treatments-xml/data/03/D5/87/03D587E0FF84FFACFE86263D4442FD6A.xml

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<document id="931952C8EF4B995D72501728BD977429" ID-DOI="10.5252/zoosystema2018v40a5" ID-GBIF-Dataset="50513da8-e5ff-4c11-bb0e-3afed74cb0da" ID-ISSN="1638-9387" ID-Zenodo-Dep="3738100" ID-ZooBank="urn:lsid:zoobank.org:pub:4ECDC922-E66F-4C63-A01B-0304C88A9CCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585833082961" checkinUser="felipe" docAuthor="Guinot, Danièle &amp; Segonzac, Michel" docDate="2018" docId="03D587E0FF84FFACFE86263D4442FD6A" docLanguage="en" docName="zoosystema.40.5.75-107.pdf.imf" docOrigin="Zoosystema 40 (5)" docStyle="DocumentStyle:AFAF64E709DD7C8340B70A24A5804402.4:Zoosystema.2018-2019.journal_article" docStyleId="AFAF64E709DD7C8340B70A24A5804402" docStyleName="Zoosystema.2018-2019.journal_article" docStyleVersion="4" docTitle="Austinograea hourdezi Guinot &amp; Segonzac 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="89" masterDocId="FFECFF98FF83FFBCFFAD2465405CFFD8" masterDocTitle="A review of the brachyuran deep-sea vent community of the western Pacific, with two new species of Austinograea Hessler &amp; Martin, 1989 (Crustacea, Decapoda, Brachyura, Bythograeidae) from the Lau and North Fiji Back-Arc Basins" masterLastPageNumber="107" masterPageNumber="75" pageNumber="80" updateTime="1698734985404" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BDA46B96BB72ACFECB3BEADF9E4B7B32">A review of the brachyuran deep-sea vent community of the western Pacific, with two new species of Austinograea Hessler &amp; Martin, 1989 (Crustacea, Decapoda, Brachyura, Bythograeidae) from the Lau and North Fiji Back-Arc Basins</mods:title>
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<mods:namePart id="707AC47A12B660F29C60CD7F23CEA20E">Segonzac, Michel</mods:namePart>
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(
<figureCitation id="13472A73FF84FFBBFEA6261D413DFD4A" box="[267,353,632,658]" captionStart="FIG" captionStartId="8.[132,143,1873,1891]" captionTargetBox="[397,1190,214,1823]" captionTargetId="figure@8.[396,927,687,1161]" captionTargetPageId="8" captionText="FIG. 3. — Austinograea hourdezi n. sp. photographed in situ: A, Lau Basin, Lau Basin 2009 cruise, dive 432, Tow Cam site, 20˚18.98S, 176˚08.19W, 2723 m, 05.VI.2009 (type locality), among sessiles barnacles Eochionelasmus ohtai ohtai Yamaguchi, 1990. The two dark spots on outer surface of palm of both chelae indicate a male; see patches of dense setae on inner surface of both chelae, and the darkly coloured two-thirds of dactylus of right cheliped; B, C, Austinograea hourdezi n. sp., due to the patch of dense setae on inner surface of both chelae: B, Lau Basin, TUIM07MV cruise, dive 163, ABE site, 20°45.65S, 176°11.45W, 2130 m, 23.VI.2005; on a bed of gastropod Ifremeria nautilei Bouchet &amp; Warén, 1991; C, Lau Basin, Lau Basin 2009 cruise, dive 431, ABE site, 20°45.65S, 176°11.45W, 2130 m, 03.VI.2009. The crab and the gastropods Alviniconcha hessleri sensu lato are blurry, the photography in situ having been taken in hydrothermal diffuse area with the emanating fluid visible as shimmering water. Photographies courtesy of C. R. Fisher (Penn State University, USA)." figureDoi="http://doi.org/10.5281/zenodo.3738106" httpUri="https://zenodo.org/record/3738106/files/figure.png" pageId="7" pageNumber="80">Figs 3A</figureCitation>
;
<figureCitation id="13472A73FF84FFBBFEC1261D41F4FD4A" box="[364,424,632,658]" captionStart="FIG" captionStartId="9.[132,143,1920,1938]" captionTargetBox="[132,1457,216,1756]" captionTargetId="figure@9.[699,984,759,1032]" captionTargetPageId="9" captionText="FIG. 4. — A-C, Austinograea hourdezi n. sp., holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, chelae with two coloured spots on outer surface of palm of right crusher (A) and left cutter (B); C, G1 and G2 photographed in situ; D, E, Austinograea hourdezi n. sp., atypical ♀ 30.0 × 48.5 mm (right-handed), Lau Back-Arc Basin, MGLN07MV cruise, dive 230, Kilo Moana site, 20°03.23S, 176°08.01W, 2623 m, MNHN-IU-2016-10752: D, crusher with one marked dark spot and the other one, only discernible by a yellow trace (wound at base of fixed finger is not a spot comparable to that of A. jolliveti n. sp.); E, cutter with two indistinct spots, only as yellow traces." figureDoi="http://doi.org/10.5281/zenodo.3738108" httpUri="https://zenodo.org/record/3738108/files/figure.png" pageId="7" pageNumber="80">4A-E</figureCitation>
;
<figureCitation id="13472A73FF84FFBBFE1E261D41A8FD4B" box="[435,500,632,659]" captionStart="FIG" captionStartId="10.[132,143,1965,1983]" captionTargetBox="[132,1455,216,1859]" captionTargetId="figure@10.[698,1267,763,1313]" captionTargetPageId="10" captionText="FIG. 5. — Austinograea hourdezi n. sp., paratype, ♂ 29.8 × 48.1 mm (left-handed, with weak heterochely and heterodonty), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738: A, dorsal view; B, frontal view; C, mxp3; D, ventral view; E, G, both chelae with two typical spots on outer surface; F, H, both chelae with patches of setae on inner surface: E, F, cutter situated at right; G, H, crusher situated at left." figureDoi="http://doi.org/10.5281/zenodo.3738110" httpUri="https://zenodo.org/record/3738110/files/figure.png" pageId="7" pageNumber="80">5A-H</figureCitation>
;
<figureCitation id="13472A73FF84FFBBFDAD261D4267FD4A" box="[512,571,632,658]" captionStart="FIG" captionStartId="12.[132,143,1942,1960]" captionTargetBox="[168,1419,251,1804]" captionTargetId="figure@12.[132,1455,215,1860]" captionTargetPageId="12" captionText="FIG. 6. — Austinograea hourdezi n. sp.: A-C, holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, right and left mxp3, with and without ornamentation; C, thoracic sternum. D, E, paratype, ♂ 29.8 × 48.1 mm (left-handed), Lau Back-Arc Basin,TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738: D, G1, two views;E, G2, two views. Scale bars: A, B, D, E, 1 mm; C, 5 mm. Note the scale bars of the two gonopods, the G2 being about half-length of G1." figureDoi="http://doi.org/10.5281/zenodo.3738112" httpUri="https://zenodo.org/record/3738112/files/figure.png" pageId="7" pageNumber="80">6A-E</figureCitation>
;
<figureCitation id="13472A73FF84FFBBFDEB261D42DBFD4A" box="[582,647,632,658]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1858]" captionTargetId="figure@14.[699,984,1036,1311]" captionTargetPageId="14" captionText="FIG. 7. — Austinograea hourdezi n. sp., paratype, ♀ 18.3 × 28.4 mm (right-handed), Lau Back-Arc Basin, BIOLAU cruise, BL 12, Vai Lili site, 23°13S, 176°38W, 1750 m, MNHN-IU-2016-10745 (= MNHN-B24060): A, dorsal view; B, C, ventral views, with pleon; D, thoracic sternum and vulvae; E, G, both chelae without spots on outer surface; F, H, both chelae with patches of setae on inner surface; E, F, right crusher; G, H, left cutter." figureDoi="http://doi.org/10.5281/zenodo.3738114" httpUri="https://zenodo.org/record/3738114/files/figure.png" pageId="7" pageNumber="80">7A-H</figureCitation>
)
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<emphasis id="B908EAE4FF84FFBBFF2926DF40A0FD09" box="[132,252,698,721]" italics="true" pageId="7" pageNumber="80">Austinograea</emphasis>
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sp.
<taxonomicName id="4C7C4D75FF84FFBBFE8926DF428DFD0A" ID-CoL="5W9JY" authority="Guinot, 1990: 898 (Addenda)" authorityName="in Guinot" authorityYear="1990" box="[292,721,698,722]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="williamsi">
aff.
<emphasis id="B908EAE4FF84FFBBFEE426DF41FCFD0A" box="[329,416,698,722]" italics="true" pageId="7" pageNumber="80">williamsi</emphasis>
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<bibRefCitation id="EFED4B07FF84FFBBFE0B26DF4203FD0A" author="GUINOT D." box="[422,607,698,722]" pageId="7" pageNumber="80" pagination="879 - 903" refId="ref26108" refString="GUINOT D. 1990. - Austinograea alayseae sp. nov., crabe hydrothermal decouvert dans le bassin de Lau, Pacifique sud-occidental (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle 4 e ser., 11, section A (4): 879 - 903." type="journal article" year="1990">Guinot, 1990: 898</bibRefCitation>
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(Addenda)
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, 900, 901. —
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, fig. 1. —
<treatmentCitation id="0ADD10E7FF84FFBBFD2526B0408DFCDF" pageId="7" pageNumber="80">
<bibRefCitation id="EFED4B07FF84FFBBFD2526B0408DFCDF" author="MATEOS M. &amp; HURTADO L. A. &amp; SANTAMARIA C. A. &amp; LEIGNEL V. &amp; GUINOT D." pageId="7" pageNumber="80" refId="ref29119" refString="MATEOS M., HURTADO L. A., SANTAMARIA C. A., LEIGNEL V. &amp; GUINOT D. 2012. - Molecular systematics of the deep-sea hydrothermal vent endemic brachyuran family Bythograeidae Williams, 1980: a comparison of three Bayesian species-tree methods. PloS ONE 7 (3): e 32066, 14 p. https: // doi. org / 10.1371 / journal. pone. 0032066" type="book" year="2012">
Mateos
<emphasis id="B908EAE4FF84FFBBFD7A26B04354FD34" box="[727,776,724,748]" italics="true" pageId="7" pageNumber="80">et al.</emphasis>
2012: 1
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, 5, 6, 10, figs 1, 2, table 1.
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MATERIAL. —
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25.7 ×
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: right-handed individual, with left cheliped and all P2- P5 detached (
<figureCitation id="13472A73FF84FFBBFEA427F4412DFC71" box="[265,369,913,937]" captionStart="FIG" captionStartId="9.[132,143,1920,1938]" captionTargetBox="[132,1457,216,1756]" captionTargetId="figure@9.[699,984,759,1032]" captionTargetPageId="9" captionText="FIG. 4. — A-C, Austinograea hourdezi n. sp., holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, chelae with two coloured spots on outer surface of palm of right crusher (A) and left cutter (B); C, G1 and G2 photographed in situ; D, E, Austinograea hourdezi n. sp., atypical ♀ 30.0 × 48.5 mm (right-handed), Lau Back-Arc Basin, MGLN07MV cruise, dive 230, Kilo Moana site, 20°03.23S, 176°08.01W, 2623 m, MNHN-IU-2016-10752: D, crusher with one marked dark spot and the other one, only discernible by a yellow trace (wound at base of fixed finger is not a spot comparable to that of A. jolliveti n. sp.); E, cutter with two indistinct spots, only as yellow traces." figureDoi="http://doi.org/10.5281/zenodo.3738108" httpUri="https://zenodo.org/record/3738108/files/figure.png" pageId="7" pageNumber="80">Figs 4A-C</figureCitation>
;
<figureCitation id="13472A73FF84FFBBFED627F441EEFC71" box="[379,434,913,937]" captionStart="FIG" captionStartId="12.[132,143,1942,1960]" captionTargetBox="[168,1419,251,1804]" captionTargetId="figure@12.[132,1455,215,1860]" captionTargetPageId="12" captionText="FIG. 6. — Austinograea hourdezi n. sp.: A-C, holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, right and left mxp3, with and without ornamentation; C, thoracic sternum. D, E, paratype, ♂ 29.8 × 48.1 mm (left-handed), Lau Back-Arc Basin,TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738: D, G1, two views;E, G2, two views. Scale bars: A, B, D, E, 1 mm; C, 5 mm. Note the scale bars of the two gonopods, the G2 being about half-length of G1." figureDoi="http://doi.org/10.5281/zenodo.3738112" httpUri="https://zenodo.org/record/3738112/files/figure.png" pageId="7" pageNumber="80">6A-E</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC336F6FF84FFBBFF2927CE4110F8CA" blockId="7.[132,776,807,1810]" pageId="7" pageNumber="80">
<materialsCitation id="3B143CABFF84FFBBFF2927CE42DAFBCB" ID-GBIF-Occurrence="2594759999" collectionCode="MNHN-IU-" country="Fiji" pageId="7" pageNumber="80" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B908EAE4FF84FFBBFF2927CE40BCFC1B" bold="true" box="[132,224,939,963]" pageId="7" pageNumber="80">
<typeStatus id="54C78854FF84FFBBFF2927CE40BCFC1B" box="[132,224,939,963]" pageId="7" pageNumber="80" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectionCode id="ED6DAE33FF84FFBBFF4627CE413AFC1B" box="[235,358,939,963]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10738, same data,
<specimenCount id="9D7AFD7FFF84FFBBFDE727C94231FC1D" box="[586,621,940,965]" pageId="7" pageNumber="80" type="male">1 ♂</specimenCount>
29.8 ×
<quantity id="4C849B13FF84FFBBFD1C27C9435BFC1B" box="[689,775,940,964]" metricMagnitude="-2" metricUnit="m" metricValue="4.81" pageId="7" pageNumber="80" unit="mm" value="48.1">48.1 mm</quantity>
(left-handed, two spots on both chelae, with weak heterochely and heterodonty) (
<figureCitation id="13472A73FF84FFBBFEBD27854124FC20" box="[272,376,992,1016]" captionStart="FIG" captionStartId="10.[132,143,1965,1983]" captionTargetBox="[132,1455,216,1859]" captionTargetId="figure@10.[698,1267,763,1313]" captionTargetPageId="10" captionText="FIG. 5. — Austinograea hourdezi n. sp., paratype, ♂ 29.8 × 48.1 mm (left-handed, with weak heterochely and heterodonty), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738: A, dorsal view; B, frontal view; C, mxp3; D, ventral view; E, G, both chelae with two typical spots on outer surface; F, H, both chelae with patches of setae on inner surface: E, F, cutter situated at right; G, H, crusher situated at left." figureDoi="http://doi.org/10.5281/zenodo.3738110" httpUri="https://zenodo.org/record/3738110/files/figure.png" pageId="7" pageNumber="80">Fig. 5A-H</figureCitation>
),
<specimenCount id="9D7AFD7FFF84FFBBFE24278441F1FC22" box="[393,429,993,1018]" pageId="7" pageNumber="80" type="male">1 ♂</specimenCount>
24.7 × 36.0 mm (left-handed, with small regenerated right chela, bearing only one spot)
</materialsCitation>
.
<materialsCitation id="3B143CABFF84FFBBFF2920704358FB4F" ID-GBIF-Occurrence="2594759992" collectingDate="1989-05-24" collectionCode="MNHN-IU-" country="Fiji" latitude="-23.216667" location="Vai Lili" longLatPrecision="1256" longitude="-176.63333" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="1758" specimenCount-female="5" specimenCount-male="1753">
<collectionCode id="ED6DAE33FF84FFBBFF29207040A2FBF5" box="[132,254,1045,1069]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10745 (= MNHN-B24060),
<specimenCount id="9D7AFD7FFF84FFBBFDEB20704235FBF7" box="[582,617,1045,1071]" pageId="7" pageNumber="80" type="male">3 ♂</specimenCount>
21.0 ×
<quantity id="4C849B13FF84FFBBFD062070435FFBF5" box="[683,771,1045,1069]" metricMagnitude="-2" metricUnit="m" metricValue="3.28" pageId="7" pageNumber="80" unit="mm" value="32.8">32.8 mm</quantity>
, 18.7 × 28.0 mm, 18.3 ×
<quantity id="4C849B13FF84FFBBFED120554186FB9F" box="[380,474,1072,1096]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="7" pageNumber="80" unit="mm" value="29.2">29.2 mm</quantity>
,
<specimenCount id="9D7AFD7FFF84FFBBFE482055425AFB91" box="[485,518,1072,1097]" pageId="7" pageNumber="80" type="female">5 ♀</specimenCount>
25.0 × 40.0 mm (two P1 lacking), 19.1 ×
<quantity id="4C849B13FF84FFBBFEB4202F412EFBBA" box="[281,370,1098,1122]" metricMagnitude="-2" metricUnit="m" metricValue="3.22" pageId="7" pageNumber="80" unit="mm" value="32.2">32.2 mm</quantity>
, 18.3 ×
<quantity id="4C849B13FF84FFBBFE17202F424EFBBA" box="[442,530,1098,1122]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="7" pageNumber="80" unit="mm" value="28.2">28.2 mm</quantity>
, 18.3 ×
<quantity id="4C849B13FF84FFBBFDF7202F42ECFBBA" box="[602,688,1098,1122]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="7" pageNumber="80" unit="mm" value="28.4">28.4 mm</quantity>
(
<figureCitation id="13472A73FF84FFBBFD16202F40C6FBA5" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1858]" captionTargetId="figure@14.[699,984,1036,1311]" captionTargetPageId="14" captionText="FIG. 7. — Austinograea hourdezi n. sp., paratype, ♀ 18.3 × 28.4 mm (right-handed), Lau Back-Arc Basin, BIOLAU cruise, BL 12, Vai Lili site, 23°13S, 176°38W, 1750 m, MNHN-IU-2016-10745 (= MNHN-B24060): A, dorsal view; B, C, ventral views, with pleon; D, thoracic sternum and vulvae; E, G, both chelae without spots on outer surface; F, H, both chelae with patches of setae on inner surface; E, F, right crusher; G, H, left cutter." figureDoi="http://doi.org/10.5281/zenodo.3738114" httpUri="https://zenodo.org/record/3738114/files/figure.png" pageId="7" pageNumber="80">Fig. 7A- H</figureCitation>
), cl
<quantity id="4C849B13FF84FFBBFF6520004178FBA4" box="[200,292,1125,1149]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="7" pageNumber="80" unit="mm" value="19.1">19.1 mm</quantity>
(bad condition),
<collectingMunicipality id="6BA7AC8CFF84FFBBFE7A200042C2FBA5" box="[471,670,1125,1149]" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
, BIOLAU cruise, BL 12,
<location id="8EA3602DFF84FFBBFEA0201A410EFB4F" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFEA0201A410EFB4F" box="[269,338,1151,1175]" country="Fiji" latitude="-23.216667" longLatPrecision="1256" longitude="-176.63333" municipality="Lau Back-Arc Basin" name="Vai Lili" pageId="7" pageNumber="80">Vai Lili</location>
site,
<geoCoordinate id="EE485031FF84FFBBFE2F201A418CFB4F" box="[386,464,1151,1175]" degrees="23" direction="south" minutes="13" orientation="latitude" pageId="7" pageNumber="80" precision="925" value="-23.216667">23°13S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFE74201A4261FB4F" box="[473,573,1151,1175]" degrees="176" direction="west" minutes="38" orientation="longitude" pageId="7" pageNumber="80" precision="925" value="-176.63333">176°38W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFDE8201A42CEFB4F" box="[581,658,1151,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="7" pageNumber="80" unit="m" value="1750.0">
<specimenCount id="9D7AFD7FFF84FFBBFDE8201A42CEFB4F" box="[581,658,1151,1175]" pageId="7" pageNumber="80" type="male">1750 m</specimenCount>
</quantity>
,
<date id="FFC21036FF84FFBBFD36201A4358FB4F" box="[667,772,1151,1175]" pageId="7" pageNumber="80" value="1989-05-24">
<collectingDate id="EF86E9DEFF84FFBBFD36201A4358FB4F" box="[667,772,1151,1175]" pageId="7" pageNumber="80" value="1989-05-24">24.V.1989</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B143CABFF84FFBBFF2920FF40B2FAD9" ID-GBIF-Occurrence="2594759985" collectingDate="1989-05-15" country="Fiji" elevation="1853" latitude="-22.533333" location="Hine Hina" longLatPrecision="1259" longitude="-176.71666" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
MNHN-IU-2016-10747 (= MNHN-B24059),
<specimenCount id="9D7AFD7FFF84FFBBFDEB20FF4235FB6B" box="[582,617,1178,1203]" pageId="7" pageNumber="80" type="male">2 ♂</specimenCount>
18.0 × 31.0 mm, 17.0 × 28.0 mm,
<specimenCount id="9D7AFD7FFF84FFBBFE8820D14119FB16" box="[293,325,1204,1230]" pageId="7" pageNumber="80" type="female">1 ♀</specimenCount>
16.4 ×
<quantity id="4C849B13FF84FFBBFE2720D141BEFB14" box="[394,482,1204,1228]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="7" pageNumber="80" unit="mm" value="26.8">26.8 mm</quantity>
,
<collectingMunicipality id="6BA7AC8CFF84FFBBFE4620D042F8FB14" box="[491,676,1204,1228]" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
, BIOLAU cruise, dive BL 03,
<location id="8EA3602DFF84FFBBFEEB20AB41EEFB3E" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFEEB20AB41EEFB3E" box="[326,434,1230,1254]" country="Fiji" latitude="-22.533333" longLatPrecision="1259" longitude="-176.71666" municipality="Lau Back-Arc Basin" name="Hine Hina" pageId="7" pageNumber="80">Hine Hina</location>
site,
<geoCoordinate id="EE485031FF84FFBBFE4520AA426BFB3F" box="[488,567,1231,1255]" degrees="22" direction="south" minutes="32" orientation="latitude" pageId="7" pageNumber="80" precision="925" value="-22.533333">22°32S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFDEF20AB42FBFB3E" box="[578,679,1230,1254]" degrees="176" direction="west" minutes="43" orientation="longitude" pageId="7" pageNumber="80" precision="925" value="-176.71666">176°43W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFD1F20AB435FFB3E" box="[690,771,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.853" pageId="7" pageNumber="80" unit="m" value="1853.0">
<elevation id="0051D1C5FF84FFBBFD1F20AB435FFB3E" box="[690,771,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.853" pageId="7" pageNumber="80" unit="m" value="1853.0">1853 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFF29208C40B2FAD9" box="[132,238,1257,1281]" pageId="7" pageNumber="80" value="1989-05-15">
<collectingDate id="EF86E9DEFF84FFBBFF29208C40B2FAD9" box="[132,238,1257,1281]" pageId="7" pageNumber="80" value="1989-05-15">15.V.1989</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B143CABFF84FFBBFF29216642DBFA88" ID-GBIF-Occurrence="2594759988" collectingDate="2009-09-07" collectionCode="MNHN-IU-" country="Fiji" elevation="2623" latitude="-20.053833" location="Lau Basin" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="1" specimenCount-female="1">
<collectionCode id="ED6DAE33FF84FFBBFF292166415FFAC3" box="[132,259,1283,1307]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10739,
<specimenCount id="9D7AFD7FFF84FFBBFE24216141F6FAC5" box="[393,426,1284,1309]" pageId="7" pageNumber="80" type="female">1 ♀</specimenCount>
(with two cutters) 28.4 ×
<quantity id="4C849B13FF84FFBBFD0A2166435FFAC3" box="[679,771,1283,1307]" metricMagnitude="-2" metricUnit="m" metricValue="4.68" pageId="7" pageNumber="80" unit="mm" value="46.8">46.8 mm</quantity>
,
<collectingMunicipality id="6BA7AC8CFF84FFBBFF29217B411BFAEE" box="[132,327,1310,1334]" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
,
<location id="8EA3602DFF84FFBBFEFD217B41ECFAEE" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFEFD217B41ECFAEE" box="[336,432,1310,1334]" country="Fiji" latitude="-20.053833" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" name="Lau Basin" pageId="7" pageNumber="80">Lau Basin</location>
2009 cruise, dive 424,
<location id="8EA3602DFF84FFBBFD3F217B435BFAED" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFD3F217B435BFAED" box="[658,775,1310,1334]" country="Fiji" latitude="-20.053833" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" name="Kilo Moana" pageId="7" pageNumber="80">Kilo Moana</location>
site,
<geoCoordinate id="EE485031FF84FFBBFF1C215D417DFA88" box="[177,289,1336,1360]" degrees="20" direction="south" minutes="03.23" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-20.053833">20°03.23S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFE87215D41F2FA88" box="[298,430,1336,1360]" degrees="176" direction="west" minutes="08.01" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.1335">176°08.01W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFE14215D4254FA88" box="[441,520,1336,1360]" metricMagnitude="3" metricUnit="m" metricValue="2.623" pageId="7" pageNumber="80" unit="m" value="2623.0">
<elevation id="0051D1C5FF84FFBBFE14215D4254FA88" box="[441,520,1336,1360]" metricMagnitude="3" metricUnit="m" metricValue="2.623" pageId="7" pageNumber="80" unit="m" value="2623.0">2623 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFDBE215D42DBFA88" box="[531,647,1336,1360]" pageId="7" pageNumber="80" value="2009-09-07">
<collectingDate id="EF86E9DEFF84FFBBFDBE215D42DBFA88" box="[531,647,1336,1360]" pageId="7" pageNumber="80" value="2009-09-07">07.IX.2009</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B143CABFF84FFBBFF292136418AFA78" ID-GBIF-Occurrence="2594759993" collectingDate="2005-05-17" collectionCode="MNHN-IU-" country="Fiji" latitude="-20.053833" location="Kilo Moana" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCode="TUIM06" specimenCount="2624" specimenCount-female="1" specimenCount-male="2623">
<collectionCode id="ED6DAE33FF84FFBBFF2921364158FAB3" box="[132,260,1363,1387]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10741,
<specimenCount id="9D7AFD7FFF84FFBBFE24213641F6FAB4" box="[393,426,1363,1388]" pageId="7" pageNumber="80" type="female">1 ♀</specimenCount>
25.3 × 40.0 mm,
<collectingMunicipality id="6BA7AC8CFF84FFBBFDF6213640FFFA5D" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
,
<specimenCode id="DBDA9E8DFF84FFBBFF0321084155FA5D" box="[174,265,1389,1413]" pageId="7" pageNumber="80">TUIM06</specimenCode>
MV cruise, dive 140,
<location id="8EA3602DFF84FFBBFE4B21084201FA5D" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFE4B21084201FA5D" box="[486,605,1389,1413]" country="Fiji" latitude="-20.053833" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" name="Kilo Moana" pageId="7" pageNumber="80">Kilo Moana</location>
site,
<geoCoordinate id="EE485031FF84FFBBFD3921084358FA5D" box="[660,772,1389,1413]" degrees="20" direction="south" minutes="03.23" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-20.053833">20°03.23S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFF2921ED4154FA78" box="[132,264,1416,1440]" degrees="176" direction="west" minutes="08.01" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.1335">176°08.01W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFEBE21ED413EFA47" box="[275,354,1416,1440]" metricMagnitude="3" metricUnit="m" metricValue="2.623" pageId="7" pageNumber="80" unit="m" value="2623.0">
<specimenCount id="9D7AFD7FFF84FFBBFEBE21ED413EFA47" box="[275,354,1416,1440]" pageId="7" pageNumber="80" type="male">2623 m</specimenCount>
</quantity>
,
<date id="FFC21036FF84FFBBFEC121ED418AFA78" box="[364,470,1416,1440]" pageId="7" pageNumber="80" value="2005-05-17">
<collectingDate id="EF86E9DEFF84FFBBFEC121ED418AFA78" box="[364,470,1416,1440]" pageId="7" pageNumber="80" value="2005-05-17">17.V.2005</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B143CABFF84FFBBFF2921C742AEFA37" ID-GBIF-Occurrence="2594760000" collectingDate="2006-09-11" collectionCode="MNHN-IU-" country="Fiji" elevation="1891" latitude="-21.989" location="Malila" longLatPrecision="12" longitude="-176.56816" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="1" specimenCount-male="1">
<collectionCode id="ED6DAE33FF84FFBBFF2921C7415AFA62" box="[132,262,1442,1466]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10740,
<specimenCount id="9D7AFD7FFF84FFBBFE3921C741E5FA64" box="[404,441,1442,1468]" pageId="7" pageNumber="80" type="male">1 ♂</specimenCount>
21.4 ×
<quantity id="4C849B13FF84FFBBFDA021C74233FA62" box="[525,623,1442,1466]" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" pageId="7" pageNumber="80" unit="mm" value="34.3">34.3 mm</quantity>
,
<collectingMunicipality id="6BA7AC8CFF84FFBBFDD621C640E0FA0D" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
, MGLN07MV cruise, dive 232, Tui
<location id="8EA3602DFF84FFBBFD8821D8423FFA0D" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFD8821D8423FFA0D" box="[549,611,1469,1493]" country="Fiji" latitude="-21.989" longLatPrecision="12" longitude="-176.56816" municipality="Lau Back-Arc Basin" name="Malila" pageId="7" pageNumber="80">Malila</location>
site,
<geoCoordinate id="EE485031FF84FFBBFD3921D9435FFA0C" box="[660,771,1468,1492]" degrees="21" direction="south" minutes="59.34" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-21.989">21°59.34S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFF2921B24155FA37" box="[132,265,1495,1519]" degrees="176" direction="west" minutes="34.09" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.56816">176°34.09W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFEBE21B2413EFA37" box="[275,354,1495,1519]" metricMagnitude="3" metricUnit="m" metricValue="1.891" pageId="7" pageNumber="80" unit="m" value="1891.0">
<elevation id="0051D1C5FF84FFBBFEBE21B2413EFA37" box="[275,354,1495,1519]" metricMagnitude="3" metricUnit="m" metricValue="1.891" pageId="7" pageNumber="80" unit="m" value="1891.0">1891 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFEC121B24182FA37" box="[364,478,1495,1519]" pageId="7" pageNumber="80" value="2006-09-11">
<collectingDate id="EF86E9DEFF84FFBBFEC121B24182FA37" box="[364,478,1495,1519]" pageId="7" pageNumber="80" value="2006-09-11">11.IX.2006</collectingDate>
</date>
(preserved in ethanol 85%)
</materialsCitation>
.
<materialsCitation id="3B143CABFF84FFBBFF2921944261F981" ID-GBIF-Occurrence="2594759990" collectingDate="2006-09-09" collectionCode="MNHN-IU-" country="Fiji" elevation="2130" latitude="-20.760834" location="Lau Back-Arc Basin" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="1" specimenCount-male="1">
<collectionCode id="ED6DAE33FF84FFBBFF292194415FF9D1" box="[132,259,1521,1545]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10743, 1 big damaged
<specimenCount id="9D7AFD7FFF84FFBBFD8C21914269F9D3" box="[545,565,1524,1547]" pageId="7" pageNumber="80" type="male"></specimenCount>
(with detached legs),
<collectingMunicipality id="6BA7AC8CFF84FFBBFF292268410EF9FC" box="[132,338,1548,1572]" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
, MGLN07MV cruise, dive 231, ABE site,
<geoCoordinate id="EE485031FF84FFBBFF29224340AFF9E6" box="[132,243,1574,1598]" degrees="20" direction="south" minutes="45.65" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-20.760834">20°45.65S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFF50224341DEF9E6" box="[253,386,1574,1598]" degrees="176" direction="west" minutes="11.45" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.19083">176°11.45W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFE2122424187F9E6" box="[396,475,1575,1599]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="80" unit="m" value="2130.0">
<elevation id="0051D1C5FF84FFBBFE2122424187F9E6" box="[396,475,1575,1599]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="80" unit="m" value="2130.0">2130 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFE4B22434204F9E6" box="[486,600,1574,1598]" pageId="7" pageNumber="80" value="2006-09-09">
<collectingDate id="EF86E9DEFF84FFBBFE4B22434204F9E6" box="[486,600,1574,1598]" pageId="7" pageNumber="80" value="2006-09-09">09.IX.2006</collectingDate>
</date>
(preserved in formalin 10% then transferred to ethanol 70%)
</materialsCitation>
.
<materialsCitation id="3B143CABFF84FFBBFF29223E4358F970" ID-GBIF-Occurrence="2594759986" collectingDate="1989-07-11" collectionCode="MNHN-IU-" country="Fiji" elevation="2000" latitude="-16.991667" location="White Lady" longLatPrecision="13" longitude="173.9245" pageId="7" pageNumber="80" specimenCount="1" specimenCount-female="1" stateProvince="North Fiji Basin">
<collectionCode id="ED6DAE33FF84FFBBFF29223E40A3F9AB" box="[132,255,1627,1651]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10746 (= MNHN-B27831),
<specimenCount id="9D7AFD7FFF84FFBBFDE122394231F9AD" box="[588,621,1628,1653]" pageId="7" pageNumber="80" type="female">1 ♀</specimenCount>
cl
<quantity id="4C849B13FF84FFBBFD2822394280F9AB" box="[645,732,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="7" pageNumber="80" unit="mm" value="28.2">28.2 mm</quantity>
(left part broken),
<collectingRegion id="49B8F814FF84FFBBFEA3221341EDF956" box="[270,433,1654,1678]" pageId="7" pageNumber="80">North Fiji Basin</collectingRegion>
, STARMER II cruise, dive PL 16,
<location id="8EA3602DFF84FFBBFF2922F540A4F970" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFF2922F540A4F970" box="[132,248,1680,1704]" country="Fiji" latitude="-16.991667" longLatPrecision="13" longitude="173.9245" name="White Lady" pageId="7" pageNumber="80" stateProvince="North Fiji Basin">White Lady</location>
site,
<geoCoordinate id="EE485031FF84FFBBFE8122F541C7F970" box="[300,411,1680,1704]" degrees="16" direction="south" minutes="59.50" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-16.991667">16°59.50S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFE0922F5427EF970" box="[420,546,1680,1704]" degrees="173" direction="east" minutes="55.47" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="173.9245">173°55.47E</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFD8122F54227F970" box="[556,635,1680,1704]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="80" unit="m" value="2000.0">
<elevation id="0051D1C5FF84FFBBFD8122F54227F970" box="[556,635,1680,1704]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="80" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFD2B22F54358F970" box="[646,772,1680,1704]" pageId="7" pageNumber="80" value="1989-07-11">
<collectingDate id="EF86E9DEFF84FFBBFD2B22F54358F970" box="[646,772,1680,1704]" pageId="7" pageNumber="80" value="1989-07-11">11.VII.1989</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B143CABFF84FFBBFF2922CE4114F8CA" ID-GBIF-Occurrence="2594759989" collectingDate="2005-05-30" collectionCode="MNHN-IU-" country="Fiji" elevation="2000" latitude="-16.991667" location="White Lady" longLatPrecision="13" longitude="173.9245" municipality="White Lady" pageId="7" pageNumber="80" specimenCount="4" specimenCount-female="4" stateProvince="North Fiji Basin">
<collectionCode id="ED6DAE33FF84FFBBFF2922CE415EF91B" box="[132,258,1707,1731]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10742,
<specimenCount id="9D7AFD7FFF84FFBBFE2722CE41F7F91C" box="[394,427,1707,1732]" pageId="7" pageNumber="80" type="female">4 ♀</specimenCount>
19.4 ×
<quantity id="4C849B13FF84FFBBFE5522CE4209F91A" box="[504,597,1707,1731]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" pageId="7" pageNumber="80" unit="mm" value="39.3">39.3 mm</quantity>
, 20.5 ×
<quantity id="4C849B13FF84FFBBFD0B22CE435FF91A" box="[678,771,1707,1731]" metricMagnitude="-2" metricUnit="m" metricValue="3.32" pageId="7" pageNumber="80" unit="mm" value="33.2">33.2 mm</quantity>
, 19.0 × 30.0 mm, 25.0 × 40.0 mm, TUIM06MV cruise, dive 151, box 2,
<collectingRegion id="49B8F814FF84FFBBFF682285413FF920" box="[197,355,1760,1784]" pageId="7" pageNumber="80">North Fiji Basin</collectingRegion>
,
<collectingMunicipality id="6BA7AC8CFF84FFBBFEC622854181F920" box="[363,477,1760,1784]" pageId="7" pageNumber="80">White Lady</collectingMunicipality>
site,
<geoCoordinate id="EE485031FF84FFBBFDA322BA4221F92F" box="[526,637,1759,1783]" degrees="16" direction="south" minutes="59.50" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-16.991667">16°59.50S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFD2822BA435FF92F" box="[645,771,1759,1783]" degrees="173" direction="east" minutes="55.47" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="173.9245">173°55.47E</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFF29229F408FF8CA" box="[132,211,1786,1810]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="80" unit="m" value="2000.0">
<elevation id="0051D1C5FF84FFBBFF29229F408FF8CA" box="[132,211,1786,1810]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="80" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFF73229F4114F8CA" box="[222,328,1786,1810]" pageId="7" pageNumber="80" value="2005-05-30">
<collectingDate id="EF86E9DEFF84FFBBFF73229F4114F8CA" box="[222,328,1786,1810]" pageId="7" pageNumber="80" value="2005-05-30">30.V.2005</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C366657DFF84FFBBFF292357421FF8BD" pageId="7" pageNumber="80" type="distribution">
<paragraph id="8BC336F6FF84FFBBFF292357421FF8BD" blockId="7.[132,774,1842,1893]" pageId="7" pageNumber="80">
<typeStatus id="54C78854FF84FFBBFF29235740E9F890" box="[132,181,1842,1866]" pageId="7" pageNumber="80">TYPE</typeStatus>
LOCALITY. — Southwestern Pacific, Lau Back-Arc Basin, Tow Cam site,
<geoCoordinate id="EE485031FF84FFBBFF452328410BF8BD" box="[232,343,1869,1893]" degrees="20" direction="south" minutes="19.07" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-20.317833">20°19.07S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFECC232841B9F8BD" box="[353,485,1869,1893]" degrees="176" direction="west" minutes="08.24" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.13733">176°08.24W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFE4223284263F8BD" box="[495,575,1869,1893]" metricMagnitude="3" metricUnit="m" metricValue="2.719" pageId="7" pageNumber="80" unit="m" value="2719.0">2719 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C366657DFF84FFBBFF2923E14462FBD2" pageId="7" pageNumber="80" type="materials_examined">
<paragraph id="8BC336F6FF84FFBBFF2923E1413CF833" blockId="7.[132,775,1924,2028]" pageId="7" pageNumber="80">
OTHER MATERIAL EXAMINED. —
<materialsCitation id="3B143CABFF84FFBBFE0723E14100F833" ID-GBIF-Occurrence="2594760005" collectionCode="MNHN-IU-" country="Fiji" elevation="2000" latitude="-16.991667" location="White Lady" longLatPrecision="13" longitude="173.9245" municipality="White Lady" pageId="7" pageNumber="80" specimenCount="1" specimenCount-female="1" stateProvince="North Fiji Basin">
<collectionCode id="ED6DAE33FF84FFBBFE0723E1427EF844" box="[426,546,1924,1948]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10748 (= MNHN- B27827),
<specimenCount id="9D7AFD7FFF84FFBBFF4A23FA4154F860" box="[231,264,1951,1976]" pageId="7" pageNumber="80" type="female">1 ♀</specimenCount>
cl 29.2 (damaged, left P1 lacking),
<collectingRegion id="49B8F814FF84FFBBFDCE23FA435FF86F" box="[611,771,1951,1975]" pageId="7" pageNumber="80">North Fiji Basin</collectingRegion>
, STARMER II cruise, dive PL 20,
<collectingMunicipality id="6BA7AC8CFF84FFBBFE4C23DC4205F809" box="[481,601,1977,2001]" pageId="7" pageNumber="80">White Lady</collectingMunicipality>
site,
<geoCoordinate id="EE485031FF84FFBBFD3F23DC435FF809" box="[658,771,1977,2001]" degrees="16" direction="south" minutes="59.50" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-16.991667">16°59.50S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFF2923B1415EF834" box="[132,258,2004,2028]" degrees="173" direction="east" minutes="55.47" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="173.9245">173°55.47E</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFEA123B14100F833" box="[268,348,2004,2028]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="80" unit="m" value="2000.0">
<elevation id="0051D1C5FF84FFBBFEA123B14100F833" box="[268,348,2004,2028]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="80" unit="m" value="2000.0">2000 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC336F6FF84FFBBFC8024BC4462FBD2" blockId="7.[813,1456,217,1034]" pageId="7" pageNumber="80">
<materialsCitation id="3B143CABFF84FFBBFC8024BC4385FE98" ID-GBIF-Occurrence="2594759984" collectingDate="2009-05-28" collectionCode="MNHN-IU-" country="Fiji" elevation="2130" latitude="-20.760834" location="Lau Basin" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="2" specimenCount-female="2">
<collectionCode id="ED6DAE33FF84FFBBFC8024BC43F3FF29" box="[813,943,217,241]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10744,
<specimenCount id="9D7AFD7FFF84FFBBFB9524BC4406FF2A" box="[1080,1114,217,242]" pageId="7" pageNumber="80" type="female">2 ♀</specimenCount>
26.0 × 42.0 mm, 26.0 × 40.0 mm,
<collectingMunicipality id="6BA7AC8CFF84FFBBFC8024914459FED3" box="[813,1029,243,267]" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
,
<location id="8EA3602DFF84FFBBFBBC24914420FED3" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFBBC24914420FED3" box="[1041,1148,243,267]" country="Fiji" latitude="-20.760834" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" name="Lau Basin" pageId="7" pageNumber="80">Lau Basin</location>
2009 cruise, dive 427, ABE site,
<geoCoordinate id="EE485031FF84FFBBFCF02568438DFEFD" box="[861,977,269,293]" degrees="20" direction="south" minutes="45.65" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-20.760834">20°45.65S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFC702568443BFEFD" box="[989,1127,269,293]" degrees="176" direction="west" minutes="11.45" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.19083">176°11.45W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFBDE256B449BFEFD" box="[1139,1223,270,294]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="80" unit="m" value="2130.0">
<elevation id="0051D1C5FF84FFBBFBDE256B449BFEFD" box="[1139,1223,270,294]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="80" unit="m" value="2130.0">2130 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFB79256B4563FEFE" box="[1236,1343,270,294]" pageId="7" pageNumber="80" value="2009-05-28">
<collectingDate id="EF86E9DEFF84FFBBFB79256B4563FEFE" box="[1236,1343,270,294]" pageId="7" pageNumber="80" value="2009-05-28">28.V.2009</collectingDate>
</date>
(preserved in ethanol 85%)
</materialsCitation>
. This sample with two typical females devoid of spots on chelae palm also contains a female 27.1 ×
<quantity id="4C849B13FF84FFBBFAFC252745F3FE82" box="[1361,1455,322,346]" metricMagnitude="-2" metricUnit="m" metricValue="4.46" pageId="7" pageNumber="80" unit="mm" value="44.6">44.6 mm</quantity>
with a spot (a unique spot) on each chela (see
<emphasis id="B908EAE4FF84FFBBFAA125384385FE57" italics="true" pageId="7" pageNumber="80">Individuals with regenerated chelae</emphasis>
).
<materialsCitation id="3B143CABFF84FFBBFC8025F74421FE21" ID-GBIF-Occurrence="2594760001" collectingDate="2006-04-07" collectionCode="MNHN-IU-" country="Fiji" elevation="2623" latitude="-20.053833" location="Kilo Moana" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="1" specimenCount-female="1">
<collectionCode id="ED6DAE33FF84FFBBFC8025F743F7FE72" box="[813,939,402,426]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10752,
<specimenCount id="9D7AFD7FFF84FFBBFB9F25F7440EFE73" box="[1074,1106,402,427]" pageId="7" pageNumber="80" type="female">1♀</specimenCount>
30.0 ×
<quantity id="4C849B13FF84FFBBFB3025F744A5FE71" box="[1181,1273,402,426]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" pageId="7" pageNumber="80" unit="mm" value="48.5">48.5 mm</quantity>
,
<collectingMunicipality id="6BA7AC8CFF84FFBBFAAE25F74317FE1C" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
, MGLN07MV cruise, dive 230,
<location id="8EA3602DFF84FFBBFB3F25C94554FE1C" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFB3F25C94554FE1C" box="[1170,1288,428,452]" country="Fiji" latitude="-20.053833" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" name="Kilo Moana" pageId="7" pageNumber="80">Kilo Moana</location>
site,
<geoCoordinate id="EE485031FF84FFBBFA9125C945F0FE1C" box="[1340,1452,428,452]" degrees="20" direction="south" minutes="03.23" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-20.053833">20°03.23S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFC8025A243ECFE07" box="[813,944,455,479]" degrees="176" direction="west" minutes="08.01" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.1335">176°08.01W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFC1525A24459FE06" box="[952,1029,455,479]" metricMagnitude="3" metricUnit="m" metricValue="2.623" pageId="7" pageNumber="80" unit="m" value="2623.0">
<elevation id="0051D1C5FF84FFBBFC1525A24459FE06" box="[952,1029,455,479]" metricMagnitude="3" metricUnit="m" metricValue="2.623" pageId="7" pageNumber="80" unit="m" value="2623.0">2623 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFBA325A24420FE07" box="[1038,1148,455,479]" pageId="7" pageNumber="80" value="2006-04-07">
<collectingDate id="EF86E9DEFF84FFBBFBA325A24420FE07" box="[1038,1148,455,479]" pageId="7" pageNumber="80" value="2006-04-07">07.IV.2006</collectingDate>
</date>
(preserved in formalin 10% and then transferred into ethanol 70%)
</materialsCitation>
. This large right-handed female, with heterochely and heterodonty (
<figureCitation id="13472A73FF84FFBBFB25259944A8FDCC" box="[1160,1268,508,532]" captionStart="FIG" captionStartId="9.[132,143,1920,1938]" captionTargetBox="[132,1457,216,1756]" captionTargetId="figure@9.[699,984,759,1032]" captionTargetPageId="9" captionText="FIG. 4. — A-C, Austinograea hourdezi n. sp., holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, chelae with two coloured spots on outer surface of palm of right crusher (A) and left cutter (B); C, G1 and G2 photographed in situ; D, E, Austinograea hourdezi n. sp., atypical ♀ 30.0 × 48.5 mm (right-handed), Lau Back-Arc Basin, MGLN07MV cruise, dive 230, Kilo Moana site, 20°03.23S, 176°08.01W, 2623 m, MNHN-IU-2016-10752: D, crusher with one marked dark spot and the other one, only discernible by a yellow trace (wound at base of fixed finger is not a spot comparable to that of A. jolliveti n. sp.); E, cutter with two indistinct spots, only as yellow traces." figureDoi="http://doi.org/10.5281/zenodo.3738108" httpUri="https://zenodo.org/record/3738108/files/figure.png" pageId="7" pageNumber="80">Fig. 4D, E</figureCitation>
), is atypical due to the presence two spots on the chelae (on the crusher, one marked dark spot plus a faint another one, only discernible as yellow trace; on the cutter, two indistinct spots, only as yellow traces), conversely to typical females that are devoid of any spot.
<materialsCitation id="3B143CABFF84FFBBFC8026E544EBFD15" ID-GBIF-Occurrence="2594759994" collectingDate="1989-07-13" collectionCode="MNHN-IU-" country="Fiji" elevation="2750" latitude="-18.833334" location="Mussel Valley" longLatPrecision="1273" longitude="173.48334" pageId="7" pageNumber="80" specimenCount="1" specimenCount-female="1" stateProvince="North Fiji Basin">
<collectionCode id="ED6DAE33FF84FFBBFC8026E543FBFD40" box="[813,935,640,664]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10768 (= MNHN-B27834),
<specimenCount id="9D7AFD7FFF84FFBBFB4226E5454CFD42" box="[1263,1296,640,666]" pageId="7" pageNumber="80" type="female">1 ♀</specimenCount>
22.7 ×
<quantity id="4C849B13FF84FFBBFAFE26E545F7FD4F" box="[1363,1451,640,664]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="7" pageNumber="80" unit="mm" value="36.5">36.5 mm</quantity>
,
<collectingRegion id="49B8F814FF84FFBBFC8026FF4393FD6A" box="[813,975,666,690]" pageId="7" pageNumber="80">North Fiji Basin</collectingRegion>
, STARMER II cruise, dive PL 18,
<location id="8EA3602DFF84FFBBFA8B26FF45F0FD6A" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFA8B26FF45F0FD6A" box="[1318,1452,666,690]" country="Fiji" latitude="-18.833334" longLatPrecision="1273" longitude="173.48334" name="Mussel Valley" pageId="7" pageNumber="80" stateProvince="North Fiji Basin">Mussel Valley</location>
,
<geoCoordinate id="EE485031FF84FFBBFC8026D04326FD15" box="[813,890,693,717]" degrees="18" direction="south" minutes="50" orientation="latitude" pageId="7" pageNumber="80" precision="925" value="-18.833334">18°50S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFC2F26D04383FD15" box="[898,991,693,717]" degrees="173" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="80" precision="925" value="173.48334">173°29E</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFC4A26D04468FD14" box="[999,1076,693,717]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="7" pageNumber="80" unit="m" value="2750.0">
<elevation id="0051D1C5FF84FFBBFC4A26D04468FD14" box="[999,1076,693,717]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="7" pageNumber="80" unit="m" value="2750.0">2750 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFB9126D044EBFD15" box="[1084,1207,693,717]" pageId="7" pageNumber="80" value="1989-07-13">
<collectingDate id="EF86E9DEFF84FFBBFB9126D044EBFD15" box="[1084,1207,693,717]" pageId="7" pageNumber="80" value="1989-07-13">13.VII.1989</collectingDate>
</date>
</materialsCitation>
. This atypical female bears a spot on each chela.
<materialsCitation id="3B143CABFF84FFBBFC80268F44ECFC5E" ID-GBIF-Occurrence="2594760002" collectingDate="2006-09-11" collectionCode="MNHN-IU-" country="Fiji" elevation="1891" latitude="-21.989" location="Collection C. R. Fisher" longLatPrecision="12" longitude="-176.56816" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="2" specimenCount-female="2">
<collectionCode id="ED6DAE33FF84FFBBFC80268F43FBFCDA" box="[813,935,746,770]" pageId="7" pageNumber="80">MNHN-IU-</collectionCode>
2016-10765, only one big crusher chela (with two spots) and several pereopods,
<collectingMunicipality id="6BA7AC8CFF84FFBBFBA427604495FCC4" box="[1033,1225,772,796]" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
, no other data.
<location id="8EA3602DFF84FFBBFC80277A444BFCEF" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFC80277A444BFCEF" box="[813,1047,799,823]" country="Fiji" latitude="-21.989" longLatPrecision="12" longitude="-176.56816" municipality="Lau Back-Arc Basin" name="Collection C. R. Fisher" pageId="7" pageNumber="80">Collection C. R. Fisher</location>
.
<specimenCount id="9D7AFD7FFF84FFBBFB8F277A441FFCE0" box="[1058,1091,799,824]" pageId="7" pageNumber="80" type="female">2 ♀</specimenCount>
, southwestern
<collectingMunicipality id="6BA7AC8CFF84FFBBFB77277A4540FCEF" box="[1242,1308,799,823]" pageId="7" pageNumber="80">Pacific</collectingMunicipality>
,
<collectingMunicipality id="6BA7AC8CFF84FFBBFA85277A433BFC89" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
, MGLN07MV, dive J2-232, Tui
<location id="8EA3602DFF84FFBBFB6C275C455DFC89" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFB6C275C455DFC89" box="[1217,1281,825,849]" country="Fiji" latitude="-21.989" longLatPrecision="12" longitude="-176.56816" municipality="Lau Back-Arc Basin" name="Malila" pageId="7" pageNumber="80">Malila</location>
site,
<geoCoordinate id="EE485031FF84FFBBFA97275C45F7FC89" box="[1338,1451,825,849]" degrees="21" direction="south" minutes="59.34" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-21.989">21°59.34S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFC80273143E8FCB4" box="[813,948,852,876]" degrees="176" direction="west" minutes="34.09" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.56816">176°34.09W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFC122731444EFCB3" box="[959,1042,852,876]" metricMagnitude="3" metricUnit="m" metricValue="1.891" pageId="7" pageNumber="80" unit="m" value="1891.0">
<elevation id="0051D1C5FF84FFBBFC122731444EFCB3" box="[959,1042,852,876]" metricMagnitude="3" metricUnit="m" metricValue="1.891" pageId="7" pageNumber="80" unit="m" value="1891.0">1891 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFBB0273144CCFCB4" box="[1053,1168,852,876]" pageId="7" pageNumber="80" value="2006-09-11">
<collectingDate id="EF86E9DEFF84FFBBFBB0273144CCFCB4" box="[1053,1168,852,876]" pageId="7" pageNumber="80" value="2006-09-11">11.IX.2006</collectingDate>
</date>
(preserved in formalin 10% and then transferred into ethanol 70%)
</materialsCitation>
;
<materialsCitation id="3B143CABFF84FFBBFB17270B43E4FC0E" ID-GBIF-Occurrence="2594759987" collectingDate="2006-09-09" country="Fiji" latitude="-20.760834" location="Lau Back-Arc Basin" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D7AFD7FFF84FFBBFB17270B4487FC50" box="[1210,1243,878,904]" pageId="7" pageNumber="80" type="female">2 ♀</specimenCount>
,
<collectingMunicipality id="6BA7AC8CFF84FFBBFB45270A45F7FC5E" box="[1256,1451,878,902]" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
, MGLN07MV, dive 231, ABE site,
<geoCoordinate id="EE485031FF84FFBBFB3227ED4548FC78" box="[1183,1300,904,928]" degrees="20" direction="south" minutes="45.65" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-20.760834">20°45.65S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFA8D27ED45F7FC78" box="[1312,1451,904,928]" degrees="176" direction="west" minutes="11.45" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.19083">176°11.45W</geoCoordinate>
,
<date id="FFC21036FF84FFBBFC8027C643C3FC63" box="[813,927,931,955]" pageId="7" pageNumber="80" value="2006-09-09">
<collectingDate id="EF86E9DEFF84FFBBFC8027C643C3FC63" box="[813,927,931,955]" pageId="7" pageNumber="80" value="2006-09-09">09.IX.2006</collectingDate>
</date>
(preserved in formalin 10% and then transferred into ethanol 70%)
</materialsCitation>
;
<materialsCitation id="3B143CABFF84FFBBFC6927DB4466FBD2" ID-GBIF-Occurrence="2594760004" collectingDate="2009-05-25" country="Fiji" elevation="2130" latitude="-20.760834" location="Lau Basin" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" pageId="7" pageNumber="80" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D7AFD7FFF84FFBBFC6927DB43B9FC0F" box="[964,997,958,983]" pageId="7" pageNumber="80" type="female">2 ♀</specimenCount>
,
<collectingMunicipality id="6BA7AC8CFF84FFBBFC5927DB44E2FC0D" box="[1012,1214,957,981]" pageId="7" pageNumber="80">Lau Back-Arc Basin</collectingMunicipality>
,
<location id="8EA3602DFF84FFBBFB6427DB4571FC0D" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF84FFBBFB6427DB4571FC0D" box="[1225,1325,957,981]" country="Fiji" latitude="-20.760834" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" name="Lau Basin" pageId="7" pageNumber="80">Lau Basin</location>
2009 cruise, dive 425, ABE site,
<geoCoordinate id="EE485031FF84FFBBFC4627BD4406FC28" box="[1003,1114,984,1008]" degrees="20" direction="south" minutes="45.65" orientation="latitude" pageId="7" pageNumber="80" precision="9" value="-20.760834">20°45.65S</geoCoordinate>
,
<geoCoordinate id="EE485031FF84FFBBFBCE27BD44B4FC28" box="[1123,1256,984,1008]" degrees="176" direction="west" minutes="11.45" orientation="longitude" pageId="7" pageNumber="80" precision="9" value="-176.19083">176°11.45W</geoCoordinate>
,
<quantity id="4C849B13FF84FFBBFB5C27BD4563FC37" box="[1265,1343,984,1008]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="80" unit="m" value="2130.0">
<elevation id="0051D1C5FF84FFBBFB5C27BD4563FC37" box="[1265,1343,984,1008]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="80" unit="m" value="2130.0">2130 m</elevation>
</quantity>
,
<date id="FFC21036FF84FFBBFAE527BD45F3FC28" box="[1352,1455,984,1008]" pageId="7" pageNumber="80" value="2009-05-25">
<collectingDate id="EF86E9DEFF84FFBBFAE527BD45F3FC28" box="[1352,1455,984,1008]" pageId="7" pageNumber="80" value="2009-05-25">25.V.2009</collectingDate>
</date>
(preserved in ethanol 85%)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C366657DFF84FFBBFC80204E4516FB1F" pageId="7" pageNumber="80" type="etymology">
<paragraph id="8BC336F6FF84FFBBFC80204E4516FB1F" blockId="7.[813,1456,1067,1223]" pageId="7" pageNumber="80">
ETYMOLOGY. — Dedicated to Stéphane Hourdez (CNRS, Station biologique de Roscoff,
<collectingCountry id="F36B7666FF84FFBBFBA22023440EFB86" box="[1039,1106,1094,1118]" name="France" pageId="7" pageNumber="80">France</collectingCountry>
) in recognition of his valuable contribution to the knowledge of hydrothermal fauna sites (taxonomy, ecology, physiology, adaptation, phylogeny). He actively participated in the taxonomic sampling during the C. R. Fishers cruises in the Lau Basin, suspecting the presence of new
<taxonomicName id="4C7C4D75FF84FFBBFB6120D54518FB1F" authorityName="Hessler &amp; Martin" authorityYear="1989" box="[1228,1348,1200,1223]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B908EAE4FF84FFBBFB6120D54518FB1F" box="[1228,1348,1200,1223]" italics="true" pageId="7" pageNumber="80">Austinograea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C366657DFF84FFBBFC80208D442DF9D1" pageId="7" pageNumber="80" type="distribution">
<paragraph id="8BC336F6FF84FFBBFC80208D445DFAB2" blockId="7.[813,1456,1256,1545]" pageId="7" pageNumber="80">
DISTRIBUTION. —
<taxonomicName id="4C7C4D75FF84FFBBFC49208D44EFFAD8" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[996,1203,1256,1280]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF84FFBBFC49208D44EFFAD8" box="[996,1203,1256,1280]" italics="true" pageId="7" pageNumber="80">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF84FFBBFB15208C44B1FAD8" box="[1208,1261,1257,1280]" pageId="7" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
is a widespread species encountered in the southwestern Pacific from numerous vent fields of the Lau and North
<collectingCountry id="F36B7666FF84FFBBFBEF2178443EFAED" box="[1090,1122,1309,1333]" name="Fiji" pageId="7" pageNumber="80">Fiji</collectingCountry>
Back-Arc Basins. It cohabits with
<taxonomicName id="4C7C4D75FF84FFBBFC80215D43CEFA97" authorityName="Guinot" authorityYear="1990" box="[813,914,1335,1359]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF84FFBBFC80215D43CEFA97" box="[813,914,1335,1359]" italics="true" pageId="7" pageNumber="80">A. alayseae</emphasis>
</taxonomicName>
at a number of sites, and the two species seem to share a similar distribution.
</paragraph>
<paragraph id="8BC336F6FF84FFBBFC802108442DF9D1" blockId="7.[813,1456,1256,1545]" pageId="7" pageNumber="80">
As
<taxonomicName id="4C7C4D75FF84FFBBFCE6210843ECFA5C" authorityName="Guinot" authorityYear="1990" box="[843,944,1388,1412]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF84FFBBFCE6210843ECFA5C" box="[843,944,1388,1412]" italics="true" pageId="7" pageNumber="80">A. alayseae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7C4D75FF84FFBBFC1021084476FA5C" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[957,1066,1388,1412]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF84FFBBFC1021084476FA5C" box="[957,1066,1388,1412]" italics="true" pageId="7" pageNumber="80">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF84FFBBFB9C2108443BFA5C" box="[1073,1127,1389,1412]" pageId="7" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
has not been found in the northwestern Pacific, at least after the revision of the MNHN collection and without, however, the examination of an extensive material from the Mariana Trough. For its part,
<taxonomicName id="4C7C4D75FF84FFBBFBDD21D944BEFA0C" authorityName="in Guinot" authorityYear="1990" box="[1136,1250,1468,1492]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="80" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF84FFBBFBDD21D944BEFA0C" box="[1136,1250,1468,1492]" italics="true" pageId="7" pageNumber="80">A. williamsi</emphasis>
</taxonomicName>
seems to be confined to the northwestern Pacific, in the Mariana Back-Arc Basin, and vent sites of the Mariana Trough.
</paragraph>
</subSubSection>
<subSubSection id="C366657DFF84FFB7FC80225145C8F996" lastPageId="11" lastPageNumber="84" pageId="7" pageNumber="80" type="description">
<paragraph id="8BC336F6FF84FFBBFC8022514453F996" blockId="7.[813,1456,1588,2030]" box="[813,1039,1588,1614]" pageId="7" pageNumber="80">DESCRIPTION (MALE)</paragraph>
<paragraph id="8BC336F6FF84FFBBFC80223043D1F9B7" blockId="7.[813,1456,1588,2030]" box="[813,909,1621,1647]" pageId="7" pageNumber="80">
<emphasis id="B908EAE4FF84FFBBFC80223043D1F9B7" box="[813,909,1621,1647]" italics="true" pageId="7" pageNumber="80">Carapace</emphasis>
</paragraph>
<paragraph id="8BC336F6FF84FFB7FC802211410DFDCA" blockId="7.[813,1456,1588,2030]" lastBlockId="11.[132,776,217,530]" lastPageId="11" lastPageNumber="84" pageId="7" pageNumber="80">Carapace transversely elliptical, flat; regions indistinct. Dorsal surface nearly smooth, except for rare small granules on frontal region and small tubercles on orbital region; numerous small pits. Anterolateral margin marked by rounded edge, slightly granulous. Posterolateral margins convergent; posterior margin slightly concave. Subhepatic regions entirely covered by patches of dense setae. Front broad, not protruded, bluntly pointed medially and with two slightly concave lobes. Eyes, antennules and antennae recessed under front. Suborbital plate absent. Orbit not delimited, only an orbital region that extends as groove lateral to region containing vestigial eyestalks and antennae. Outer side of orbital region tuberculated. Eyestalk not moveable, showing as fixed piece fused to floor of orbital region; no cornea. Antennules folded horizontally. Antenna inserted in wide supraorbital notch; urinary article fixed, broad, recessed; basal article (2 +3) cylindrical, moveable; article 4 slightly elongated, inclined; flagellum of about 10 articles. Proepistome incomplete. Margin of epistome with moderate median projection and formed of two moderately concave lobes, marked by row of low granules. Pterygostomial lobe tuberculous; pterygostomial region smooth, except for small curved ridge.</paragraph>
<caption id="DF03667EFF8BFFB4FF2923344579F833" ID-DOI="http://doi.org/10.5281/zenodo.3738106" ID-Zenodo-Dep="3738106" httpUri="https://zenodo.org/record/3738106/files/figure.png" pageId="8" pageNumber="81" startId="8.[132,143,1873,1891]" targetBox="[397,1190,214,1823]" targetPageId="8">
<paragraph id="8BC336F6FF8BFFB4FF2923344579F833" blockId="8.[132,1457,1872,2027]" pageId="8" pageNumber="81">
FIG. 3. —
<taxonomicName id="4C7C4D75FF8BFFB4FF7B233441CCF8BB" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[214,400,1873,1891]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8BFFB4FF7B233441CCF8BB" box="[214,400,1873,1891]" italics="true" pageId="8" pageNumber="81">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8BFFB4FE382334419AF8BB" box="[405,454,1873,1891]" pageId="8" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
photographed
<emphasis id="B908EAE4FF8BFFB4FDEB23344224F8BB" box="[582,632,1873,1891]" italics="true" pageId="8" pageNumber="81">in situ</emphasis>
:
<emphasis id="B908EAE4FF8BFFB4FD2E233442CCF8BB" bold="true" box="[643,656,1873,1891]" pageId="8" pageNumber="81">A</emphasis>
, Lau Basin, Lau Basin 2009 cruise, dive 432, Tow Cam site, 20˚18.98S, 176˚08.19W, 2723 m, 05.VI.2009 (type locality), among sessiles barnacles
<taxonomicName id="4C7C4D75FF8BFFB4FD94230D43E9F8A2" authority="ohtai Yamaguchi, 1990" authorityName="Yamaguchi" authorityYear="1990" box="[569,949,1896,1914]" class="Hexanauplia" family="Pachylasmatidae" genus="Eochionelasmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="81" phylum="Arthropoda" rank="subSpecies" species="ohtai" subSpecies="ohtai">
<emphasis id="B908EAE4FF8BFFB4FD94230D4341F8A2" box="[569,797,1896,1914]" italics="true" pageId="8" pageNumber="81">Eochionelasmus ohtai ohtai</emphasis>
Yamaguchi, 1990
</taxonomicName>
. The two dark spots on outer surface of palm of both chelae indicate a male; see patches of dense setae on inner surface of both chelae, and the darkly coloured two-thirds of dactylus of right cheliped;
<emphasis id="B908EAE4FF8BFFB4FABE231A457CF849" bold="true" box="[1299,1312,1919,1937]" pageId="8" pageNumber="81">B</emphasis>
,
<emphasis id="B908EAE4FF8BFFB4FA87231A4564F849" bold="true" box="[1322,1336,1919,1937]" pageId="8" pageNumber="81">C</emphasis>
,
<taxonomicName id="4C7C4D75FF8BFFB4FAEF231B4097F87F" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="81" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8BFFB4FAEF231B4097F87F" italics="true" pageId="8" pageNumber="81">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8BFFB4FF7D23F0415EF87F" box="[208,258,1941,1959]" pageId="8" pageNumber="81" rank="species">n. sp.</taxonomicNameLabel>
, due to the patch of dense setae on inner surface of both chelae:
<emphasis id="B908EAE4FF8BFFB4FC8E23F0436CF87F" bold="true" box="[803,816,1941,1959]" pageId="8" pageNumber="81">B</emphasis>
, Lau Basin, TUIM07MV cruise, dive 163, ABE site, 20°45.65S, 176°11.45W, 2130 m, 23.VI.2005; on a bed of gastropod
<taxonomicName id="4C7C4D75FF8BFFB4FE5623C94305F866" authority="Bouchet &amp; Waren, 1991" authorityName="Bouchet &amp; Waren" authorityYear="1991" box="[507,857,1964,1982]" class="Gastropoda" family="Provannidae" genus="Ifremeria" kingdom="Animalia" pageId="8" pageNumber="81" phylum="Mollusca" rank="species" species="nautilei">
<emphasis id="B908EAE4FF8BFFB4FE5623C942DAF866" box="[507,646,1964,1982]" italics="true" pageId="8" pageNumber="81">Ifremeria nautilei</emphasis>
Bouchet &amp; Warén, 1991
</taxonomicName>
;
<emphasis id="B908EAE4FF8BFFB4FCCF23C9432CF866" bold="true" box="[866,880,1964,1982]" pageId="8" pageNumber="81">C</emphasis>
, Lau Basin, Lau Basin 2009 cruise, dive 431, ABE site, 20°45.65S, 176°11.45W, 2130 m, 03.VI.2009. The crab and the gastropods
<emphasis id="B908EAE4FF8BFFB4FD3023A643F8F80D" box="[669,932,1987,2005]" italics="true" pageId="8" pageNumber="81">
<taxonomicName id="4C7C4D75FF8BFFB4FD3023A64316F80D" authorityName="Okutani &amp; Ohta" authorityYear="1988" box="[669,842,1987,2005]" class="Gastropoda" family="Provannidae" genus="Alviniconcha" kingdom="Animalia" pageId="8" pageNumber="81" phylum="Mollusca" rank="species" sensu="lato" species="hessleri">Alviniconcha hessleri</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8BFFB4FCFD23A643F8F80D" box="[848,932,1987,2005]" pageId="8" pageNumber="81" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
are blurry, the photography
<emphasis id="B908EAE4FF8BFFB4FB3D23A6449DF80D" box="[1168,1217,1987,2005]" italics="true" pageId="8" pageNumber="81">in situ</emphasis>
having been taken in hydrothermal diffuse area with the emanating fluid visible as shimmering water. Photographies courtesy of C. R. Fisher (Penn State University, USA).
</paragraph>
</caption>
<caption id="DF03667EFF8AFFB5FF2923E545FCF835" ID-DOI="http://doi.org/10.5281/zenodo.3738108" ID-Zenodo-Dep="3738108" httpUri="https://zenodo.org/record/3738108/files/figure.png" pageId="9" pageNumber="82" startId="9.[132,143,1920,1938]" targetBox="[132,1457,216,1756]" targetPageId="9">
<paragraph id="8BC336F6FF8AFFB5FF2923E545FCF835" blockId="9.[132,1457,1920,2029]" pageId="9" pageNumber="82">
FIG. 4. —
<emphasis id="B908EAE4FF8AFFB5FF7A23E540A6F84A" bold="true" box="[215,250,1920,1938]" pageId="9" pageNumber="82">A -C</emphasis>
,
<taxonomicName id="4C7C4D75FF8AFFB5FEAB23E5419FF84A" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[262,451,1920,1938]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8AFFB5FEAB23E5419FF84A" box="[262,451,1920,1938]" italics="true" pageId="9" pageNumber="82">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8AFFB5FE6723E541A2F84A" box="[458,510,1920,1938]" pageId="9" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
, holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737:
<emphasis id="B908EAE4FF8AFFB5FDDC23F24222F871" bold="true" box="[625,638,1943,1961]" pageId="9" pageNumber="82">A</emphasis>
,
<emphasis id="B908EAE4FF8AFFB5FD2423F242CAF871" bold="true" box="[649,662,1943,1961]" pageId="9" pageNumber="82">B</emphasis>
, chelae with two coloured spots on outer surface of palm of right crusher (
<emphasis id="B908EAE4FF8AFFB5FAAE23F2454CF871" bold="true" box="[1283,1296,1943,1961]" pageId="9" pageNumber="82">A</emphasis>
) and left cutter (
<emphasis id="B908EAE4FF8AFFB5FA3523F245F9F871" bold="true" box="[1432,1445,1943,1961]" pageId="9" pageNumber="82">B</emphasis>
);
<emphasis id="B908EAE4FF8AFFB5FF2923C840CEF867" bold="true" box="[132,146,1965,1983]" pageId="9" pageNumber="82">C</emphasis>
, G1 and G2 photographed
<emphasis id="B908EAE4FF8AFFB5FED223C841EDF867" box="[383,433,1965,1983]" italics="true" pageId="9" pageNumber="82">in situ</emphasis>
;
<emphasis id="B908EAE4FF8AFFB5FE1023C84197F867" bold="true" box="[445,459,1965,1983]" pageId="9" pageNumber="82">D</emphasis>
,
<emphasis id="B908EAE4FF8AFFB5FE7A23C841BFF867" bold="true" box="[471,483,1965,1983]" pageId="9" pageNumber="82">E</emphasis>
,
<taxonomicName id="4C7C4D75FF8AFFB5FE4223C842F6F867" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[495,682,1965,1983]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8AFFB5FE4223C842F6F867" box="[495,682,1965,1983]" italics="true" pageId="9" pageNumber="82">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8AFFB5FD1D23C842B8F867" box="[688,740,1965,1983]" pageId="9" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
, atypical ♀ 30.0 × 48.5 mm (right-handed), Lau Back-Arc Basin, MGLN07MV cruise, dive 230, Kilo Moana site, 20°03.23S, 176°08.01W, 2623 m, MNHN-IU-2016-10752:
<emphasis id="B908EAE4FF8AFFB5FCE323A14300F80E" bold="true" box="[846,860,1988,2006]" pageId="9" pageNumber="82">D</emphasis>
, crusher with one marked dark spot and the other one, only discernible by a yellow trace (wound at base of fixed finger is not a spot comparable to that of
<taxonomicName id="4C7C4D75FF8AFFB5FC9523BE43DBF835" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[824,903,2011,2029]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="82" phylum="Arthropoda" rank="species" species="jolliveti" status="sp. nov.">
<emphasis id="B908EAE4FF8AFFB5FC9523BE43DBF835" box="[824,903,2011,2029]" italics="true" pageId="9" pageNumber="82">A. jolliveti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8AFFB5FC2123BE43E2F835" box="[908,958,2011,2029]" pageId="9" pageNumber="82" rank="species">n. sp.</taxonomicNameLabel>
);
<emphasis id="B908EAE4FF8AFFB5FC6723BE438AF835" bold="true" box="[970,982,2011,2029]" pageId="9" pageNumber="82">E</emphasis>
, cutter with two indistinct spots, only as yellow traces.
</paragraph>
</caption>
<caption id="DF03667EFF89FFB6FF2923C844D5F834" ID-DOI="http://doi.org/10.5281/zenodo.3738110" ID-Zenodo-Dep="3738110" httpUri="https://zenodo.org/record/3738110/files/figure.png" pageId="10" pageNumber="83" startId="10.[132,143,1965,1983]" targetBox="[132,1455,216,1859]" targetPageId="10">
<paragraph id="8BC336F6FF89FFB6FF2923C844D5F834" blockId="10.[132,1455,1965,2029]" pageId="10" pageNumber="83">
FIG. 5. —
<taxonomicName id="4C7C4D75FF89FFB6FF7A23C841CEF867" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[215,402,1965,1983]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="83" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF89FFB6FF7A23C841CEF867" box="[215,402,1965,1983]" italics="true" pageId="10" pageNumber="83">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF89FFB6FE3423C84190F867" box="[409,460,1965,1983]" pageId="10" pageNumber="83" rank="species">n. sp.</taxonomicNameLabel>
, paratype, ♂ 29.8 × 48.1 mm (left-handed, with weak heterochely and heterodonty), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738:
<emphasis id="B908EAE4FF89FFB6FDFD23A14201F80E" bold="true" box="[592,605,1988,2006]" pageId="10" pageNumber="83">A</emphasis>
, dorsal view;
<emphasis id="B908EAE4FF89FFB6FD6323A14287F80E" bold="true" box="[718,731,1988,2006]" pageId="10" pageNumber="83">B</emphasis>
, frontal view;
<emphasis id="B908EAE4FF89FFB6FCE023A14307F80E" bold="true" box="[845,859,1988,2006]" pageId="10" pageNumber="83">C</emphasis>
, mxp3;
<emphasis id="B908EAE4FF89FFB6FC3023A143F7F80E" bold="true" box="[925,939,1988,2006]" pageId="10" pageNumber="83">D</emphasis>
, ventral view;
<emphasis id="B908EAE4FF89FFB6FB8D23A14470F80E" bold="true" box="[1056,1068,1988,2006]" pageId="10" pageNumber="83">E</emphasis>
,
<emphasis id="B908EAE4FF89FFB6FB9823A1441FF80E" bold="true" box="[1077,1091,1988,2006]" pageId="10" pageNumber="83">G</emphasis>
, both chelae with two typical spots on outer surface;
<emphasis id="B908EAE4FF89FFB6FF6123BE408BF835" bold="true" box="[204,215,2011,2029]" pageId="10" pageNumber="83">F</emphasis>
,
<emphasis id="B908EAE4FF89FFB6FF4F23BE40ACF835" bold="true" box="[226,240,2011,2029]" pageId="10" pageNumber="83">H</emphasis>
, both chelae with patches of setae on inner surface:
<emphasis id="B908EAE4FF89FFB6FD0423BE42E9F835" bold="true" box="[681,693,2011,2029]" pageId="10" pageNumber="83">E</emphasis>
,
<emphasis id="B908EAE4FF89FFB6FD1223BE4296F835" bold="true" box="[703,714,2011,2029]" pageId="10" pageNumber="83">F</emphasis>
, cutter situated at right;
<emphasis id="B908EAE4FF89FFB6FC3A23BF43F9F834" bold="true" box="[919,933,2010,2028]" pageId="10" pageNumber="83">G</emphasis>
,
<emphasis id="B908EAE4FF89FFB6FC1D23BE43E2F835" bold="true" box="[944,958,2011,2029]" pageId="10" pageNumber="83">H</emphasis>
, crusher situated at left.
</paragraph>
</caption>
<paragraph id="8BC336F6FF88FFB7FF29265D415EFD8A" blockId="11.[132,776,568,2030]" box="[132,258,568,594]" pageId="11" pageNumber="84">
<emphasis id="B908EAE4FF88FFB7FF29265D415EFD8A" box="[132,258,568,594]" italics="true" pageId="11" pageNumber="84">Thoracopods</emphasis>
</paragraph>
<paragraph id="8BC336F6FF88FFB7FF29263D41F2F836" blockId="11.[132,776,568,2030]" pageId="11" pageNumber="84">
Third maxillipeds completely closing buccal cavity. Ischium long, with marked longitudinal internal groove. Merus: external margin regularly curved, oblique, without marked angle, with distal part markedly narrow and produced; internal margin with prominent lobe in half proximal portion. Carpus inserted in notch of antero-internal margin of merus; propodus thick; dactylus about as long as merus; inner margins of propodus and dactylus with brush-like setae. Exopod longer than endopod ischium. Mxp3 coxa with only proximal portion visible, its lateral projection hidden by junction of thoracic sternum (sternite 4) with pterygostome. Chelipeds heteromorphic, marked heterochely (also in females, see below), with two
<typeStatus id="54C78854FF88FFB7FEE527B241DCFC29" box="[328,384,983,1009]" pageId="11" pageNumber="84">types</typeStatus>
of chelae: major cheliped blunt-tip (crusher), stouter and shorter than pointed-tip minor chela (cutter). Carpus with setae on superior margin and inner surface; merus long, triangular in cross section, anterior border straight, with blunt teeth, smaller distally. Propodus of both chelae with two conspicuous, deep depressions at outer surface of palm near base of dactylus (absent in females), both as two dark spots, even well visible in specimens photographed
<emphasis id="B908EAE4FF88FFB7FE8520B34131FB37" box="[296,365,1238,1263]" italics="true" pageId="11" pageNumber="84">in situ</emphasis>
. Inner surface of both chelae dished on half posterior part; thick patches of dense setae largely extending on half anterior part of propodus and partially extending along occluding margins. Dark colour extending on about three-quarters of fixed finger and practically on whole part of dactylus of both crusher and cutter. Crusher: palm convex, inflated, smooth, except for two depressions located near base of dactylus: one horizontally directed and the largest; the other obliquely directed, smaller. Fingers thick, slightly gaping at occluding margins; two weak, blunt teeth, proximal and median, on occluding margin of dactylus; a blunt but marked subproximal tooth and smaller ones on occluding margin of fixed finger. Cutter: palm elongate, with subparallel borders, smooth, except for two depressions near base of dactylus, similar to those of the crusher; both fingers not gaping at the occluding margins; dactylus elongate, hollowed, with occluding margin smooth; fixed finger very thick, dished on inner surface; occluding margin nearly straight, with 2-3 small teeth, size progressively diminishing forwards. P2-P5 shorter than chelipeds; P3 and P4 longest; meri with patch of dense setae on ventral margins, carpi, propodi fringed on dorsal and ventral margins with felt of short, stiff setae, mixed with sparse longer setae, the patch being dense, thicker on inferior margin. Dactyli rather stout, with patches of short setae.
</paragraph>
<paragraph id="8BC336F6FF88FFB7FC8024BC4382FF2B" blockId="11.[813,1456,217,435]" box="[813,990,217,243]" pageId="11" pageNumber="84">
<emphasis id="B908EAE4FF88FFB7FC8024BC4382FF2B" box="[813,990,217,243]" italics="true" pageId="11" pageNumber="84">Thoracic sternum</emphasis>
</paragraph>
<paragraph id="8BC336F6FF88FFB7FC80249C44ABFE6B" blockId="11.[813,1456,217,435]" pageId="11" pageNumber="84">Thoracic sternum with sutures 4/5-7/8 incomplete but separared by short gap; suture 2/3 complete. Median line only along sternite 8, hardly continuying along sternite 7. Junction of sternite 4 with pterygostome represented by only short juxtaposition, without patch of setae. Press-button of locking mechanism acute, very close to suture 5/6.</paragraph>
<paragraph id="8BC336F6FF88FFB7FC8025BD433AFE2A" blockId="11.[813,1456,472,753]" box="[813,870,472,498]" pageId="11" pageNumber="84">
<emphasis id="B908EAE4FF88FFB7FC8025BD433AFE2A" box="[813,870,472,498]" italics="true" pageId="11" pageNumber="84">Pleon</emphasis>
</paragraph>
<paragraph id="8BC336F6FF88FFB7FC80259D451BFD29" blockId="11.[813,1456,472,753]" pageId="11" pageNumber="84">Pleon of six free segments and telson regularly triangular; pleonal somite 3 widest; pleonal somite 6 longest, anterointernally delineated by raised, thick margin; telson triangular, rounded distally. Both G1 joining at tips. G1: slender, faintly curved, with only short, spiniform setae arranged in two rows along mid-part only. G2: about half the length of G1, with bend about two-thirds length at level of small setiferous area; flagellum rather long, curved, flattened, bladelike.</paragraph>
<paragraph id="8BC336F6FF88FFB7FC802772447AFCE9" blockId="11.[813,1456,791,1615]" box="[813,1062,791,817]" pageId="11" pageNumber="84">DESCRIPTION (FEMALE)</paragraph>
<paragraph id="8BC336F6FF88FFB7FC80275245C8F996" blockId="11.[813,1456,791,1615]" pageId="11" pageNumber="84">Chelipeds weakly dimorphic (versus dimorphic in males), having almost cutters on both sides. On both chelipeds: inner surface markedly dished on half posterior part of propodus and on fixed finger; thick patches of dense setae extending on half or anterior two-thirds part of propodus and along occluding margins of dactyli. Dark colour extending on about threequarters of fixed finger and on two-thirds of dactylus of both crusher and cutter. Weak heterochely and heterodonty; cutter however smaller than crusher. Crusher: palm hardly inflated, elongated, smooth, without depressions; fingers thick, not or only slightly gaping at the occluding margins; two tiny blunt teeth on occluding margin of dactylus; two tooth and a few smaller ones on occluding margin of fixed finger.Cutter: palm similar to crusher but fingers more elongated, not gaping at occluding margins; palm smooth, without depressions; both fingers; dactylus elongate, hollowed, with occluding margin practically smooth; fixed finger very thick, with occluding margin nearly straight, bearing small teeth, two being more marked. P2-P5 with felt of short, stiff setae, mixed with numerous longer setae. Median line along sternite 8, hardly continuying along sternite 7. Dense patches of setae on thoracic sternum near junction of sternite 4 with pterygostome. No pleonal locking mechanism in mature females. Margins of pleon densely fringed with setae. Vulvae large, rounded, with thick membranous area all around median opening.</paragraph>
</subSubSection>
<subSubSection id="C366657DFF88FFB1FC80221144AAFAA8" lastPageId="13" lastPageNumber="86" pageId="11" pageNumber="84" type="diagnosis">
<paragraph id="8BC336F6FF88FFB7FC8022114484F954" blockId="11.[813,1456,1652,2030]" box="[813,1240,1652,1678]" pageId="11" pageNumber="84">REMARKS AND COMPARATIVE DIAGNOSIS</paragraph>
<paragraph id="8BC336F6FF88FFB7FC8022F1457EF836" blockId="11.[813,1456,1652,2030]" pageId="11" pageNumber="84">
The two characteristic depressions on the propodus of both male chelae, crusher and cutter, of
<taxonomicName id="4C7C4D75FF88FFB7FB3E22D1452EF916" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[1171,1394,1716,1742]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="84" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF88FFB7FB3E22D1452EF916" box="[1171,1394,1716,1742]" italics="true" pageId="11" pageNumber="84">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF88FFB7FADA22D045ECF917" box="[1399,1456,1717,1743]" pageId="11" pageNumber="84" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13472A73FF88FFB7FC9922B143D5F936" box="[820,905,1748,1774]" captionStart="FIG" captionStartId="8.[132,143,1873,1891]" captionTargetBox="[397,1190,214,1823]" captionTargetId="figure@8.[396,927,687,1161]" captionTargetPageId="8" captionText="FIG. 3. — Austinograea hourdezi n. sp. photographed in situ: A, Lau Basin, Lau Basin 2009 cruise, dive 432, Tow Cam site, 20˚18.98S, 176˚08.19W, 2723 m, 05.VI.2009 (type locality), among sessiles barnacles Eochionelasmus ohtai ohtai Yamaguchi, 1990. The two dark spots on outer surface of palm of both chelae indicate a male; see patches of dense setae on inner surface of both chelae, and the darkly coloured two-thirds of dactylus of right cheliped; B, C, Austinograea hourdezi n. sp., due to the patch of dense setae on inner surface of both chelae: B, Lau Basin, TUIM07MV cruise, dive 163, ABE site, 20°45.65S, 176°11.45W, 2130 m, 23.VI.2005; on a bed of gastropod Ifremeria nautilei Bouchet &amp; Warén, 1991; C, Lau Basin, Lau Basin 2009 cruise, dive 431, ABE site, 20°45.65S, 176°11.45W, 2130 m, 03.VI.2009. The crab and the gastropods Alviniconcha hessleri sensu lato are blurry, the photography in situ having been taken in hydrothermal diffuse area with the emanating fluid visible as shimmering water. Photographies courtesy of C. R. Fisher (Penn State University, USA)." figureDoi="http://doi.org/10.5281/zenodo.3738106" httpUri="https://zenodo.org/record/3738106/files/figure.png" pageId="11" pageNumber="84">Figs 3A</figureCitation>
;
<figureCitation id="13472A73FF88FFB7FC3E22B1438FF936" box="[915,979,1748,1774]" captionStart="FIG" captionStartId="9.[132,143,1920,1938]" captionTargetBox="[132,1457,216,1756]" captionTargetId="figure@9.[699,984,759,1032]" captionTargetPageId="9" captionText="FIG. 4. — A-C, Austinograea hourdezi n. sp., holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, chelae with two coloured spots on outer surface of palm of right crusher (A) and left cutter (B); C, G1 and G2 photographed in situ; D, E, Austinograea hourdezi n. sp., atypical ♀ 30.0 × 48.5 mm (right-handed), Lau Back-Arc Basin, MGLN07MV cruise, dive 230, Kilo Moana site, 20°03.23S, 176°08.01W, 2623 m, MNHN-IU-2016-10752: D, crusher with one marked dark spot and the other one, only discernible by a yellow trace (wound at base of fixed finger is not a spot comparable to that of A. jolliveti n. sp.); E, cutter with two indistinct spots, only as yellow traces." figureDoi="http://doi.org/10.5281/zenodo.3738108" httpUri="https://zenodo.org/record/3738108/files/figure.png" pageId="11" pageNumber="84">4A, B</figureCitation>
;
<figureCitation id="13472A73FF88FFB7FC7322B1447CF936" box="[990,1056,1748,1775]" captionStart="FIG" captionStartId="10.[132,143,1965,1983]" captionTargetBox="[132,1455,216,1859]" captionTargetId="figure@10.[698,1267,763,1313]" captionTargetPageId="10" captionText="FIG. 5. — Austinograea hourdezi n. sp., paratype, ♂ 29.8 × 48.1 mm (left-handed, with weak heterochely and heterodonty), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738: A, dorsal view; B, frontal view; C, mxp3; D, ventral view; E, G, both chelae with two typical spots on outer surface; F, H, both chelae with patches of setae on inner surface: E, F, cutter situated at right; G, H, crusher situated at left." figureDoi="http://doi.org/10.5281/zenodo.3738110" httpUri="https://zenodo.org/record/3738110/files/figure.png" pageId="11" pageNumber="84">5E, G</figureCitation>
), typically absent in females (
<figureCitation id="13472A73FF88FFB7FAF722B14318F8D6" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1858]" captionTargetId="figure@14.[699,984,1036,1311]" captionTargetPageId="14" captionText="FIG. 7. — Austinograea hourdezi n. sp., paratype, ♀ 18.3 × 28.4 mm (right-handed), Lau Back-Arc Basin, BIOLAU cruise, BL 12, Vai Lili site, 23°13S, 176°38W, 1750 m, MNHN-IU-2016-10745 (= MNHN-B24060): A, dorsal view; B, C, ventral views, with pleon; D, thoracic sternum and vulvae; E, G, both chelae without spots on outer surface; F, H, both chelae with patches of setae on inner surface; E, F, right crusher; G, H, left cutter." figureDoi="http://doi.org/10.5281/zenodo.3738114" httpUri="https://zenodo.org/record/3738114/files/figure.png" pageId="11" pageNumber="84">Fig. 7E, G</figureCitation>
), are similarly located in all the individuals examined, but they are variously delineated by a raised margin and thus may appear quite deep or shallower but practically always coloured as dark or coloured spots, always with a different texture and colour but commonly referred to as “dark spots”. The function of the pigmented pits or spots present on the palm of chelipeds (also on the subocular regions in others species) of certain bythograeids is supposed to be sensory.
</paragraph>
<caption id="DF03667EFF8FFFB0FF2923F3446DF834" ID-DOI="http://doi.org/10.5281/zenodo.3738112" ID-Zenodo-Dep="3738112" httpUri="https://zenodo.org/record/3738112/files/figure.png" pageId="12" pageNumber="85" startId="12.[132,143,1942,1960]" targetBox="[168,1419,251,1804]" targetPageId="12">
<paragraph id="8BC336F6FF8FFFB0FF2923F3446DF834" blockId="12.[132,1456,1942,2028]" pageId="12" pageNumber="85">
FIG. 6. —
<taxonomicName id="4C7C4D75FF8FFFB0FF7A23F341C9F870" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[215,405,1942,1960]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="85" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8FFFB0FF7A23F341C9F870" box="[215,405,1942,1960]" italics="true" pageId="12" pageNumber="85">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8FFFB0FE3123F3418CF870" box="[412,464,1942,1960]" pageId="12" pageNumber="85" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B908EAE4FF8FFFB0FE7523F341A6F870" bold="true" box="[472,506,1942,1960]" pageId="12" pageNumber="85">A -C</emphasis>
, holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737:
<emphasis id="B908EAE4FF8FFFB0FDDB23C842DFF867" bold="true" box="[630,643,1965,1983]" pageId="12" pageNumber="85">A</emphasis>
,
<emphasis id="B908EAE4FF8FFFB0FD2323C842C7F867" bold="true" box="[654,667,1965,1983]" pageId="12" pageNumber="85">B</emphasis>
, right and left mxp3, with and without ornamentation;
<emphasis id="B908EAE4FF8FFFB0FBC523C8442AF867" bold="true" box="[1128,1142,1965,1983]" pageId="12" pageNumber="85">C</emphasis>
, thoracic sternum.
<emphasis id="B908EAE4FF8FFFB0FAB623C84575F867" bold="true" box="[1307,1321,1965,1983]" pageId="12" pageNumber="85">D</emphasis>
,
<emphasis id="B908EAE4FF8FFFB0FA9923C8451CF867" bold="true" box="[1332,1344,1965,1983]" pageId="12" pageNumber="85">E</emphasis>
, paratype, ♂ 29.8 × 48.1 mm (left-handed), Lau Back-Arc Basin,TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738:
<emphasis id="B908EAE4FF8FFFB0FBF523A1443AF80E" bold="true" box="[1112,1126,1988,2006]" pageId="12" pageNumber="85">D</emphasis>
, G1, two views;
<emphasis id="B908EAE4FF8FFFB0FB4023A144A5F80E" bold="true" box="[1261,1273,1988,2006]" pageId="12" pageNumber="85">E</emphasis>
, G2, two views. Scale bars: A, B, D, E, 1 mm; C, 5 mm. Note the scale bars of the two gonopods, the G2 being about half-length of G1.
</paragraph>
</caption>
<paragraph id="8BC336F6FF8EFFB1FF3624BF4183FECC" blockId="13.[132,776,217,2030]" pageId="13" pageNumber="86">
For atypical specimens of
<taxonomicName id="4C7C4D75FF8EFFB1FE1F24BF4277FF2B" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[434,555,217,244]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FE1F24BF4277FF2B" box="[434,555,217,244]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FD9F24BF4232FF2C" box="[562,622,218,244]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
, see below
<emphasis id="B908EAE4FF8EFFB1FD4A24BF418BFECB" italics="true" pageId="13" pageNumber="86">Individuals with regenerated chelae</emphasis>
.
</paragraph>
<paragraph id="8BC336F6FF8EFFB1FF36257C4153FEAB" blockId="13.[132,776,217,2030]" pageId="13" pageNumber="86">
<taxonomicName id="4C7C4D75FF8EFFB1FF36257C41DBFEEB" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[155,391,281,307]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FF36257C41DBFEEB" box="[155,391,281,307]" italics="true" pageId="13" pageNumber="86">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FE3D257F4192FEEC" box="[400,462,282,308]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
may be easily distinguished from
<taxonomicName id="4C7C4D75FF8EFFB1FF6C255C416DFE8B" authorityName="Guinot" authorityYear="1990" box="[193,305,313,339]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FF6C255C416DFE8B" box="[193,305,313,339]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
(
<figureCitation id="13472A73FF8EFFB1FEED255C41EFFE8B" box="[320,435,313,339]" captionStart="FIG" captionStartId="15.[132,143,1851,1869]" captionTargetBox="[426,1176,216,1815]" captionTargetId="figure@15.[426,1176,216,1816]" captionTargetPageId="15" captionText="FIG. 8. — A, B, Austinograea alayseae Guinot, 1990,photographed in situ: see absence of patches of setae on inner surface of chelae:A, crab with white carapace, Lau Basin 2009 cruise, Kilo Moana site, 20°03.23S, 176°08.01W, 2621 m, among mussels Bathymodiolus brevior Cosel, Métivier &amp; Hashimoto, 1994 and snails Ifremeria nautilei Bouchet &amp; Warén, 1991; B, crab with carapace stained orange due to iron or manganese oxide deposits, and cluster of alvinocaridid shrimps, Lau Basin, TUIM07MV, Kilo Moana site, 20°03.23S, 176°08.01W, 2621 m; C, Austinograea hourdezi n. sp. (presumed identification, see p. 89), Lau Basin, Lau Basin 2009 cruise, Kilo Moana site, among mussels Bathymodiolus brevior, gastropods Ifremeria nautilei, sea anemone probably Cyananthea hourdezi Zelnio, Rodríguez &amp; Daly, 2009, and alvinocaridid shrimp, Alvinocaris komaii Zelnio &amp; Hourdez, 2009. This invidual with palm of preserved left cheliped (regenerated?) bearing only a single dark spot is atypical such as many Austinograea species showing regenerated chelipeds; patches of setae discernible just above on inner surface of chela and on right subhepatic region. Photographies courtesy of C. R. Fisher (Penn State University, USA)." figureDoi="http://doi.org/10.5281/zenodo.3738116" httpUri="https://zenodo.org/record/3738116/files/figure.png" pageId="13" pageNumber="86">Fig. 8A, B</figureCitation>
), with which it cohabits in several sites, by:
</paragraph>
<paragraph id="8BC336F6FF8EFFB1FF36251C4199FAC8" blockId="13.[132,776,217,2030]" pageId="13" pageNumber="86">
1) subhepatic region covered by patch of dense setae in both sexes (no setae in
<taxonomicName id="4C7C4D75FF8EFFB1FEDA25FC41BAFE6A" authorityName="Guinot" authorityYear="1990" box="[375,486,408,435]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FEDA25FC41BAFE6A" box="[375,486,408,435]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
); 2) mxp3: external border oblique, distal end strongly produced (in
<taxonomicName id="4C7C4D75FF8EFFB1FD9D25DC42C7FE0A" authorityName="Guinot" authorityYear="1990" box="[560,667,440,467]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FD9D25DC42C7FE0A" box="[560,667,440,467]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
forming a marked angle:proximal portion vertically oriented, with slightly concave border, half distal portion abruptly tilted; distal end weakly produced); inner margin produced in marked lobe (without lobe in
<taxonomicName id="4C7C4D75FF8EFFB1FE93265D41F3FD8A" authorityName="Guinot" authorityYear="1990" box="[318,431,568,594]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FE93265D41F3FD8A" box="[318,431,568,594]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
); 3) both male chelae with two deep, dark depressions, marked as spots (without any spot in
<taxonomicName id="4C7C4D75FF8EFFB1FF29261D40A8FD4A" authorityName="Guinot" authorityYear="1990" box="[132,244,632,658]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FF29261D40A8FD4A" box="[132,244,632,658]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
); 4) in both sexes patches of dense setae on inner surface of palm and along fingers (without setae in
<taxonomicName id="4C7C4D75FF8EFFB1FD1926FD40F7FD09" authorityName="Guinot" authorityYear="1990" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FD1926FD40F7FD09" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
); 5) marked heterochely: male crusher with short, stout propodus, and short, thick fingers; 6) fixed finger coloured; dactylus coloured on two thirds (in
<taxonomicName id="4C7C4D75FF8EFFB1FDA1269D4220FCC9" authorityName="Guinot" authorityYear="1990" box="[524,636,759,786]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FDA1269D4220FCC9" box="[524,636,759,786]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
pronounced heterochely but propodus of male crusher more elongated; fingers more elongated, narrower); 7) male cutter with occluding margin of dactylus smooth; 8) fixed finger very thick, almost entirely coloured, occluding margin with three small teeth, size progressively diminishing forwards (in
<taxonomicName id="4C7C4D75FF8EFFB1FD3A27F2435BFC69" authorityName="Guinot" authorityYear="1990" box="[663,775,919,945]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FD3A27F2435BFC69" box="[663,775,919,945]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
fingers elongated, armed on both occluding margins with several small teeth interspersed with smaller ones; dactylus coloured at distal end in males and only at tip in females); 9) thoracic sternum with median line at level of sternite 8 and practically not continuying along sternite
<quantity id="4C849B13FF8EFFB1FD3720524295FB88" box="[666,713,1078,1105]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="13" pageNumber="86" unit="in" value="7.0">7 in</quantity>
both sexes (extending on most part of sternite
<quantity id="4C849B13FF8EFFB1FDEB2032422EFBA8" box="[582,626,1110,1137]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="13" pageNumber="86" unit="in" value="7.0">7 in</quantity>
both sexes of
<taxonomicName id="4C7C4D75FF8EFFB1FF29201340A9FB48" authorityName="Guinot" authorityYear="1990" box="[132,245,1142,1168]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FF29201340A9FB48" box="[132,245,1142,1168]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EFED4B07FF8EFFB1FE87201341E5FB48" author="GUINOT D." box="[298,441,1142,1168]" pageId="13" pageNumber="86" pagination="879 - 903" refId="ref26108" refString="GUINOT D. 1990. - Austinograea alayseae sp. nov., crabe hydrothermal decouvert dans le bassin de Lau, Pacifique sud-occidental (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle 4 e ser., 11, section A (4): 879 - 903." type="journal article" year="1990">Guinot 1990</bibRefCitation>
: fig. 2C); 10) male pleon with triangular telson (rounded in
<taxonomicName id="4C7C4D75FF8EFFB1FE6D20F34273FB68" authorityName="Guinot" authorityYear="1990" box="[448,559,1174,1200]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FE6D20F34273FB68" box="[448,559,1174,1200]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
); and 11) G1 rather straigt, with only minute setae along subdistal half (more curved, with several rows of rather long, acute setae along most of length in
<taxonomicName id="4C7C4D75FF8EFFB1FEE8209341E9FAD7" authorityName="Guinot" authorityYear="1990" box="[325,437,1269,1296]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FEE8209341E9FAD7" box="[325,437,1269,1296]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC336F6FF8EFFB1FF3621704448FEAB" blockId="13.[132,776,217,2030]" lastBlockId="13.[813,1456,217,722]" pageId="13" pageNumber="86">
The most distinctive character between
<taxonomicName id="4C7C4D75FF8EFFB1FD8021734354FAF7" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[557,776,1301,1328]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FD8021734354FAF7" box="[557,776,1301,1328]" italics="true" pageId="13" pageNumber="86">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FF29215340E1FA88" box="[132,189,1334,1360]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C7C4D75FF8EFFB1FF4221534136FA97" authorityName="in Guinot" authorityYear="1990" box="[239,362,1333,1360]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FF4221534136FA97" box="[239,362,1333,1360]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
is in the presence of a two spots in male chelae of
<taxonomicName id="4C7C4D75FF8EFFB1FF4621334138FAB7" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[235,356,1365,1392]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FF4621334138FAB7" box="[235,356,1365,1392]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FEC6213341FAFAA8" box="[363,422,1366,1392]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13472A73FF8EFFB1FE1921304256FAB7" box="[436,522,1365,1391]" captionStart="FIG" captionStartId="8.[132,143,1873,1891]" captionTargetBox="[397,1190,214,1823]" captionTargetId="figure@8.[396,927,687,1161]" captionTargetPageId="8" captionText="FIG. 3. — Austinograea hourdezi n. sp. photographed in situ: A, Lau Basin, Lau Basin 2009 cruise, dive 432, Tow Cam site, 20˚18.98S, 176˚08.19W, 2723 m, 05.VI.2009 (type locality), among sessiles barnacles Eochionelasmus ohtai ohtai Yamaguchi, 1990. The two dark spots on outer surface of palm of both chelae indicate a male; see patches of dense setae on inner surface of both chelae, and the darkly coloured two-thirds of dactylus of right cheliped; B, C, Austinograea hourdezi n. sp., due to the patch of dense setae on inner surface of both chelae: B, Lau Basin, TUIM07MV cruise, dive 163, ABE site, 20°45.65S, 176°11.45W, 2130 m, 23.VI.2005; on a bed of gastropod Ifremeria nautilei Bouchet &amp; Warén, 1991; C, Lau Basin, Lau Basin 2009 cruise, dive 431, ABE site, 20°45.65S, 176°11.45W, 2130 m, 03.VI.2009. The crab and the gastropods Alviniconcha hessleri sensu lato are blurry, the photography in situ having been taken in hydrothermal diffuse area with the emanating fluid visible as shimmering water. Photographies courtesy of C. R. Fisher (Penn State University, USA)." figureDoi="http://doi.org/10.5281/zenodo.3738106" httpUri="https://zenodo.org/record/3738106/files/figure.png" pageId="13" pageNumber="86">Figs 3A</figureCitation>
;
<figureCitation id="13472A73FF8EFFB1FDB82130420AFAA8" box="[533,598,1365,1392]" captionStart="FIG" captionStartId="9.[132,143,1920,1938]" captionTargetBox="[132,1457,216,1756]" captionTargetId="figure@9.[699,984,759,1032]" captionTargetPageId="9" captionText="FIG. 4. — A-C, Austinograea hourdezi n. sp., holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, chelae with two coloured spots on outer surface of palm of right crusher (A) and left cutter (B); C, G1 and G2 photographed in situ; D, E, Austinograea hourdezi n. sp., atypical ♀ 30.0 × 48.5 mm (right-handed), Lau Back-Arc Basin, MGLN07MV cruise, dive 230, Kilo Moana site, 20°03.23S, 176°08.01W, 2623 m, MNHN-IU-2016-10752: D, crusher with one marked dark spot and the other one, only discernible by a yellow trace (wound at base of fixed finger is not a spot comparable to that of A. jolliveti n. sp.); E, cutter with two indistinct spots, only as yellow traces." figureDoi="http://doi.org/10.5281/zenodo.3738108" httpUri="https://zenodo.org/record/3738108/files/figure.png" pageId="13" pageNumber="86">4A, B</figureCitation>
;
<figureCitation id="13472A73FF8EFFB1FDCF213042F9FAB7" box="[610,677,1365,1392]" captionStart="FIG" captionStartId="10.[132,143,1965,1983]" captionTargetBox="[132,1455,216,1859]" captionTargetId="figure@10.[698,1267,763,1313]" captionTargetPageId="10" captionText="FIG. 5. — Austinograea hourdezi n. sp., paratype, ♂ 29.8 × 48.1 mm (left-handed, with weak heterochely and heterodonty), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738: A, dorsal view; B, frontal view; C, mxp3; D, ventral view; E, G, both chelae with two typical spots on outer surface; F, H, both chelae with patches of setae on inner surface: E, F, cutter situated at right; G, H, crusher situated at left." figureDoi="http://doi.org/10.5281/zenodo.3738110" httpUri="https://zenodo.org/record/3738110/files/figure.png" pageId="13" pageNumber="86">5E, G</figureCitation>
) ([versus one spot on the palm of crusher and cutter male chelipeds of
<taxonomicName id="4C7C4D75FF8EFFB1FF0F21F0417DFA77" authorityName="in Guinot" authorityYear="1990" box="[162,289,1429,1455]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FF0F21F0417DFA77" box="[162,289,1429,1455]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
, described by
<bibRefCitation id="EFED4B07FF8EFFB1FE1321F0435EFA68" author="HESSLER R. R. &amp; MARTIN J. W." box="[446,770,1429,1456]" pageId="13" pageNumber="86" pagination="645 - 661" refId="ref27182" refString="HESSLER R. R. &amp; MARTIN J. W. 1989. - Austinograea williamsi, new genus, new species, a hydrothermal vent crab (Decapoda: Bythograeidea) from the Mariana Back-Arc Basin, western Pacific. Journal of Crustacean Biology 9 (4): 645 - 661. https: // doi. org / 10.2307 / 1548594" type="journal article" year="1989">Hessler &amp; Martin (1989: 651</bibRefCitation>
, figs 1b, 9a-c) as “small, often brown-stained, pitted area just proximal to, and in line with, ventral border of dactylus” in “larger males” and absent in females (see also
<bibRefCitation id="EFED4B07FF8EFFB1FD292190417AF9F7" author="TSUCHIDA S. &amp; FUJIKURA K." pageId="13" pageNumber="86" pagination="407 - 414" refId="ref30827" refString="TSUCHIDA S. &amp; FUJIKURA K. 2000. - Heterochely, relative growth, and gonopod morphology in the bythograeid crab, Austinograea williamsi (Decapoda, Brachyura). Journal of Crustacean Biology 20 (2): 407 - 414. https: // doi. org / 10.1163 / 20021975 - 99990052" type="journal article" year="2000">Tsuchida &amp; Fujikura 2000</bibRefCitation>
: fig. 2]). Heterochely and heterodonty are pronounced in males of
<taxonomicName id="4C7C4D75FF8EFFB1FE232250425BF996" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[398,519,1588,1615]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FE232250425BF996" box="[398,519,1588,1615]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FDA322504216F997" box="[526,586,1589,1615]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
and
<emphasis id="B908EAE4FF8EFFB1FD2C22504354F996" box="[641,776,1588,1615]" italics="true" pageId="13" pageNumber="86">
<taxonomicName id="4C7C4D75FF8EFFB1FD2C22504358F996" authorityName="in Guinot" authorityYear="1990" box="[641,772,1588,1615]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">A. williamsi</taxonomicName>
.
</emphasis>
The male cutter chela of
<taxonomicName id="4C7C4D75FF8EFFB1FE2122304258F9B6" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[396,516,1620,1647]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FE2122304258F9B6" box="[396,516,1620,1647]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FDA722304218F9B7" box="[522,580,1621,1647]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
has a thick, broad fixed finger, and an occluding margin armed with a few teeth. In
<taxonomicName id="4C7C4D75FF8EFFB1FF0822F1417AF976" authorityName="in Guinot" authorityYear="1990" box="[165,294,1684,1710]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FF0822F1417AF976" box="[165,294,1684,1710]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
the cutter chela is more elongate; the fixed finger is more inflated and broader, with a deeply dished inner surface, and a markedly convex, much toothed occluding margin that fits flush against dactylus. In
<taxonomicName id="4C7C4D75FF8EFFB1FDE722914298F8D6" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[586,708,1780,1806]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FDE722914298F8D6" box="[586,708,1780,1806]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FD662291435BF8D6" box="[715,775,1780,1806]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
the chelae show a sexual dimorphism that is distinctive from that of
<taxonomicName id="4C7C4D75FF8EFFB1FF6323514115F895" authorityName="in Guinot" authorityYear="1990" box="[206,329,1843,1870]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FF6323514115F895" box="[206,329,1843,1870]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFED4B07FF8EFFB1FED62356422DF896" author="HESSLER R. R. &amp; MARTIN J. W." box="[379,625,1843,1870]" pageId="13" pageNumber="86" pagination="645 - 661" refId="ref27182" refString="HESSLER R. R. &amp; MARTIN J. W. 1989. - Austinograea williamsi, new genus, new species, a hydrothermal vent crab (Decapoda: Bythograeidea) from the Mariana Back-Arc Basin, western Pacific. Journal of Crustacean Biology 9 (4): 645 - 661. https: // doi. org / 10.2307 / 1548594" type="journal article" year="1989">Hessler &amp; Martin 1989</bibRefCitation>
: figs 9, 10), in which females have cutters for both chelipeds with a markedly concave propodal inner margin and a well dished fixed finger at occluding margin that bears numerous, sharper, delicate teeth. The mxp3 merus of
<taxonomicName id="4C7C4D75FF8EFFB1FE3023D64248F815" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[413,532,1971,1997]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FE3023D64248F815" box="[413,532,1971,1997]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FDB723D14208F816" box="[538,596,1972,1998]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
is very similar to that of
<taxonomicName id="4C7C4D75FF8EFFB1FF7F23B6410DF835" authorityName="in Guinot" authorityYear="1990" box="[210,337,2003,2029]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FF7F23B6410DF835" box="[210,337,2003,2029]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
, both having an oblique external border, a produced ending and a pronounced inner lobe. Another marked difference is the subhepatic region: covered by dense setae in
<taxonomicName id="4C7C4D75FF8EFFB1FC2C257C43A4FEEB" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[897,1016,281,307]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FC2C257C43A4FEEB" box="[897,1016,281,307]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FC53257F4465FEEC" box="[1022,1081,282,308]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
, whereas the patch of setae is much more larger in
<taxonomicName id="4C7C4D75FF8EFFB1FC67255C4415FE8B" authorityName="in Guinot" authorityYear="1990" box="[970,1097,313,339]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FC67255C4415FE8B" box="[970,1097,313,339]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
, in which it considerably extends below the lateral line.
</paragraph>
<paragraph id="8BC336F6FF8EFFB1FCEE251C4413FD09" blockId="13.[813,1456,217,722]" pageId="13" pageNumber="86">
For the distinction between
<taxonomicName id="4C7C4D75FF8EFFB1FBD1251C44B1FE4A" authorityName="Guinot" authorityYear="1990" box="[1148,1261,376,403]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FBD1251C44B1FE4A" box="[1148,1261,376,403]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7C4D75FF8EFFB1FA85251C45F4FE4A" authorityName="in Guinot" authorityYear="1990" box="[1320,1448,376,403]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FA85251C45F4FE4A" box="[1320,1448,376,403]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
, see the comparative illustrations in
<bibRefCitation id="EFED4B07FF8EFFB1FB0025FD4325FE0B" author="TSUCHIDA S. &amp; HASHIMOTO J." pageId="13" pageNumber="86" pagination="642 - 650" refId="ref30883" refString="TSUCHIDA S. &amp; HASHIMOTO J. 2002. - A new species of bythograeid crab, Austinograea rodriguezensis (Decapoda, Brachyura), associated with active hydrothermal vents from the Indian Ocean. Journal of Crustacean Biology 22 (3): 642 - 650. https: // doi. org / 10.1163 / 20021975 - 99990276" type="journal article" year="2002">Tsuchida &amp; Hashimoto (2002)</bibRefCitation>
. The inner surface of palm and most inner surface of the dactylus of the two chelae bear patches of setae in both sexes of
<taxonomicName id="4C7C4D75FF8EFFB1FC24259C445FFDCA" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[905,1027,504,531]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FC24259C445FFDCA" box="[905,1027,504,531]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FBA1259C4415FDCB" box="[1036,1097,505,531]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13472A73FF8EFFB1FBF4259D44C7FDCA" box="[1113,1179,504,530]" captionStart="FIG" captionStartId="8.[132,143,1873,1891]" captionTargetBox="[397,1190,214,1823]" captionTargetId="figure@8.[396,927,687,1161]" captionTargetPageId="8" captionText="FIG. 3. — Austinograea hourdezi n. sp. photographed in situ: A, Lau Basin, Lau Basin 2009 cruise, dive 432, Tow Cam site, 20˚18.98S, 176˚08.19W, 2723 m, 05.VI.2009 (type locality), among sessiles barnacles Eochionelasmus ohtai ohtai Yamaguchi, 1990. The two dark spots on outer surface of palm of both chelae indicate a male; see patches of dense setae on inner surface of both chelae, and the darkly coloured two-thirds of dactylus of right cheliped; B, C, Austinograea hourdezi n. sp., due to the patch of dense setae on inner surface of both chelae: B, Lau Basin, TUIM07MV cruise, dive 163, ABE site, 20°45.65S, 176°11.45W, 2130 m, 23.VI.2005; on a bed of gastropod Ifremeria nautilei Bouchet &amp; Warén, 1991; C, Lau Basin, Lau Basin 2009 cruise, dive 431, ABE site, 20°45.65S, 176°11.45W, 2130 m, 03.VI.2009. The crab and the gastropods Alviniconcha hessleri sensu lato are blurry, the photography in situ having been taken in hydrothermal diffuse area with the emanating fluid visible as shimmering water. Photographies courtesy of C. R. Fisher (Penn State University, USA)." figureDoi="http://doi.org/10.5281/zenodo.3738106" httpUri="https://zenodo.org/record/3738106/files/figure.png" pageId="13" pageNumber="86">Figs 3</figureCitation>
A-C; 5A, F, H; 7A, B, F, H), as in
<taxonomicName id="4C7C4D75FF8EFFB1FC23267D4457FDEA" authorityName="in Guinot" authorityYear="1990" box="[910,1035,536,562]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FC23267D4457FDEA" box="[910,1035,536,562]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
, whereas in both sexes of
<taxonomicName id="4C7C4D75FF8EFFB1FAB4267D45DBFDEA" authorityName="Guinot" authorityYear="1990" box="[1305,1415,536,562]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FAB4267D45DBFDEA" box="[1305,1415,536,562]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
the chelae of both sides are glabrous on palm and dactylus. The G1 and G2 of
<taxonomicName id="4C7C4D75FF8EFFB1FC63263D4414FDAA" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[974,1096,600,626]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FC63263D4414FDAA" box="[974,1096,600,626]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FBE2263D44D6FDAA" box="[1103,1162,600,626]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C7C4D75FF8EFFB1FB6C263D451CFDAA" authorityName="in Guinot" authorityYear="1990" box="[1217,1344,600,626]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FB6C263D451CFDAA" box="[1217,1344,600,626]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
(in which the G2 is about less than half the G1s length) are similar, whereas in
<taxonomicName id="4C7C4D75FF8EFFB1FC0C26FD4453FD6A" authorityName="Guinot" authorityYear="1990" box="[929,1039,664,690]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FC0C26FD4453FD6A" box="[929,1039,664,690]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
the G1 bears numerous long setae scattered over its entire length.
</paragraph>
<paragraph id="8BC336F6FF8EFFB1FC802692441EFCD7" blockId="13.[813,1456,759,1392]" box="[813,1090,759,785]" pageId="13" pageNumber="86">DISTINCTION OF FEMALES</paragraph>
<paragraph id="8BC336F6FF8EFFB1FC80277244AAFAA8" blockId="13.[813,1456,759,1392]" pageId="13" pageNumber="86">
As the spots on the cheliped propodus are typically absent in female
<taxonomicName id="4C7C4D75FF8EFFB1FCD4275243ADFC89" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[889,1009,823,849]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FCD4275243ADFC89" box="[889,1009,823,849]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FC5A275D446DFC8A" box="[1015,1073,824,850]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
and in both sexes of
<taxonomicName id="4C7C4D75FF8EFFB1FABC275245DCFC89" authorityName="Guinot" authorityYear="1990" box="[1297,1408,823,849]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FABC275245DCFC89" box="[1297,1408,823,849]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
, the more useful significant character to differentiate the females of the two species is the patch of setae at the inner surface of the palm of females of
<taxonomicName id="4C7C4D75FF8EFFB1FB8B27F244C3FC69" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[1062,1183,919,945]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FB8B27F244C3FC69" box="[1062,1183,919,945]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FB0B27F244BDFC69" box="[1190,1249,919,945]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13472A73FF8EFFB1FB5D27F245FDFC69" box="[1264,1441,919,945]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1858]" captionTargetId="figure@14.[699,984,1036,1311]" captionTargetPageId="14" captionText="FIG. 7. — Austinograea hourdezi n. sp., paratype, ♀ 18.3 × 28.4 mm (right-handed), Lau Back-Arc Basin, BIOLAU cruise, BL 12, Vai Lili site, 23°13S, 176°38W, 1750 m, MNHN-IU-2016-10745 (= MNHN-B24060): A, dorsal view; B, C, ventral views, with pleon; D, thoracic sternum and vulvae; E, G, both chelae without spots on outer surface; F, H, both chelae with patches of setae on inner surface; E, F, right crusher; G, H, left cutter." figureDoi="http://doi.org/10.5281/zenodo.3738114" httpUri="https://zenodo.org/record/3738114/files/figure.png" pageId="13" pageNumber="86">Fig. 7A, B, F, H</figureCitation>
), as in the males (
<figureCitation id="13472A73FF8EFFB1FC7927D24475FC09" box="[980,1065,951,977]" captionStart="FIG" captionStartId="8.[132,143,1873,1891]" captionTargetBox="[397,1190,214,1823]" captionTargetId="figure@8.[396,927,687,1161]" captionTargetPageId="8" captionText="FIG. 3. — Austinograea hourdezi n. sp. photographed in situ: A, Lau Basin, Lau Basin 2009 cruise, dive 432, Tow Cam site, 20˚18.98S, 176˚08.19W, 2723 m, 05.VI.2009 (type locality), among sessiles barnacles Eochionelasmus ohtai ohtai Yamaguchi, 1990. The two dark spots on outer surface of palm of both chelae indicate a male; see patches of dense setae on inner surface of both chelae, and the darkly coloured two-thirds of dactylus of right cheliped; B, C, Austinograea hourdezi n. sp., due to the patch of dense setae on inner surface of both chelae: B, Lau Basin, TUIM07MV cruise, dive 163, ABE site, 20°45.65S, 176°11.45W, 2130 m, 23.VI.2005; on a bed of gastropod Ifremeria nautilei Bouchet &amp; Warén, 1991; C, Lau Basin, Lau Basin 2009 cruise, dive 431, ABE site, 20°45.65S, 176°11.45W, 2130 m, 03.VI.2009. The crab and the gastropods Alviniconcha hessleri sensu lato are blurry, the photography in situ having been taken in hydrothermal diffuse area with the emanating fluid visible as shimmering water. Photographies courtesy of C. R. Fisher (Penn State University, USA)." figureDoi="http://doi.org/10.5281/zenodo.3738106" httpUri="https://zenodo.org/record/3738106/files/figure.png" pageId="13" pageNumber="86">Figs 3A</figureCitation>
;
<figureCitation id="13472A73FF8EFFB1FB9E27D344CCFC09" box="[1075,1168,950,977]" captionStart="FIG" captionStartId="10.[132,143,1965,1983]" captionTargetBox="[132,1455,216,1859]" captionTargetId="figure@10.[698,1267,763,1313]" captionTargetPageId="10" captionText="FIG. 5. — Austinograea hourdezi n. sp., paratype, ♂ 29.8 × 48.1 mm (left-handed, with weak heterochely and heterodonty), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738: A, dorsal view; B, frontal view; C, mxp3; D, ventral view; E, G, both chelae with two typical spots on outer surface; F, H, both chelae with patches of setae on inner surface: E, F, cutter situated at right; G, H, crusher situated at left." figureDoi="http://doi.org/10.5281/zenodo.3738110" httpUri="https://zenodo.org/record/3738110/files/figure.png" pageId="13" pageNumber="86">5A, F, H</figureCitation>
). In contrast, in
<taxonomicName id="4C7C4D75FF8EFFB1FAED27D245F3FC09" authorityName="Guinot" authorityYear="1990" box="[1344,1455,951,977]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FAED27D245F3FC09" box="[1344,1455,951,977]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
(
<figureCitation id="13472A73FF8EFFB1FC9927B343F4FC29" box="[820,936,982,1009]" captionStart="FIG" captionStartId="15.[132,143,1851,1869]" captionTargetBox="[426,1176,216,1815]" captionTargetId="figure@15.[426,1176,216,1816]" captionTargetPageId="15" captionText="FIG. 8. — A, B, Austinograea alayseae Guinot, 1990,photographed in situ: see absence of patches of setae on inner surface of chelae:A, crab with white carapace, Lau Basin 2009 cruise, Kilo Moana site, 20°03.23S, 176°08.01W, 2621 m, among mussels Bathymodiolus brevior Cosel, Métivier &amp; Hashimoto, 1994 and snails Ifremeria nautilei Bouchet &amp; Warén, 1991; B, crab with carapace stained orange due to iron or manganese oxide deposits, and cluster of alvinocaridid shrimps, Lau Basin, TUIM07MV, Kilo Moana site, 20°03.23S, 176°08.01W, 2621 m; C, Austinograea hourdezi n. sp. (presumed identification, see p. 89), Lau Basin, Lau Basin 2009 cruise, Kilo Moana site, among mussels Bathymodiolus brevior, gastropods Ifremeria nautilei, sea anemone probably Cyananthea hourdezi Zelnio, Rodríguez &amp; Daly, 2009, and alvinocaridid shrimp, Alvinocaris komaii Zelnio &amp; Hourdez, 2009. This invidual with palm of preserved left cheliped (regenerated?) bearing only a single dark spot is atypical such as many Austinograea species showing regenerated chelipeds; patches of setae discernible just above on inner surface of chela and on right subhepatic region. Photographies courtesy of C. R. Fisher (Penn State University, USA)." figureDoi="http://doi.org/10.5281/zenodo.3738116" httpUri="https://zenodo.org/record/3738116/files/figure.png" pageId="13" pageNumber="86">Fig. 8A, B</figureCitation>
) the chelae are devoid of setae, in females as in males. The same character allows identification of females of
<taxonomicName id="4C7C4D75FF8EFFB1FCE620724397FBE8" authorityName="in Guinot" authorityYear="1990" box="[843,971,1046,1073]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FCE620724397FBE8" box="[843,971,1046,1073]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
, with setose patch at the inner chela surface (
<bibRefCitation id="EFED4B07FF8EFFB1FC9820534472FB88" author="HESSLER R. R. &amp; MARTIN J. W." box="[821,1070,1078,1104]" pageId="13" pageNumber="86" pagination="645 - 661" refId="ref27182" refString="HESSLER R. R. &amp; MARTIN J. W. 1989. - Austinograea williamsi, new genus, new species, a hydrothermal vent crab (Decapoda: Bythograeidea) from the Mariana Back-Arc Basin, western Pacific. Journal of Crustacean Biology 9 (4): 645 - 661. https: // doi. org / 10.2307 / 1548594" type="journal article" year="1989">Hessler &amp; Martin 1989</bibRefCitation>
: fig. 10), from females of
<taxonomicName id="4C7C4D75FF8EFFB1FA96205245F4FB88" authorityName="Guinot" authorityYear="1990" box="[1339,1448,1078,1105]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FA96205245F4FB88" box="[1339,1448,1078,1105]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
, which have glabrous chelae. See the comparative illustration of female
<taxonomicName id="4C7C4D75FF8EFFB1FC3220134475FB48" authorityName="Hessler &amp; Martin" authorityYear="1989" box="[927,1065,1142,1168]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="genus">
<emphasis id="B908EAE4FF8EFFB1FC3220134475FB48" box="[927,1065,1142,1168]" italics="true" pageId="13" pageNumber="86">Austinograea</emphasis>
</taxonomicName>
chelae by
<bibRefCitation id="EFED4B07FF8EFFB1FB0F20134331FB68" author="TSUCHIDA S. &amp; HASHIMOTO J." pageId="13" pageNumber="86" pagination="642 - 650" refId="ref30883" refString="TSUCHIDA S. &amp; HASHIMOTO J. 2002. - A new species of bythograeid crab, Austinograea rodriguezensis (Decapoda, Brachyura), associated with active hydrothermal vents from the Indian Ocean. Journal of Crustacean Biology 22 (3): 642 - 650. https: // doi. org / 10.1163 / 20021975 - 99990276" type="journal article" year="2002">Tsuchida &amp; Hashimoto (2002</bibRefCitation>
: fig. 6), which does not show any spot in
<taxonomicName id="4C7C4D75FF8EFFB1FA9C20F345ECFB68" authorityName="in Guinot" authorityYear="1990" box="[1329,1456,1174,1200]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF8EFFB1FA9C20F345ECFB68" box="[1329,1456,1174,1200]" italics="true" pageId="13" pageNumber="86">A. williamsi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7C4D75FF8EFFB1FCF020D34393FB08" authorityName="Guinot" authorityYear="1990" box="[861,975,1206,1232]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FCF020D34393FB08" box="[861,975,1206,1232]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
, in contrast with
<taxonomicName id="4C7C4D75FF8EFFB1FB3820D34519FB08" authorityName="Tsuchida &amp; Hashimoto" authorityYear="2002" box="[1173,1349,1206,1232]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="rodriguezensis">
<emphasis id="B908EAE4FF8EFFB1FB3820D34519FB08" box="[1173,1349,1206,1232]" italics="true" pageId="13" pageNumber="86">A. rodriguezensis</emphasis>
</taxonomicName>
, which is shown bearing a spot, as in the males. In
<taxonomicName id="4C7C4D75FF8EFFB1FB4020B34502FB28" authorityName="Guinot" authorityYear="1990" box="[1261,1374,1238,1264]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8EFFB1FB4020B34502FB28" box="[1261,1374,1238,1264]" italics="true" pageId="13" pageNumber="86">A. alayseae</emphasis>
</taxonomicName>
a thick posterior membrane may cover most of the vulva, leaving a variously developed opening, sometimes only a lateral slit, whereas in
<taxonomicName id="4C7C4D75FF8EFFB1FC0A2153447CFA97" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[935,1056,1333,1360]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FC0A2153447CFA97" box="[935,1056,1333,1360]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FB8521534438FA88" box="[1064,1124,1334,1360]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
the medial opening is entirely surrounded by thick membrane (
<figureCitation id="13472A73FF8EFFB1FB3C213044B5FAA8" box="[1169,1257,1365,1392]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1858]" captionTargetId="figure@14.[699,984,1036,1311]" captionTargetPageId="14" captionText="FIG. 7. — Austinograea hourdezi n. sp., paratype, ♀ 18.3 × 28.4 mm (right-handed), Lau Back-Arc Basin, BIOLAU cruise, BL 12, Vai Lili site, 23°13S, 176°38W, 1750 m, MNHN-IU-2016-10745 (= MNHN-B24060): A, dorsal view; B, C, ventral views, with pleon; D, thoracic sternum and vulvae; E, G, both chelae without spots on outer surface; F, H, both chelae with patches of setae on inner surface; E, F, right crusher; G, H, left cutter." figureDoi="http://doi.org/10.5281/zenodo.3738114" httpUri="https://zenodo.org/record/3738114/files/figure.png" pageId="13" pageNumber="86">Fig. 7D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C366657DFF8EFFACFC8021F14120FEEC" lastPageId="16" lastPageNumber="89" pageId="13" pageNumber="86" type="description">
<paragraph id="8BC336F6FF8EFFB1FC8021F144B0FA75" blockId="13.[813,1456,1428,2030]" box="[813,1260,1428,1454]" pageId="13" pageNumber="86">INDIVIDUALS WITH REGENERATED CHELAE</paragraph>
<paragraph id="8BC336F6FF8EFFB1FC8021D043E5F876" blockId="13.[813,1456,1428,2030]" pageId="13" pageNumber="86">
A few male specimens that display most characters of
<taxonomicName id="4C7C4D75FF8EFFB1FAF721D04304FA37" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FAF721D04304FA37" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FCCD21B043C1FA37" box="[864,925,1493,1519]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
(including mxp3 shape and patches of setae on inner surface of chelae) exhibit a single spot on the outer surface chelae palm, instead of two. The
<typeStatus id="54C78854FF8EFFB1FB7122714567F9F6" box="[1244,1339,1556,1582]" pageId="13" pageNumber="86" type="holotype">holotype</typeStatus>
(MNHN- IU-2016-10737), from the Lau Basin, Tow Cam site, a large right-handed male 25.7 ×
<quantity id="4C849B13FF8EFFB1FBE7223144EDF9B7" box="[1098,1201,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="4.02" pageId="13" pageNumber="86" unit="mm" value="40.2">40.2 mm</quantity>
, typically bears the two characteristic spots on each chela (
<figureCitation id="13472A73FF8EFFB1FB322211454EF957" box="[1183,1298,1652,1679]" captionStart="FIG" captionStartId="9.[132,143,1920,1938]" captionTargetBox="[132,1457,216,1756]" captionTargetId="figure@9.[699,984,759,1032]" captionTargetPageId="9" captionText="FIG. 4. — A-C, Austinograea hourdezi n. sp., holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, chelae with two coloured spots on outer surface of palm of right crusher (A) and left cutter (B); C, G1 and G2 photographed in situ; D, E, Austinograea hourdezi n. sp., atypical ♀ 30.0 × 48.5 mm (right-handed), Lau Back-Arc Basin, MGLN07MV cruise, dive 230, Kilo Moana site, 20°03.23S, 176°08.01W, 2623 m, MNHN-IU-2016-10752: D, crusher with one marked dark spot and the other one, only discernible by a yellow trace (wound at base of fixed finger is not a spot comparable to that of A. jolliveti n. sp.); E, cutter with two indistinct spots, only as yellow traces." figureDoi="http://doi.org/10.5281/zenodo.3738108" httpUri="https://zenodo.org/record/3738108/files/figure.png" pageId="13" pageNumber="86">Fig. 4A, B</figureCitation>
), such as most of other males that we have examined. Although left-handed and with weak heterochely and heterodonty, a larger male from the same site 29.8 ×
<quantity id="4C849B13FF8EFFB1FBF022B14495F936" box="[1117,1225,1748,1774]" metricMagnitude="-2" metricUnit="m" metricValue="4.81" pageId="13" pageNumber="86" unit="mm" value="48.1">48.1 mm</quantity>
,
<typeStatus id="54C78854FF8EFFB1FB7A22B14566F936" box="[1239,1338,1748,1774]" pageId="13" pageNumber="86" type="paratype">paratype</typeStatus>
, MNHN- IU-2016-10738, also shows the two typical spots on chelae, even on regenerated cheliped (
<figureCitation id="13472A73FF8EFFB1FBC3237144BDF8F6" box="[1134,1249,1811,1838]" captionStart="FIG" captionStartId="10.[132,143,1965,1983]" captionTargetBox="[132,1455,216,1859]" captionTargetId="figure@10.[698,1267,763,1313]" captionTargetPageId="10" captionText="FIG. 5. — Austinograea hourdezi n. sp., paratype, ♂ 29.8 × 48.1 mm (left-handed, with weak heterochely and heterodonty), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site,MNHN-IU-2016-10738: A, dorsal view; B, frontal view; C, mxp3; D, ventral view; E, G, both chelae with two typical spots on outer surface; F, H, both chelae with patches of setae on inner surface: E, F, cutter situated at right; G, H, crusher situated at left." figureDoi="http://doi.org/10.5281/zenodo.3738110" httpUri="https://zenodo.org/record/3738110/files/figure.png" pageId="13" pageNumber="86">Fig. 5E, G</figureCitation>
). But a smaller lefthanded male 24.7 × 36.0 mm, from the same sample, has an obviously regenerated minor chela that bears only one spot; its left chela, having become the major chela, shows the two typical spots.
</paragraph>
<paragraph id="8BC336F6FF8EFFACFCEE23D14120FEEC" blockId="13.[813,1456,1428,2030]" lastBlockId="16.[132,776,217,2030]" lastPageId="16" lastPageNumber="89" pageId="13" pageNumber="86">
Two samples of female
<taxonomicName id="4C7C4D75FF8EFFB1FB9023D644EAF815" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[1085,1206,1971,1997]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="86" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8EFFB1FB9023D644EAF815" box="[1085,1206,1971,1997]" italics="true" pageId="13" pageNumber="86">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8EFFB1FB1123D144A5F816" box="[1212,1273,1972,1998]" pageId="13" pageNumber="86" rank="species">n. sp.</taxonomicNameLabel>
, instead of being devoid of any spot on the chela, bear a spot on both chelae: they are left-handed, which means that one chela was regenerated. These females that bear a spot on each cheliped, crusher and cutter, are:
</paragraph>
</subSubSection>
<caption id="DF03667EFF8DFFB2FF2923C94410F833" ID-DOI="http://doi.org/10.5281/zenodo.3738114" ID-Zenodo-Dep="3738114" httpUri="https://zenodo.org/record/3738114/files/figure.png" pageId="14" pageNumber="87" startId="14.[132,143,1964,1982]" targetBox="[132,1455,215,1858]" targetPageId="14">
<paragraph id="8BC336F6FF8DFFB2FF2923C94410F833" blockId="14.[132,1457,1964,2027]" pageId="14" pageNumber="87">
FIG. 7. —
<taxonomicName id="4C7C4D75FF8DFFB2FF7923C941D2F866" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[212,398,1964,1982]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="87" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8DFFB2FF7923C941D2F866" box="[212,398,1964,1982]" italics="true" pageId="14" pageNumber="87">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8DFFB2FE3E23C94199F866" box="[403,453,1964,1982]" pageId="14" pageNumber="87" rank="species">n. sp.</taxonomicNameLabel>
, paratype, ♀ 18.3 × 28.4 mm (right-handed), Lau Back-Arc Basin, BIOLAU cruise, BL 12, Vai Lili site, 23°13S, 176°38W, 1750 m, MNHN-IU-2016-10745 (= MNHN-B24060):
<emphasis id="B908EAE4FF8DFFB2FD9423A6421AF80D" bold="true" box="[569,582,1987,2005]" pageId="14" pageNumber="87">A</emphasis>
, dorsal view;
<emphasis id="B908EAE4FF8DFFB2FD1623A64294F80D" bold="true" box="[699,712,1987,2005]" pageId="14" pageNumber="87">B</emphasis>
,
<emphasis id="B908EAE4FF8DFFB2FD7923A642BEF80D" bold="true" box="[724,738,1987,2005]" pageId="14" pageNumber="87">C</emphasis>
, ventral views, with pleon;
<emphasis id="B908EAE4FF8DFFB2FC6B23A64388F80D" bold="true" box="[966,980,1987,2005]" pageId="14" pageNumber="87">D</emphasis>
, thoracic sternum and vulvae;
<emphasis id="B908EAE4FF8DFFB2FB7423A644B9F80D" bold="true" box="[1241,1253,1987,2005]" pageId="14" pageNumber="87">E</emphasis>
,
<emphasis id="B908EAE4FF8DFFB2FB5D23A744A2F80C" bold="true" box="[1264,1278,1986,2004]" pageId="14" pageNumber="87">G</emphasis>
, both chelae without spots on outer surface;
<emphasis id="B908EAE4FF8DFFB2FEE723BC4109F833" bold="true" box="[330,341,2009,2027]" pageId="14" pageNumber="87">F</emphasis>
,
<emphasis id="B908EAE4FF8DFFB2FECD23BC4132F833" bold="true" box="[352,366,2009,2027]" pageId="14" pageNumber="87">H</emphasis>
, both chelae with patches of setae on inner surface;
<emphasis id="B908EAE4FF8DFFB2FC8A23BC436FF833" bold="true" box="[807,819,2009,2027]" pageId="14" pageNumber="87">E</emphasis>
,
<emphasis id="B908EAE4FF8DFFB2FC9323BC4315F833" bold="true" box="[830,841,2009,2027]" pageId="14" pageNumber="87">F</emphasis>
, right crusher;
<emphasis id="B908EAE4FF8DFFB2FC6A23BC4389F833" bold="true" box="[967,981,2009,2027]" pageId="14" pageNumber="87">G</emphasis>
,
<emphasis id="B908EAE4FF8DFFB2FC7223BC43B1F833" bold="true" box="[991,1005,2009,2027]" pageId="14" pageNumber="87">H</emphasis>
, left cutter.
</paragraph>
</caption>
<caption id="DF03667EFF8CFFB3FF29235E440FF833" ID-DOI="http://doi.org/10.5281/zenodo.3738116" ID-Zenodo-Dep="3738116" httpUri="https://zenodo.org/record/3738116/files/figure.png" pageId="15" pageNumber="88" startId="15.[132,143,1851,1869]" targetBox="[426,1176,216,1815]" targetPageId="15">
<paragraph id="8BC336F6FF8CFFB3FF29235E440FF833" blockId="15.[132,1457,1851,2028]" pageId="15" pageNumber="88">
FIG. 8. —
<emphasis id="B908EAE4FF8CFFB3FF7F235E4083F895" bold="true" box="[210,223,1851,1869]" pageId="15" pageNumber="88">A</emphasis>
,
<emphasis id="B908EAE4FF8CFFB3FF4A235E40A8F895" bold="true" box="[231,244,1851,1869]" pageId="15" pageNumber="88">B</emphasis>
,
<taxonomicName id="4C7C4D75FF8CFFB3FF53235E427BF895" authority="Guinot" authorityName="Guinot" authorityYear="1990" box="[254,551,1851,1869]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF8CFFB3FF53235E41EAF895" box="[254,438,1851,1869]" italics="true" pageId="15" pageNumber="88">Austinograea alayseae</emphasis>
<bibRefCitation id="EFED4B07FF8CFFB3FE17235E427BF895" author="GUINOT D." box="[442,551,1851,1869]" pageId="15" pageNumber="88" pagination="879 - 903" refId="ref26108" refString="GUINOT D. 1990. - Austinograea alayseae sp. nov., crabe hydrothermal decouvert dans le bassin de Lau, Pacifique sud-occidental (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle 4 e ser., 11, section A (4): 879 - 903." type="journal article" year="1990">Guinot, 1990</bibRefCitation>
</taxonomicName>
,photographed
<emphasis id="B908EAE4FF8CFFB3FD0A235E4280F895" box="[679,732,1851,1869]" italics="true" pageId="15" pageNumber="88">in situ:</emphasis>
see absence of patches of setae on inner surface of chelae:
<emphasis id="B908EAE4FF8CFFB3FB6A235E4488F895" bold="true" box="[1223,1236,1851,1869]" pageId="15" pageNumber="88">A</emphasis>
, crab with white carapace, Lau Basin 2009 cruise, Kilo Moana site, 20°03.23S, 176°08.01W, 2621 m, among mussels
<taxonomicName id="4C7C4D75FF8CFFB3FCD42337450BF8BC" authority="Cosel" authorityName="Cosel" authorityYear="1994" box="[889,1367,1873,1892]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytiloida" pageId="15" pageNumber="88" phylum="Mollusca" rank="species" species="brevior">
<emphasis id="B908EAE4FF8CFFB3FCD42337446FF8BC" box="[889,1075,1874,1892]" italics="true" pageId="15" pageNumber="88">Bathymodiolus brevior</emphasis>
<bibRefCitation id="EFED4B07FF8CFFB3FB952334450BF8BC" author="COSEL R. &amp; VON &amp; METIVIER B. &amp; HASHIMOTO J." box="[1080,1367,1873,1892]" pageId="15" pageNumber="88" pagination="374 - 392" refId="ref24757" refString="COSEL R. VON, METIVIER B. &amp; HASHIMOTO J. 1994. - Three new species of Bathymodiolus (Bivalvia: Mytilidae) from hydrothermal vents in the Lau basin and the North Fiji Basin, Western Pacific, and the Snake Pit Area, Mid-Atlantic Ridge. Veliger 37: 374 - 392." type="journal article" year="1994">Cosel, Métivier &amp; Hashimoto, 1994</bibRefCitation>
</taxonomicName>
and snails
<taxonomicName id="4C7C4D75FF8CFFB3FF29230D4186F8A2" authority="Bouchet &amp; Waren, 1991" authorityName="Bouchet &amp; Waren" authorityYear="1991" box="[132,474,1896,1914]" class="Gastropoda" family="Provannidae" genus="Ifremeria" kingdom="Animalia" pageId="15" pageNumber="88" phylum="Mollusca" rank="species" species="nautilei">
<emphasis id="B908EAE4FF8CFFB3FF29230D4151F8A2" box="[132,269,1896,1914]" italics="true" pageId="15" pageNumber="88">Ifremeria nautilei</emphasis>
Bouchet &amp; Warén, 1991
</taxonomicName>
;
<emphasis id="B908EAE4FF8CFFB3FE4F230C41B3F8A3" bold="true" box="[482,495,1897,1915]" pageId="15" pageNumber="88">B</emphasis>
, crab with carapace stained orange due to iron or manganese oxide deposits, and cluster of alvinocaridid shrimps, Lau Basin, TUIM07MV, Kilo Moana site, 20°03.23S, 176°08.01W, 2621 m;
<emphasis id="B908EAE4FF8CFFB3FD55231A435AF849" bold="true" box="[760,774,1919,1937]" pageId="15" pageNumber="88">C</emphasis>
,
<taxonomicName id="4C7C4D75FF8CFFB3FCBC231A4396F849" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[785,970,1919,1937]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF8CFFB3FCBC231A4396F849" box="[785,970,1919,1937]" italics="true" pageId="15" pageNumber="88">Austinograea hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF8CFFB3FC7D231A445CF849" box="[976,1024,1919,1937]" pageId="15" pageNumber="88" rank="species">n. sp.</taxonomicNameLabel>
(presumed identification, see p. 89), Lau Basin, Lau Basin 2009 cruise, Kilo Moana site, among mussels
<taxonomicName id="4C7C4D75FF8CFFB3FD9023F342A4F870" authorityName="Cosel, Metivier &amp; Hashimoto" authorityYear="1994" box="[573,760,1942,1960]" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytiloida" pageId="15" pageNumber="88" phylum="Mollusca" rank="species" species="brevior">
<emphasis id="B908EAE4FF8CFFB3FD9023F342A4F870" box="[573,760,1942,1960]" italics="true" pageId="15" pageNumber="88">Bathymodiolus brevior</emphasis>
</taxonomicName>
, gastropods
<taxonomicName id="4C7C4D75FF8CFFB3FCC523F343ADF870" authorityName="Bouchet &amp; Waren" authorityYear="1991" box="[872,1009,1942,1960]" class="Gastropoda" family="Provannidae" genus="Ifremeria" kingdom="Animalia" pageId="15" pageNumber="88" phylum="Mollusca" rank="species" species="nautilei">
<emphasis id="B908EAE4FF8CFFB3FCC523F343ADF870" box="[872,1009,1942,1960]" italics="true" pageId="15" pageNumber="88">Ifremeria nautilei</emphasis>
</taxonomicName>
, sea anemone probably
<taxonomicName id="4C7C4D75FF8CFFB3FB6F23F04110F866" authority="Zelnio, Rodriguez &amp; Daly, 2009" authorityName="Zelnio, Rodriguez &amp; Daly" authorityYear="2009" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="15" pageNumber="88" phylum="Cnidaria" rank="species" species="hourdezi">
<emphasis id="B908EAE4FF8CFFB3FB6F23F0452DF870" box="[1218,1393,1941,1960]" italics="true" pageId="15" pageNumber="88">Cyananthea hourdezi</emphasis>
<bibRefCitation id="EFED4B07FF8CFFB3FADA23F34110F866" author="ZELNIO K. A. &amp; RODRIGUEZ R. &amp; DALY M." pageId="15" pageNumber="88" pagination="547 - 571" refId="ref32533" refString="ZELNIO K. A., RODRIGUEZ R. &amp; DALY M. 2009. - Hexacorals (Anthozoa: Actiniaria, Zoanthidea) from hydrothermal vents in the south-western Pacific. Marine Biology Research 5: 547 - 571. https: // doi. org / 10.1080 / 17451000902729662" type="journal article" year="2009">Zelnio, Rodríguez &amp; Daly, 2009</bibRefCitation>
</taxonomicName>
, and alvinocaridid shrimp,
<taxonomicName id="4C7C4D75FF8CFFB3FD8123C943D3F866" authority="Zelnio &amp; Hourdez, 2009" authorityName="Zelnio &amp; Hourdez" authorityYear="2009" box="[556,911,1964,1982]" class="Malacostraca" family="Alvinocarididae" genus="Alvinocaris" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="88" phylum="Arthropoda" rank="species" species="komaii">
<emphasis id="B908EAE4FF8CFFB3FD8123C9429DF866" box="[556,705,1964,1982]" italics="true" pageId="15" pageNumber="88">Alvinocaris komaii</emphasis>
<bibRefCitation id="EFED4B07FF8CFFB3FD6A23C943D3F866" author="ZELNIO K. A. &amp; HOURDEZ S." box="[711,911,1964,1982]" pageId="15" pageNumber="88" pagination="52 - 71" refId="ref32459" refString="ZELNIO K. A. &amp; HOURDEZ S. 2009. - A new species of Alvinocaris (Crustacea: Decapoda: Caridea: Alvinocarididae) from hydrothermal vents at the Lau Basin, southwest Pacific, and a key to the species of Alvinocarididae. Proceedings of the Biological Society of Washington 122 (1): 52 - 71. https: // doi. org / 10.2988 / 07 - 28.1" type="journal article" year="2009">Zelnio &amp; Hourdez, 2009</bibRefCitation>
</taxonomicName>
. This invidual with palm of preserved left cheliped (regenerated?) bearing only a single dark spot is atypical such as many
<taxonomicName id="4C7C4D75FF8CFFB3FDF723A6429BF80D" authorityName="Hessler &amp; Martin" authorityYear="1989" box="[602,711,1987,2005]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="B908EAE4FF8CFFB3FDF723A6429BF80D" box="[602,711,1987,2005]" italics="true" pageId="15" pageNumber="88">Austinograea</emphasis>
</taxonomicName>
species showing regenerated chelipeds; patches of setae discernible just above on inner surface of chela and on right subhepatic region. Photographies courtesy of C. R. Fisher (Penn State University, USA).
</paragraph>
</caption>
<subSubSection id="C366657DFF93FFACFF36255C4260FD8B" pageId="16" pageNumber="89" type="materials_examined">
<paragraph id="8BC336F6FF93FFACFF36255C42F0FE0A" blockId="16.[132,776,217,2030]" pageId="16" pageNumber="89">
<materialsCitation id="3B143CABFF93FFACFF36255C416AFE6B" ID-GBIF-Occurrence="2594759997" collectingDate="2009-05-28" collectionCode="MNHN-IU-" elevation="2130" latitude="-20.760834" location="Lau Back-Arc Basin" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" pageId="16" pageNumber="89" specimenCount="1" specimenCount-female="1" specimenCount-male="2130">
<collectionCode id="ED6DAE33FF93FFACFF36255C417CFE8B" box="[155,288,313,339]" pageId="16" pageNumber="89">MNHN-IU-</collectionCode>
2016-10744,
<specimenCount id="9D7AFD7FFF93FFACFE01255C418CFE8D" box="[428,464,313,341]" pageId="16" pageNumber="89" type="female">1 ♀</specimenCount>
27.1 ×
<quantity id="4C849B13FF93FFACFDB4255C422AFE8B" box="[537,630,313,339]" metricMagnitude="-2" metricUnit="m" metricValue="4.46" pageId="16" pageNumber="89" unit="mm" value="44.6">44.6 mm</quantity>
(left-handed),
<location id="8EA3602DFF93FFACFF29253C4102FEAB" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF93FFACFF29253C4102FEAB" box="[132,350,345,371]" latitude="-20.760834" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" name="Lau Back-Arc Basin" pageId="16" pageNumber="89">Lau Back-Arc Basin</location>
,
<location id="8EA3602DFF93FFACFEC4253C4188FEAB" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF93FFACFEC4253C4188FEAB" box="[361,468,345,371]" latitude="-20.760834" longLatPrecision="12" longitude="-176.19083" municipality="Lau Back-Arc Basin" name="Lau Basin" pageId="16" pageNumber="89">Lau Basin</location>
2009 cruise, dive 427, ABE site,
<geoCoordinate id="EE485031FF93FFACFF1E251D4175FE4B" box="[179,297,376,403]" degrees="20" direction="south" minutes="45.65" orientation="latitude" pageId="16" pageNumber="89" precision="9" value="-20.760834">20°45.65S</geoCoordinate>
,
<geoCoordinate id="EE485031FF93FFACFE9E251D419EFE4B" box="[307,450,376,403]" degrees="176" direction="west" minutes="11.45" orientation="longitude" pageId="16" pageNumber="89" precision="9" value="-176.19083">176°11.45W</geoCoordinate>
,
<quantity id="4C849B13FF93FFACFE66251C4242FE4B" box="[459,542,377,403]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="16" pageNumber="89" unit="m" value="2130.0">
<elevation id="0051D1C5FF93FFACFE66251C4242FE4B" box="[459,542,377,403]" metricMagnitude="3" metricUnit="m" metricValue="2.13" pageId="16" pageNumber="89" unit="m" value="2130.0">2130 m</elevation>
</quantity>
,
<date id="FFC21036FF93FFACFD85251C42CAFE4B" box="[552,662,377,403]" pageId="16" pageNumber="89" value="2009-05-28">
<collectingDate id="EF86E9DEFF93FFACFD85251C42CAFE4B" box="[552,662,377,403]" pageId="16" pageNumber="89" value="2009-05-28">28.V.2009</collectingDate>
</date>
(preserved in ethanol 85%)
</materialsCitation>
. It belongs to the same sample that contains two typical females devoid of spots on chelae palm;
</paragraph>
<paragraph id="8BC336F6FF93FFACFF3625BD4260FD8B" blockId="16.[132,776,217,2030]" pageId="16" pageNumber="89">
<materialsCitation id="3B143CABFF93FFACFF3625BD4260FD8B" ID-GBIF-Occurrence="2594759998" collectingDate="1989-07-13" collectionCode="MNHN-IU-" elevation="2750" latitude="-18.833334" location="Mussel Valley" longLatPrecision="1273" longitude="173.48334" municipality="Mussel Valley" pageId="16" pageNumber="89" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="North Fiji Basin">
<collectionCode id="ED6DAE33FF93FFACFF3625BD417AFE2A" box="[155,294,472,498]" pageId="16" pageNumber="89">MNHN-IU-</collectionCode>
2016-10768 (= MNHN-B27834),
<specimenCount id="9D7AFD7FFF93FFACFD3725BD42E2FE2C" box="[666,702,472,500]" pageId="16" pageNumber="89" type="female">1♀</specimenCount>
22.7 ×
<quantity id="4C849B13FF93FFACFF29259D40B5FDCB" box="[132,233,504,531]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="16" pageNumber="89" unit="mm" value="36.5">36.5 mm</quantity>
,
<collectingRegion id="49B8F814FF93FFACFF5F259D41FDFDCA" box="[242,417,504,530]" pageId="16" pageNumber="89">North Fiji Basin</collectingRegion>
, STARMER II cruise, dive PL 18,
<collectingMunicipality id="6BA7AC8CFF93FFACFF29267D4148FDEA" box="[132,276,536,562]" latitude="-18.833334" longLatPrecision="1273" longitude="173.48334" name="Mussel Valley" pageId="16" pageNumber="89" stateProvince="North Fiji Basin">Mussel Valley</collectingMunicipality>
,
<geoCoordinate id="EE485031FF93FFACFEB0267D4128FDEB" box="[285,372,536,563]" degrees="18" direction="south" minutes="50" orientation="latitude" pageId="16" pageNumber="89" precision="925" value="-18.833334">18°50S</geoCoordinate>
,
<geoCoordinate id="EE485031FF93FFACFED0267D41BFFDEA" box="[381,483,536,562]" degrees="173" direction="east" minutes="29" orientation="longitude" pageId="16" pageNumber="89" precision="925" value="173.48334">173°29E</geoCoordinate>
,
<quantity id="4C849B13FF93FFACFE40267D421EFDEB" box="[493,578,536,563]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="16" pageNumber="89" unit="m" value="2750.0">
<elevation id="0051D1C5FF93FFACFE40267D421EFDEB" box="[493,578,536,563]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="16" pageNumber="89" unit="m" value="2750.0">2750 m</elevation>
</quantity>
,
<date id="FFC21036FF93FFACFDE1267D428FFDEA" box="[588,723,536,562]" pageId="16" pageNumber="89" value="1989-07-13">
<collectingDate id="EF86E9DEFF93FFACFDE1267D428FFDEA" box="[588,723,536,562]" pageId="16" pageNumber="89" value="1989-07-13">13.VII.1989</collectingDate>
</date>
(was found with a typical
<specimenCount id="9D7AFD7FFF93FFACFEC8265D41C4FD8A" box="[357,408,568,594]" pageId="16" pageNumber="89" type="male">male</specimenCount>
of
<taxonomicName id="4C7C4D75FF93FFACFE11265D4270FD8A" authorityName="Guinot" authorityYear="1990" box="[444,556,568,594]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF93FFACFE11265D4270FD8A" box="[444,556,568,594]" italics="true" pageId="16" pageNumber="89">A. alayseae</emphasis>
</taxonomicName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C366657DFF93FFACFF36263D4134F836" pageId="16" pageNumber="89" type="description">
<paragraph id="8BC336F6FF93FFACFF36263D42EEFCA9" blockId="16.[132,776,217,2030]" pageId="16" pageNumber="89">
<materialsCitation id="3B143CABFF93FFACFF36263D4137FD09" ID-GBIF-Occurrence="2594760003" collectingDate="2006-09-07" collectionCode="MNHN-" elevation="2623" latitude="-20.053833" location="Lau Back-Arc Basin" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" pageId="16" pageNumber="89" specimenCode="MGLN07" specimenCount="1" specimenCount-female="1" specimenCount-male="2623">
A large, right-handed
<specimenCount id="9D7AFD7FFF93FFACFE09263E41E4FDAC" box="[420,440,603,628]" pageId="16" pageNumber="89" type="female"></specimenCount>
30.0 ×
<quantity id="4C849B13FF93FFACFDB6263D42D5FDAA" box="[539,649,600,627]" metricMagnitude="-2" metricUnit="m" metricValue="4.85" pageId="16" pageNumber="89" unit="mm" value="48.5">48.5 mm</quantity>
(
<collectionCode id="ED6DAE33FF93FFACFD30263D4354FDAA" box="[669,776,600,626]" pageId="16" pageNumber="89">MNHN-</collectionCode>
IU-2016-10752) from the
<location id="8EA3602DFF93FFACFE35261D423AFD4A" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF93FFACFE35261D423AFD4A" box="[408,614,632,658]" latitude="-20.053833" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" name="Lau Back-Arc Basin" pageId="16" pageNumber="89">Lau Back-Arc Basin</location>
, MGLN07MV cruise, dive 230,
<location id="8EA3602DFF93FFACFE9A26FD41EBFD6A" LSID="urn:lsid:plazi:treatment:03D587E0FF84FFACFE86263D4442FD6A:8EA3602DFF93FFACFE9A26FD41EBFD6A" box="[311,439,664,690]" latitude="-20.053833" longLatPrecision="12" longitude="-176.1335" municipality="Lau Back-Arc Basin" name="Kilo Moana" pageId="16" pageNumber="89">Kilo Moana</location>
site,
<geoCoordinate id="EE485031FF93FFACFE4126FD423BFD6A" box="[492,615,664,690]" degrees="20" direction="south" minutes="03.23" orientation="latitude" pageId="16" pageNumber="89" precision="9" value="-20.053833">20°03.23S</geoCoordinate>
,
<geoCoordinate id="EE485031FF93FFACFDDD26FD435EFD6A" box="[624,770,664,690]" degrees="176" direction="west" minutes="08.01" orientation="longitude" pageId="16" pageNumber="89" precision="9" value="-176.1335">176°08.01W</geoCoordinate>
,
<quantity id="4C849B13FF93FFACFF2926D24080FD0A" box="[132,220,695,722]" metricMagnitude="3" metricUnit="m" metricValue="2.623" pageId="16" pageNumber="89" unit="m" value="2623.0">
<elevation id="0051D1C5FF93FFACFF2926D24080FD0A" box="[132,220,695,722]" metricMagnitude="3" metricUnit="m" metricValue="2.623" pageId="16" pageNumber="89" unit="m" value="2623.0">2623 m</elevation>
</quantity>
,
<date id="FFC21036FF93FFACFF4A26D24134FD09" box="[231,360,695,721]" pageId="16" pageNumber="89" value="2006-09-07">
<collectingDate id="EF86E9DEFF93FFACFF4A26D24134FD09" box="[231,360,695,721]" pageId="16" pageNumber="89" value="2006-09-07">07.IX.2006</collectingDate>
</date>
,
</materialsCitation>
and identified as
<taxonomicName id="4C7C4D75FF93FFACFD8726DD42FFFD09" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[554,675,695,722]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFD8726DD42FFFD09" box="[554,675,695,722]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFD0426DD42B8FD0A" box="[681,740,696,722]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
on the basis of other characters, has the right chela of a crusher
<typeStatus id="54C78854FF93FFACFF29269D40EEFCCA" box="[132,178,760,786]" pageId="16" pageNumber="89">type</typeStatus>
with one marked spot plus a second spot only as a yellow trace, and the cutter with two yellowish spots (
<figureCitation id="13472A73FF93FFACFD2F277242A6FCEA" box="[642,762,791,818]" captionStart="FIG" captionStartId="9.[132,143,1920,1938]" captionTargetBox="[132,1457,216,1756]" captionTargetId="figure@9.[699,984,759,1032]" captionTargetPageId="9" captionText="FIG. 4. — A-C, Austinograea hourdezi n. sp., holotype, ♂ 25.7 × 40.2 mm (right-handed), Lau Back-Arc Basin, TUIM06MV cruise, dive 142, Tow Cam site, 20°19.07S, 176°08.24W, 2719 m, MNHN-IU-2016-10737: A, B, chelae with two coloured spots on outer surface of palm of right crusher (A) and left cutter (B); C, G1 and G2 photographed in situ; D, E, Austinograea hourdezi n. sp., atypical ♀ 30.0 × 48.5 mm (right-handed), Lau Back-Arc Basin, MGLN07MV cruise, dive 230, Kilo Moana site, 20°03.23S, 176°08.01W, 2623 m, MNHN-IU-2016-10752: D, crusher with one marked dark spot and the other one, only discernible by a yellow trace (wound at base of fixed finger is not a spot comparable to that of A. jolliveti n. sp.); E, cutter with two indistinct spots, only as yellow traces." figureDoi="http://doi.org/10.5281/zenodo.3738108" httpUri="https://zenodo.org/record/3738108/files/figure.png" pageId="16" pageNumber="89">Fig. 4D, E</figureCitation>
), This male chela pattern is unusual for female
<taxonomicName id="4C7C4D75FF93FFACFDFC27524298FC89" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[593,708,823,849]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFDFC27524298FC89" box="[593,708,823,849]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFD64275D435EFC8A" box="[713,770,824,850]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
, which are generally devoid of any spot (
<figureCitation id="13472A73FF93FFACFD9D273242F9FCA9" box="[560,677,855,882]" captionStart="FIG" captionStartId="14.[132,143,1964,1982]" captionTargetBox="[132,1455,215,1858]" captionTargetId="figure@14.[699,984,1036,1311]" captionTargetPageId="14" captionText="FIG. 7. — Austinograea hourdezi n. sp., paratype, ♀ 18.3 × 28.4 mm (right-handed), Lau Back-Arc Basin, BIOLAU cruise, BL 12, Vai Lili site, 23°13S, 176°38W, 1750 m, MNHN-IU-2016-10745 (= MNHN-B24060): A, dorsal view; B, C, ventral views, with pleon; D, thoracic sternum and vulvae; E, G, both chelae without spots on outer surface; F, H, both chelae with patches of setae on inner surface; E, F, right crusher; G, H, left cutter." figureDoi="http://doi.org/10.5281/zenodo.3738114" httpUri="https://zenodo.org/record/3738114/files/figure.png" pageId="16" pageNumber="89">Fig. 7E, G</figureCitation>
).
</paragraph>
<paragraph id="8BC336F6FF93FFACFF362712417FFA17" blockId="16.[132,776,217,2030]" pageId="16" pageNumber="89">
A bythograeid that bears only a left cheliped, photographed
<emphasis id="B908EAE4FF93FFACFF4827F24174FC68" box="[229,296,919,944]" italics="true" pageId="16" pageNumber="89">in situ</emphasis>
(
<figureCitation id="13472A73FF93FFACFE9827F241D6FC69" box="[309,394,919,945]" captionStart="FIG" captionStartId="15.[132,143,1851,1869]" captionTargetBox="[426,1176,216,1815]" captionTargetId="figure@15.[426,1176,216,1816]" captionTargetPageId="15" captionText="FIG. 8. — A, B, Austinograea alayseae Guinot, 1990,photographed in situ: see absence of patches of setae on inner surface of chelae:A, crab with white carapace, Lau Basin 2009 cruise, Kilo Moana site, 20°03.23S, 176°08.01W, 2621 m, among mussels Bathymodiolus brevior Cosel, Métivier &amp; Hashimoto, 1994 and snails Ifremeria nautilei Bouchet &amp; Warén, 1991; B, crab with carapace stained orange due to iron or manganese oxide deposits, and cluster of alvinocaridid shrimps, Lau Basin, TUIM07MV, Kilo Moana site, 20°03.23S, 176°08.01W, 2621 m; C, Austinograea hourdezi n. sp. (presumed identification, see p. 89), Lau Basin, Lau Basin 2009 cruise, Kilo Moana site, among mussels Bathymodiolus brevior, gastropods Ifremeria nautilei, sea anemone probably Cyananthea hourdezi Zelnio, Rodríguez &amp; Daly, 2009, and alvinocaridid shrimp, Alvinocaris komaii Zelnio &amp; Hourdez, 2009. This invidual with palm of preserved left cheliped (regenerated?) bearing only a single dark spot is atypical such as many Austinograea species showing regenerated chelipeds; patches of setae discernible just above on inner surface of chela and on right subhepatic region. Photographies courtesy of C. R. Fisher (Penn State University, USA)." figureDoi="http://doi.org/10.5281/zenodo.3738116" httpUri="https://zenodo.org/record/3738116/files/figure.png" pageId="16" pageNumber="89">Fig. 8C</figureCitation>
) at the site Kilo Moana during the Lau Basin 2009 cruise and with a single dark spot on the chela could be
<taxonomicName id="4C7C4D75FF93FFACFE8027B241F0FC28" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[301,428,982,1009]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFE8027B241F0FC28" box="[301,428,982,1009]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFE1E27B241AFFC29" box="[435,499,983,1009]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
, based on the patches of setae discernible just at the suface of the inner chela palm and on setose patches on the right subhepatic region. This could be an atypical specimen of
<taxonomicName id="4C7C4D75FF93FFACFDAD20524223FB88" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[512,639,1078,1105]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFDAD20524223FB88" box="[512,639,1078,1105]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFD2A2052429BFB89" box="[647,711,1079,1105]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
, with a regenerated cheliped. It was found associated with mussels
<taxonomicName id="4C7C4D75FF93FFACFF1C201340E3FB68" authority="Cosel, Metivier &amp; Hashimoto, 1994" authorityName="Cosel, Metivier &amp; Hashimoto" authorityYear="1994" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytiloida" pageId="16" pageNumber="89" phylum="Mollusca" rank="species" species="brevior">
<emphasis id="B908EAE4FF93FFACFF1C201341F9FB48" box="[177,421,1142,1168]" italics="true" pageId="16" pageNumber="89">Bathymodiolus brevior</emphasis>
<bibRefCitation id="EFED4B07FF93FFACFE06201340E3FB68" author="COSEL R. &amp; VON &amp; METIVIER B. &amp; HASHIMOTO J." pageId="16" pageNumber="89" pagination="374 - 392" refId="ref24757" refString="COSEL R. VON, METIVIER B. &amp; HASHIMOTO J. 1994. - Three new species of Bathymodiolus (Bivalvia: Mytilidae) from hydrothermal vents in the Lau basin and the North Fiji Basin, Western Pacific, and the Snake Pit Area, Mid-Atlantic Ridge. Veliger 37: 374 - 392." type="journal article" year="1994">Cosel, Métivier &amp; Hashimoto, 1994</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFED4B07FF93FFACFF5B20F341ECFB68" author="COSEL R. &amp; VON &amp; METIVIER B. &amp; HASHIMOTO J." box="[246,432,1174,1200]" pageId="16" pageNumber="89" pagination="374 - 392" refId="ref24757" refString="COSEL R. VON, METIVIER B. &amp; HASHIMOTO J. 1994. - Three new species of Bathymodiolus (Bivalvia: Mytilidae) from hydrothermal vents in the Lau basin and the North Fiji Basin, Western Pacific, and the Snake Pit Area, Mid-Atlantic Ridge. Veliger 37: 374 - 392." type="journal article" year="1994">
Cosel
<emphasis id="B908EAE4FF93FFACFE9620F2412CFB68" box="[315,368,1174,1200]" italics="true" pageId="16" pageNumber="89">et al.</emphasis>
1994
</bibRefCitation>
), gastropods
<taxonomicName id="4C7C4D75FF93FFACFDEA20F341C3FB08" authority="Bouchet &amp; Waren, 1991" authorityName="Bouchet &amp; Waren" authorityYear="1991" class="Gastropoda" family="Provannidae" genus="Ifremeria" kingdom="Animalia" pageId="16" pageNumber="89" phylum="Mollusca" rank="species" species="nautilei">
<emphasis id="B908EAE4FF93FFACFDEA20F3435BFB68" box="[583,775,1174,1200]" italics="true" pageId="16" pageNumber="89">Ifremeria nautilei</emphasis>
Bouchet &amp; Warén, 1991
</taxonomicName>
, sea anemone
<taxonomicName id="4C7C4D75FF93FFACFDEC20D34267FB28" authority="Zelnio, Rodriguez &amp; Daly, 2009" authorityName="Zelnio, Rodriguez &amp; Daly" authorityYear="2009" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi">
<emphasis id="B908EAE4FF93FFACFDEC20D340EDFB28" italics="true" pageId="16" pageNumber="89">Cyananthea hourdezi</emphasis>
<bibRefCitation id="EFED4B07FF93FFACFF1620B34267FB28" author="ZELNIO K. A. &amp; RODRIGUEZ R. &amp; DALY M." box="[187,571,1238,1264]" pageId="16" pageNumber="89" pagination="547 - 571" refId="ref32533" refString="ZELNIO K. A., RODRIGUEZ R. &amp; DALY M. 2009. - Hexacorals (Anthozoa: Actiniaria, Zoanthidea) from hydrothermal vents in the south-western Pacific. Marine Biology Research 5: 547 - 571. https: // doi. org / 10.1080 / 17451000902729662" type="journal article" year="2009">Zelnio, Rodríguez &amp; Daly, 2009</bibRefCitation>
</taxonomicName>
(see Zelnio
<emphasis id="B908EAE4FF93FFACFD6320B3435BFB28" box="[718,775,1238,1264]" italics="true" pageId="16" pageNumber="89">et al.</emphasis>
2009; E. Rodríguez, pers. comm. 2017). Due to the same features, the seemingly right-handed crab collected at Tow Cam site, dive 240, and photographed
<emphasis id="B908EAE4FF93FFACFD9F21534228FA97" box="[562,628,1334,1359]" italics="true" pageId="16" pageNumber="89">in situ</emphasis>
(Zelnio
<emphasis id="B908EAE4FF93FFACFD792153435BFA97" box="[724,775,1333,1359]" italics="true" pageId="16" pageNumber="89">et al.</emphasis>
2009: fig. 14B) associated with an unidentifiable actiniarian zoanthidean, could be also identified to
<taxonomicName id="4C7C4D75FF93FFACFD282110435BFA57" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[645,775,1397,1423]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFD282110435BFA57" box="[645,775,1397,1423]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFF2921F3409AFA68" box="[132,198,1430,1456]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
, but it is difficult to discern the presence of spots on the chelae.
</paragraph>
<paragraph id="8BC336F6FF93FFACFF3621B04134F836" blockId="16.[132,776,217,2030]" pageId="16" pageNumber="89">
All of these specimens with atypical spot(s) on the chelae exhibit all other morphological traits of
<taxonomicName id="4C7C4D75FF93FFACFD8F219042CAF9D7" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[546,662,1525,1551]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFD8F219042CAF9D7" box="[546,662,1525,1551]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFD3621904289F9D7" box="[667,725,1525,1551]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
, and are not
<taxonomicName id="4C7C4D75FF93FFACFF7A2270410BF9F6" authorityName="in Guinot" authorityYear="1990" box="[215,343,1556,1583]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF93FFACFF7A2270410BF9F6" box="[215,343,1556,1583]" italics="true" pageId="16" pageNumber="89">A. williamsi</emphasis>
</taxonomicName>
nor
<taxonomicName id="4C7C4D75FF93FFACFE2722704266F9F6" authorityName="Tsuchida &amp; Hashimoto" authorityYear="2002" box="[394,570,1556,1583]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="rodriguezensis">
<emphasis id="B908EAE4FF93FFACFE2722704266F9F6" box="[394,570,1556,1583]" italics="true" pageId="16" pageNumber="89">A. rodriguezensis</emphasis>
</taxonomicName>
, both of which are characterised by one spot on the outer chela palm, near the dactylus base. We assume that most of these atypical crabs were initially right-handed and that left-handers represent individuals whose asymmetry has been reversed following autotomy and regeneration of the lost cheliped (see
<emphasis id="B908EAE4FF93FFACFD1422D1425DF936" italics="true" pageId="16" pageNumber="89">Chelae: crusher and cutter. Regenerated chelae</emphasis>
). This reversion of chelipeds, which causes a dramatic rearrangement and triggers a new allometric growth, is thought to have induced a novel implantation of the spots on the chelae, mainly when autotomy or loss occurs at a large size of crabs. These atypical individuals, without other equivocal morphological evidence and presumably not genetically cryptic, are here treated as
<taxonomicName id="4C7C4D75FF93FFACFF2923D640A3F815" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[132,255,1971,1997]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFF2923D640A3F815" box="[132,255,1971,1997]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFEAB23D1411EF816" box="[262,322,1972,1998]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
subject to their recognition by molecular sequences (in study).
</paragraph>
</subSubSection>
<subSubSection id="C366657DFF93FFACFC8024BC4442FD6A" pageId="16" pageNumber="89" type="discussion">
<paragraph id="8BC336F6FF93FFACFC8024BC444AFF29" blockId="16.[813,1456,217,690]" box="[813,1046,217,243]" pageId="16" pageNumber="89">MOLECULAR ANALYSIS</paragraph>
<paragraph id="8BC336F6FF93FFACFC80249F4442FD6A" blockId="16.[813,1456,217,690]" pageId="16" pageNumber="89">
In the early 2000s fragments of
<taxonomicName id="4C7C4D75FF93FFACFB22249C4549FECB" authorityName="Hessler &amp; Martin" authorityYear="1989" box="[1167,1301,249,275]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="B908EAE4FF93FFACFB22249C4549FECB" box="[1167,1301,249,275]" italics="true" pageId="16" pageNumber="89">Austinograea</emphasis>
</taxonomicName>
from the Lau Back-Arc Basin, Valu Fa Ridge, recognised by
<bibRefCitation id="EFED4B07FF93FFACFABC257C45F3FEEC" author="GUINOT D." box="[1297,1455,281,308]" pageId="16" pageNumber="89" pagination="879 - 903" refId="ref26108" refString="GUINOT D. 1990. - Austinograea alayseae sp. nov., crabe hydrothermal decouvert dans le bassin de Lau, Pacifique sud-occidental (Crustacea Decapoda Brachyura). Bulletin du Museum national d'Histoire naturelle 4 e ser., 11, section A (4): 879 - 903." type="journal article" year="1990">Guinot (1990)</bibRefCitation>
as “
<taxonomicName id="4C7C4D75FF93FFACFCFF255C443FFE8B" authorityName="in Guinot" authorityYear="1990" box="[850,1123,313,339]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF93FFACFCFF255C438BFE8B" box="[850,983,313,339]" italics="true" pageId="16" pageNumber="89">Austinograea</emphasis>
aff.
<emphasis id="B908EAE4FF93FFACFBAE255C443FFE8B" box="[1027,1123,313,339]" italics="true" pageId="16" pageNumber="89">williamsi</emphasis>
</taxonomicName>
” and here described as
<taxonomicName id="4C7C4D75FF93FFACFAF4255C4304FEAB" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFAF4255C4304FEAB" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFCF3253C43C6FEAB" box="[862,922,345,371]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
were sent by D. Guinot to L. A. Hurtado (
<collectingRegion id="49B8F814FF93FFACFADF253C45ECFEAB" box="[1394,1456,345,371]" country="United States of America" name="Texas" pageId="16" pageNumber="89">Texas</collectingRegion>
A&amp;M University,
<collectingCountry id="F36B7666FF93FFACFC46251C447CFE4B" box="[1003,1056,377,403]" name="United States of America" pageId="16" pageNumber="89">USA</collectingCountry>
) and V. Leignel (Université du
<collectingRegion id="49B8F814FF93FFACFAC9251C45F0FE4B" box="[1380,1452,377,403]" country="United States of America" name="Maine" pageId="16" pageNumber="89">Maine</collectingRegion>
, Le Mans,
<collectingCountry id="F36B7666FF93FFACFC3825FC4383FE6B" box="[917,991,409,435]" name="France" pageId="16" pageNumber="89">France</collectingCountry>
) for molecular analyses. Genetic divergence warranted recognition of a new species. The sequences obtained for the 16S rDNA, Cytb, and 28S rDNA recovered that the uncorrected nucleotide divergence for the mitochondrial 16S rDNA gene was 5.5% between
<taxonomicName id="4C7C4D75FF93FFACFA9E267D45F3FDEA" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[1331,1455,536,562]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFA9E267D45F3FDEA" box="[1331,1455,536,562]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFC80265C4335FD8B" box="[813,873,569,595]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C7C4D75FF93FFACFC0C265D447EFD8A" authorityName="in Guinot" authorityYear="1990" box="[929,1058,568,594]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF93FFACFC0C265D447EFD8A" box="[929,1058,568,594]" italics="true" pageId="16" pageNumber="89">A. williamsi</emphasis>
</taxonomicName>
; 7% between
<taxonomicName id="4C7C4D75FF93FFACFB6D265D4567FD8A" authority="Guinot &amp; Segonzac, 2018" authorityName="Guinot &amp; Segonzac" authorityYear="2018" box="[1216,1339,568,594]" class="Anthozoa" family="Actinostolidae" genus="Cyananthea" kingdom="Animalia" order="Actiniaria" pageId="16" pageNumber="89" phylum="Cnidaria" rank="species" species="hourdezi" status="sp. nov.">
<emphasis id="B908EAE4FF93FFACFB6D265D4567FD8A" box="[1216,1339,568,594]" italics="true" pageId="16" pageNumber="89">A. hourdezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B579FFF93FFACFAEE265C4523FD8B" box="[1347,1407,569,595]" pageId="16" pageNumber="89" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C7C4D75FF93FFACFC80263D43C3FDAA" authorityName="Guinot" authorityYear="1990" box="[813,927,600,626]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF93FFACFC80263D43C3FDAA" box="[813,927,600,626]" italics="true" pageId="16" pageNumber="89">A. alayseae</emphasis>
</taxonomicName>
; and 5.9% between
<taxonomicName id="4C7C4D75FF93FFACFB29263D4559FDAA" authorityName="in Guinot" authorityYear="1990" box="[1156,1285,600,626]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF93FFACFB29263D4559FDAA" box="[1156,1285,600,626]" italics="true" pageId="16" pageNumber="89">A. williamsi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7C4D75FF93FFACFA90263D45F3FDAA" authorityName="Guinot" authorityYear="1990" box="[1341,1455,600,626]" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="alayseae">
<emphasis id="B908EAE4FF93FFACFA90263D45F3FDAA" box="[1341,1455,600,626]" italics="true" pageId="16" pageNumber="89">A. alayseae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFED4B07FF93FFACFC9B261D447CFD4A" author="MATEOS M. &amp; HURTADO L. A. &amp; SANTAMARIA C. A. &amp; LEIGNEL V. &amp; GUINOT D." box="[822,1056,632,658]" pageId="16" pageNumber="89" refId="ref29119" refString="MATEOS M., HURTADO L. A., SANTAMARIA C. A., LEIGNEL V. &amp; GUINOT D. 2012. - Molecular systematics of the deep-sea hydrothermal vent endemic brachyuran family Bythograeidae Williams, 1980: a comparison of three Bayesian species-tree methods. PloS ONE 7 (3): e 32066, 14 p. https: // doi. org / 10.1371 / journal. pone. 0032066" type="book" year="2012">
Mateos
<emphasis id="B908EAE4FF93FFACFC20261D439EFD49" box="[909,962,632,658]" italics="true" pageId="16" pageNumber="89">et al.</emphasis>
2012: 1
</bibRefCitation>
, 3, 5, 6, 10, fig. 2, table 1, as
<taxonomicName id="4C7C4D75FF93FFACFADD261D4452FD6A" authorityName="in Guinot" authorityYear="1990" class="Malacostraca" family="Bythograeidae" genus="Austinograea" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="89" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B908EAE4FF93FFACFADD261D4321FD69" italics="true" pageId="16" pageNumber="89">Austinograea</emphasis>
aff.
<emphasis id="B908EAE4FF93FFACFC0026FD4452FD6A" box="[941,1038,664,690]" italics="true" pageId="16" pageNumber="89">williamsi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>