697611B340FFD4FCFD42A3FDEF47A0" box="[853,958,718,744]" name="Malaysia" pageId="8" pageNumber="47">Sarawak</collectingCountry>
; 
<figureCitation id="13452A04B340FFD4FC6142A3FA5C47A0" box="[969,1037,718,744]" captionStart="FIGURE 9" captionStartId="7.[151,255,1948,1972]" captionTargetBox="[185,1401,1595,1927]" captionTargetId="figure@7.[185,1401,1595,1928]" captionTargetPageId="7" captionText="FIGURE 9. Nemacheilus saravacensis, ZRC 39433, 38.8 mm SL; Borneo: Sarawak: Sadong drainage (photograph by Tan Heok Hui)." httpUri="https://zenodo.org/record/196946/files/figure.png" pageId="8" pageNumber="47">Fig. 9</figureCitation>
) has a colour pattern made of 13– 17 irregular brown blotches along the lateral line, most extending to dorsal midline and those on caudal peduncle extending to ventral midline; 13–17 dark brown saddles along back, connected or not to midlateral blotches. 
<taxonomicName id="4C7E4D02B340FFD4FEAE4352FFAE4610" box="[262,511,831,856]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B340FFD4FEAE4352FFAE4610" box="[262,511,831,856]" italics="true" pageId="8" pageNumber="47">Nemacheilus marang</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName id="4C7E4D02B340FFD4FCF34352FA424610" box="[859,1043,831,856]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B340FFD4FCF34352FA424610" box="[859,1043,831,856]" italics="true" pageId="8" pageNumber="47">N. saravacensis</emphasis>
</taxonomicName>
in having 17 branched caudal-fin rays (vs. 
<quantity id="4C869B64B340FFD4FEAA430EFF6F4635" box="[258,318,867,893]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="8" pageNumber="47" unit="in" value="16.0">16 in</quantity>
most specimens), 8½ branched dorsal-fin rays (vs. 9–10½), and the anterior nostril at the front side of a pointed flap (vs. at extremity of an obliquely cut tube with pointed tip).
</paragraph>
<paragraph id="8BC13681B340FFD4FF6E43C2FFDA41E0" blockId="8.[151,1437,718,2014]" pageId="8" pageNumber="47">
<taxonomicName id="4C7E4D02B340FFD4FF6E43C2FFAD4680" box="[198,508,943,968]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B340FFD4FF6E43C2FFAD4680" box="[198,508,943,968]" italics="true" pageId="8" pageNumber="47">Nemacheilus selangoricus</emphasis>
</taxonomicName>
(doubtfully present in Sabah, see above; 
<figureCitation id="13452A04B340FFD4FC4D43C3FA6B4680" box="[997,1082,942,968]" captionStart="FIGURE 10" captionStartId="8.[151,255,649,673]" captionTargetBox="[190,1396,194,624]" captionTargetId="figure@8.[190,1396,194,627]" captionTargetPageId="8" captionText="FIGURE 10. Nemacheilus selangoricus, CMK 9636, 48.0 mm SL; Indonesia: Bangka." httpUri="https://zenodo.org/record/196947/files/figure.png" pageId="8" pageNumber="47">Fig. 10</figureCitation>
) has a colour pattern made of conspicuous dark brown bars, much wider than interspaces, a conspicuous blackish bar across eye, and a large black spot at the base of the anterior rays of the dorsal fin. 
<taxonomicName id="4C7E4D02B340FFD4FCE34394FA14415A" box="[843,1093,1017,1042]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="marang">
<emphasis id="B90AEA93B340FFD4FCE34394FA14415A" box="[843,1093,1017,1042]" italics="true" pageId="8" pageNumber="47">Nemacheilus marang</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName id="4C7E4D02B340FFD4FF3F4472FF014170" box="[151,336,1055,1080]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="selangoricus">
<emphasis id="B90AEA93B340FFD4FF3F4472FF014170" box="[151,336,1055,1080]" italics="true" pageId="8" pageNumber="47">N. selangoricus</emphasis>
</taxonomicName>
in having a less deeply forked caudal fin (upper lobe 1.0-1.5 times median rays, vs. 1.5–2.4) with subequal lobes (vs. upper lobe 1.1–1.3 times longer than lower one), anterior nostril pierced in front side of a pointed flap (vs. at tip of a small tube), and without acuminate scales (vs. conspicuous acuminate scales on caudal peduncle).
</paragraph>
<paragraph id="8BC13681B340FFD4FF6E44DEFFF541BB" blockId="8.[151,1437,718,2014]" pageId="8" pageNumber="47">
<emphasis id="B90AEA93B340FFD4FF6E44DEFFB24185" bold="true" box="[198,483,1203,1229]" pageId="8" pageNumber="47">Comparative material.</emphasis>
Morphometric data are in part from 
<bibRefCitation id="EFEF4B70B340FFD4FC3944D9FA154186" author="Kottelat" box="[913,1092,1204,1230]" pageId="8" pageNumber="47" refString="Kottelat, M. (1984) Revision of the Indonesian and Malaysian loaches of the subfamily Noemacheilinae. Japanese Journal of Ichthyology, 31, 225 - 260." type="journal article" year="1984">Kottelat (1984)</bibRefCitation>
and from our examination of the following material:
</paragraph>
<paragraph id="8BC13681B340FFD4FF6E4492FDE1402B" blockId="8.[151,1437,718,2014]" pageId="8" pageNumber="47">
<taxonomicName id="4C7E4D02B340FFD4FF6E4492FC454050" box="[198,532,1279,1304]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="elegantissimus">
<emphasis id="B90AEA93B340FFD4FF6E4492FC454050" box="[198,532,1279,1304]" italics="true" pageId="8" pageNumber="47">Nemacheilus elegantissimus</emphasis>
</taxonomicName>
: 
<collectionCode id="ED6FAE44B340FFD4FD804493FC304050" box="[552,609,1278,1304]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
40381, 1, 
<quantity id="4C869B64B340FFD4FD404493FD054050" box="[744,852,1278,1304]" metricMagnitude="-2" metricUnit="m" metricValue="4.779999999999999" pageId="8" pageNumber="47" unit="mm" value="47.8">47.8 mm</quantity>
SL; Sabah: Lahad Datu district: Danum Valley, Sungai Lonpodas: K. Martin-Smith, 
<date id="FFC01041B340FFD4FDE04549FC8B4076" box="[584,730,1316,1342]" pageId="8" pageNumber="47" value="1996-06-04">4 June 1996</date>
. — CMK 7937, 
<typeStatus id="54C58823B340FFD4FC094549FA584076" box="[929,1033,1316,1342]" pageId="8" pageNumber="47" type="paratype">paratype</typeStatus>
, 46.0 mm SL; Sabah: Lahad Datu district: Danum Valley, Sungai Payau; A. Samat, 
<date id="FFC01041B340FFD4FD704524FDFD402B" box="[728,940,1353,1379]" pageId="8" pageNumber="47" value="1990-12-07">7 December 1990</date>
.
</paragraph>
<paragraph id="8BC13681B340FFD4FF6E4502FC2E4356" blockId="8.[151,1437,718,2014]" pageId="8" pageNumber="47">
<taxonomicName id="4C7E4D02B340FFD4FF6E4502FFA540C0" box="[198,500,1391,1416]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="kapuasensis">
<emphasis id="B90AEA93B340FFD4FF6E4502FFA540C0" box="[198,500,1391,1416]" italics="true" pageId="8" pageNumber="47">Nemacheilus kapuasensis</emphasis>
</taxonomicName>
: 
<collectionCode id="ED6FAE44B340FFD4FDAC4503FC6C40C0" box="[516,573,1390,1416]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
44047, 1, 
<quantity id="4C869B64B340FFD4FD144503FD4140C0" box="[700,784,1390,1416]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="47" unit="mm" value="60.0">60 mm</quantity>
SL; 
<collectingCountry id="F3697611B340FFD4FCE24503FDEB40C0" box="[842,954,1390,1416]" name="Malaysia" pageId="8" pageNumber="47">Malaysia</collectingCountry>
: 
<collectingCountry id="F3697611B340FFD4FC6E4503FA7E40C0" box="[966,1071,1390,1416]" name="Malaysia" pageId="8" pageNumber="47">Sarawak</collectingCountry>
: Bau area; H. H. Tan, 
<date id="FFC01041B340FFD4FA914503FE8740E6" pageId="8" pageNumber="47" value="2004-04-24">24 April 2004</date>
. — 
<collectionCode id="ED6FAE44B340FFD4FEAF45F9FF1140E6" box="[263,320,1428,1454]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
38823, 2, 
<quantity id="4C869B64B340FFD4FE6945F9FC3C40E6" box="[449,621,1428,1454]" metricMagnitude="-2" metricUnit="m" metricValue="5.215" metricValueMax="5.29" metricValueMin="5.14" pageId="8" pageNumber="47" unit="mm" value="52.15" valueMax="52.9" valueMin="51.4">51.4–52.9 mm</quantity>
SL, 
<collectingCountry id="F3697611B340FFD4FD0F45F9FD4F40E6" box="[679,798,1428,1454]" name="Indonesia" pageId="8" pageNumber="47">Indonesia</collectingCountry>
: Borneo: Kalimantan Barat: Kapuas drainage, Sungai Lanjak in Lanjak and Sungai Lanjak Deras 
<quantity id="4C869B64B340FFD4FD3D45D4FC80409B" box="[661,721,1465,1491]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="47" unit="km" value="1.0">1 km</quantity>
east of Lanjak; M. Kottelat, 
<date id="FFC01041B340FFD4FB8B45D4FA93409B" box="[1059,1218,1465,1491]" pageId="8" pageNumber="47" value="1995-06-11">11 June 1995</date>
. — CMK 3187, 3, 41.2-54.0 mm SL; Borneo: Kalimantan Barat: Kapuas drainage: Sungai Pinoh at Nangu Saian, 
<quantity id="4C869B64B340FFD4FABE45B3FB3340B0" box="[1302,1378,1502,1528]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="47" unit="km" value="45.0">45 km</quantity>
S of Nangapinoh; T. R. Roberts, 
<date id="FFC01041B340FFD4FE484669FC2A4356" box="[480,635,1540,1566]" pageId="8" pageNumber="47" value="1976-07-26">26 July 1976</date>
.
</paragraph>
<paragraph id="8BC13681B340FFD4FF6E4644FA3143B6" blockId="8.[151,1437,718,2014]" pageId="8" pageNumber="47">
<taxonomicName id="4C7E4D02B340FFD4FF6E4644FC4C430A" box="[198,541,1577,1602]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="longipectoralis">
<emphasis id="B90AEA93B340FFD4FF6E4644FC4C430A" box="[198,541,1577,1602]" italics="true" pageId="8" pageNumber="47">Nemacheilus longipectoralis</emphasis>
</taxonomicName>
: 
<collectionCode id="ED6FAE44B340FFD4FD9B4644FCDA430B" LSID="urn:lsid:biocol.org:col:34992" box="[563,651,1577,1603]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="8" pageNumber="47">RMNH</collectionCode>
7641, 
<typeStatus id="54C58823B340FFD4FD414644FD06430B" box="[745,855,1577,1603]" pageId="8" pageNumber="47" type="lectotype">lectotype</typeStatus>
, 
<quantity id="4C869B64B340FFD4FCC14644FD88430B" box="[873,985,1577,1603]" metricMagnitude="-2" metricUnit="m" metricValue="3.44" pageId="8" pageNumber="47" unit="mm" value="34.4">34.4 mm</quantity>
SL; 
<collectingCountry id="F3697611B340FFD4FBB74644FAC4430B" box="[1055,1173,1577,1603]" name="Indonesia" pageId="8" pageNumber="47">Indonesia</collectingCountry>
: Borneo: Kalimantan Timur: upper Mahakam; A. W. Nieuwenhuis, 
<date id="FFC01041B340FFD4FD194623FD234320" box="[689,882,1614,1640]" pageId="8" pageNumber="47" value="1898-11">November 1898</date>
. — 
<collectionCode id="ED6FAE44B340FFD4FC0A4623FDAB4320" LSID="urn:lsid:biocol.org:col:34992" box="[930,1018,1614,1640]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="8" pageNumber="47">RMNH</collectionCode>
27350, 2 
<typeStatus id="54C58823B340FFD4FBC74623FB4A4320" box="[1135,1307,1614,1640]" pageId="8" pageNumber="47" type="paralectotype">paralectotypes</typeStatus>
, 
<quantity id="4C869B64B340FFD4FA8F4623FE9443C6" metricMagnitude="-2" metricUnit="m" metricValue="3.28" metricValueMax="3.67" metricValueMin="2.8899999999999997" pageId="8" pageNumber="47" unit="mm" value="32.8" valueMax="36.7" valueMin="28.9">28.9–36.7 mm</quantity>
SL; 
<collectingCountry id="F3697611B340FFD4FEA94619FF2B43C6" box="[257,378,1652,1678]" name="Indonesia" pageId="8" pageNumber="47">Indonesia</collectingCountry>
: Borneo: Kalimantan Timur: upper Mahakam; A. W. Nieuwenhuis, 
<date id="FFC01041B340FFD4FB054619FB2243C6" box="[1197,1395,1652,1678]" pageId="8" pageNumber="47" value="1898-11">November 1898</date>
. — 
<collectionCode id="ED6FAE44B340FFD4FF3F46F4FE8643FB" box="[151,215,1689,1715]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="8" pageNumber="47">MZB</collectionCode>
16479, 1, 
<quantity id="4C869B64B340FFD4FEFA46F4FFEB43FB" box="[338,442,1689,1715]" metricMagnitude="-2" metricUnit="m" metricValue="3.94" pageId="8" pageNumber="47" unit="mm" value="39.4">39.4 mm</quantity>
SL; 
<collectingCountry id="F3697611B340FFD4FE5A46F4FC3843FB" box="[498,617,1689,1715]" name="Indonesia" pageId="8" pageNumber="47">Indonesia</collectingCountry>
: Borneo: Kalimantan Timur: Mahakam drainage: Sungai Pleo, Sungai Senget Kiri; R. K. Hadiaty et al., 
<date id="FFC01041B340FFD4FD9F46D3FC814390" box="[567,720,1726,1752]" pageId="8" pageNumber="47" value="2008-06-06">6 June 2008</date>
. — 
<collectionCode id="ED6FAE44B340FFD4FCAB46D3FD6F4390" box="[771,830,1726,1752]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
40383, 3, 
<quantity id="4C869B64B340FFD4FC6A46D3FA254390" box="[962,1140,1726,1752]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.8299999999999996" metricValueMin="3.37" pageId="8" pageNumber="47" unit="mm" value="36.0" valueMax="38.3" valueMin="33.7">33.7–38.3 mm</quantity>
SL; Sabah: Lahad Datu district: Danum Valley, Sungai Lonpodas; H. H. Tan &amp; 
<collectingCountry id="F3697611B340FFD4FC894689FD6743B6" box="[801,822,1764,1790]" name="Italy" pageId="8" pageNumber="47">Y</collectingCountry>
. 
<collectingCountry id="F3697611B340FFD4FCE94689FD0743B6" box="[833,854,1764,1790]" name="Italy" pageId="8" pageNumber="47">Y</collectingCountry>
. Goh, 
<date id="FFC01041B340FFD4FC0A4689FA0D43B6" box="[930,1116,1764,1790]" pageId="8" pageNumber="47" value="1996-10-03">3 October 1996</date>
.
</paragraph>
<paragraph id="8BC13681B340FFD4FF6E4764FDBB42DB" blockId="8.[151,1437,718,2014]" pageId="8" pageNumber="47">
<taxonomicName id="4C7E4D02B340FFD4FF6E4764FF84426A" box="[198,469,1801,1826]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="olivaceus">
<emphasis id="B90AEA93B340FFD4FF6E4764FF84426A" box="[198,469,1801,1826]" italics="true" pageId="8" pageNumber="47">Nemacheilus olivaceus</emphasis>
</taxonomicName>
: 
<collectionCode id="ED6FAE44B340FFD4FE4D4764FC4F426B" box="[485,542,1801,1827]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
43986, 1, 
<quantity id="4C869B64B340FFD4FD374764FD58426B" box="[671,777,1801,1827]" metricMagnitude="-2" metricUnit="m" metricValue="4.9799999999999995" pageId="8" pageNumber="47" unit="mm" value="49.8">49.8 mm</quantity>
SL; 
<collectingCountry id="F3697611B340FFD4FCEC4764FDE5426B" box="[836,948,1801,1827]" name="Malaysia" pageId="8" pageNumber="47">Malaysia</collectingCountry>
: Sabah: Danum Valley, small tributary to Sungai Bole, ca. 
<quantity id="4C869B64B340FFD4FECA4743FFFF4200" box="[354,430,1838,1864]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="47" unit="m" value="500.0">500 m</quantity>
into coupe 93, H. H. Tan &amp; 
<collectingCountry id="F3697611B340FFD4FCAD4743FD4B4200" box="[773,794,1838,1864]" name="Italy" pageId="8" pageNumber="47">Y</collectingCountry>
. 
<collectingCountry id="F3697611B340FFD4FC8F4743FD6D4200" box="[807,828,1838,1864]" name="Italy" pageId="8" pageNumber="47">Y</collectingCountry>
. Goh, 
<date id="FFC01041B340FFD4FC224743FA194200" box="[906,1096,1838,1864]" pageId="8" pageNumber="47" value="1996-10-02">2 October 1996</date>
. — 
<collectionCode id="ED6FAE44B340FFD4FBD14743FAE34200" box="[1145,1202,1838,1864]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
45411, 1, 
<quantity id="4C869B64B340FFD4FA9A4743FBCD4200" box="[1330,1436,1838,1864]" metricMagnitude="-2" metricUnit="m" metricValue="4.39" pageId="8" pageNumber="47" unit="mm" value="43.9">43.9 mm</quantity>
SL; Borneo: Sabah: Danum Valley; H. H. Tan &amp; 
<collectingCountry id="F3697611B340FFD4FD7B4739FCB94226" box="[723,744,1876,1902]" name="Italy" pageId="8" pageNumber="47">Y</collectingCountry>
. 
<collectingCountry id="F3697611B340FFD4FD5B4739FD594226" box="[755,776,1876,1902]" name="Italy" pageId="8" pageNumber="47">Y</collectingCountry>
. Goh; 
<date id="FFC01041B340FFD4FCFD4739FA5F4226" box="[853,1038,1876,1902]" pageId="8" pageNumber="47" value="1996-10-01">1 October 1996</date>
. — 
<collectionCode id="ED6FAE44B340FFD4FB954739FA274226" box="[1085,1142,1876,1902]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="47">ZRC</collectionCode>
45485, 5, 
<quantity id="4C869B64B340FFD4FB5A4739FBCD4226" box="[1266,1436,1876,1902]" metricMagnitude="-2" metricUnit="m" metricValue="4.455" metricValueMax="4.85" metricValueMin="4.06" pageId="8" pageNumber="47" unit="mm" value="44.55" valueMax="48.5" valueMin="40.6">40.6–48.5 mm</quantity>
SL; Borneo: Sabah: tributary of Segama; 
<collectingCountry id="F3697611B340FFD4FDD54714FCC342DB" box="[637,658,1913,1939]" name="Italy" pageId="8" pageNumber="47">Y</collectingCountry>
. 
<collectingCountry id="F3697611B340FFD4FD354714FCE342DB" box="[669,690,1913,1939]" name="Italy" pageId="8" pageNumber="47">Y</collectingCountry>
. Goh, 
<date id="FFC01041B340FFD4FD574714FDB442DB" box="[767,997,1913,1939]" pageId="8" pageNumber="47" value="1998-09-23">23 September 1998</date>
.
</paragraph>
<paragraph id="8BC13681B340FFD4FF6E47F2FEA24296" blockId="8.[151,1437,718,2014]" pageId="8" pageNumber="47">
<taxonomicName id="4C7E4D02B340FFD4FF6E47F2FC5242F0" box="[198,515,1951,1976]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="47" phylum="Chordata" rank="species" species="saravacensis">
<emphasis id="B90AEA93B340FFD4FF6E47F2FC5242F0" box="[198,515,1951,1976]" italics="true" pageId="8" pageNumber="47">Nemacheilus saravacensis</emphasis>
</taxonomicName>
: 
<collectionCode id="ED6FAE44B340FFD4FDB147F3FC2042F0" box="[537,625,1950,1976]" pageId="8" pageNumber="47">BMNH</collectionCode>
1893.3.2.277, 
<typeStatus id="54C58823B340FFD4FC8747F3FDCC42F0" box="[815,925,1950,1976]" pageId="8" pageNumber="47" type="lectotype">lectotype</typeStatus>
, 
<quantity id="4C869B64B340FFD4FC0747F3FA4E42F0" box="[943,1055,1950,1976]" metricMagnitude="-2" metricUnit="m" metricValue="4.21" pageId="8" pageNumber="47" unit="mm" value="42.1">42.1 mm</quantity>
SL; Borneo: 
<collectingCountry id="F3697611B340FFD4FB7947F3FB6A42F0" box="[1233,1339,1950,1976]" name="Malaysia" pageId="8" pageNumber="47">Sarawak</collectingCountry>
: Senah; Everett.
</paragraph>
</subSubSection>
</treatment>
</document>