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<document id="E18B4CE74D12E26F150BF0A866891D18" ID-DOI="10.1111/j.1096-3642.2009.00527.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443075" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454762105" checkinUser="felipe" docAuthor="García-Paredes, Israel, Peláez-Campomanes, Pablo &amp; Álvarez-Sierra, M. Ángeles" docDate="2009" docId="03D5878B1752AB64FF62602C9A131D80" docLanguage="en" docName="j.1096-3642.2009.00527.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00527.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Pseudodryomys De Bruijn 1966" docType="treatment" docVersion="3" lastPageNumber="645" masterDocId="FFECFFF31745AB73FF9464629E0B1F6A" masterDocTitle="Gliridae (Rodentia, Mammalia) with a simple dental pattern: a new genus and new species from the European Early and Middle Miocene" masterLastPageNumber="652" masterPageNumber="622" pageNumber="645" updateTime="1699276861509" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="D4230D5A32ECC29E8AEEDA933E75BD40">Gliridae (Rodentia, Mammalia) with a simple dental pattern: a new genus and new species from the European Early and Middle Miocene</mods:title>
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<mods:namePart id="ED5C5A7BFEEDF0A506E30032CA37F56C">García-Paredes, Israel</mods:namePart>
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<mods:namePart id="7B9C2D901CB54711F32485D0088731BF">Peláez-Campomanes, Pablo</mods:namePart>
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<mods:title id="8CB2B2E8F8D9F9A60518AF90AF538E92">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="94FE4F2B142B6ED71B8E94B14D03A556">2009</mods:date>
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<subSubSection id="C36665161752AB64FF62602C9C841B0E" box="[246,655,1102,1124]" pageId="23" pageNumber="645" type="nomenclature">
<paragraph id="8BC3369D1752AB64FF62602C9C841B0E" blockId="23.[246,655,1102,1124]" box="[246,655,1102,1124]" pageId="23" pageNumber="645">
<heading id="D08B81F11752AB64FF62602C9C841B0E" box="[246,655,1102,1124]" centered="true" fontSize="9" level="2" pageId="23" pageNumber="645" reason="2">
<taxonomicName id="4C7C4D1E1752AB64FF62602C9C841B0E" authority="DE BRUIJN, 1966" authorityName="De Bruijn" authorityYear="1966" box="[246,655,1102,1124]" class="Mammalia" family="Gliridae" genus="Pseudodryomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="645" phylum="Chordata" rank="genus">
<emphasis id="B908EA8F1752AB64FF62602C9FB01B0B" box="[246,443,1102,1123]" italics="true" pageId="23" pageNumber="645">PSEUDODRYOMYS</emphasis>
<bibRefCitation id="EFED4B6C1752AB64FE52602C9C841B0E" author="De Bruijn H" box="[454,655,1102,1124]" pageId="23" pageNumber="645" pagination="58 - 71" refId="ref17196" refString="De Bruijn H. 1966. Some new Miocene Gliridae (Rodentia, Mammalia) from the Calatayud area (prov. Zaragoza, Spain), I. Proceedings of the Koninklijke Nederlandse Akademie Van Wetenschappen Series B 69: 58 - 71." type="journal article" year="1966">DE BRUIJN, 1966</bibRefCitation>
</taxonomicName>
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<subSubSection id="C36665161752AB64FF0460179A131D80" pageId="23" pageNumber="645" type="materials_examined">
<paragraph id="8BC3369D1752AB64FF0460179EE31BC3" blockId="23.[144,759,1140,1193]" pageId="23" pageNumber="645">
<emphasis id="B908EA8F1752AB64FF0460179F141BE0" box="[144,287,1141,1162]" italics="true" pageId="23" pageNumber="645">
<typeStatus id="54C7883F1752AB64FF0460179ECF1BE0" box="[144,196,1141,1162]" pageId="23" pageNumber="645">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C7C4D1E1752AB64FEB360169CFC1BE0" authority="De Bruijn, 1966" authorityName="De Bruijn" authorityYear="1966" box="[295,759,1140,1162]" class="Mammalia" family="Gliridae" genus="Pseudodryomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="645" phylum="Chordata" rank="species" species="ibericus">
<emphasis id="B908EA8F1752AB64FEB360169C331BE3" box="[295,568,1140,1161]" italics="true" pageId="23" pageNumber="645">Pseudodryomys ibericus</emphasis>
<bibRefCitation id="EFED4B6C1752AB64FDD460179CFC1BE0" author="De Bruijn H" box="[576,759,1140,1162]" pageId="23" pageNumber="645" pagination="58 - 71" refId="ref17196" refString="De Bruijn H. 1966. Some new Miocene Gliridae (Rodentia, Mammalia) from the Calatayud area (prov. Zaragoza, Spain), I. Proceedings of the Koninklijke Nederlandse Akademie Van Wetenschappen Series B 69: 58 - 71." type="journal article" year="1966">De Bruijn, 1966</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13472A181752AB64FF0C60F19ED11BC3" box="[152,218,1171,1193]" captionStart="Figure 9" captionStartId="16.[164,243,1310,1329]" captionTargetBox="[387,1221,195,1281]" captionTargetId="figure-195@16.[387,1221,195,1281]" captionTargetPageId="16" captionText="Figure 9. Pseudodryomys ibericus De Bruijn, 1966 from La Col C (collection MNCN). A, P4 COL C-2178. B, M1 COL C-1986. C, M2 COL C-2021. D, M3 COL C-2040. E, P4 COL C-2205. F, M1 COL C-2096. G, M2 COL C-2064. H, M3 COL C-2123. I, p4 COL C-2181. J, m1 COL C-1989. K, m2 COL C-2024. L, m3 COL C-2042. M, p4 COL C-2206. N, m1 COL C-2068. O, m2 COL C-2108. P, m3 COL C-2138. Right side specimens underlined." figureDoi="http://doi.org/10.5281/zenodo.5443103" httpUri="https://zenodo.org/record/5443103/files/figure.png" pageId="23" pageNumber="645">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BC3369D1752AB64FF0460B19CA41ACB" blockId="23.[144,759,1235,1441]" pageId="23" pageNumber="645">
<emphasis id="B908EA8F1752AB64FF0460B19C631B82" box="[144,616,1235,1257]" italics="true" pageId="23" pageNumber="645">
Original diagnosis (
<bibRefCitation id="EFED4B6C1752AB64FEEF60B19C511B82" author="De Bruijn H" box="[379,602,1235,1257]" pageId="23" pageNumber="645" pagination="58 - 71" refId="ref17196" refString="De Bruijn H. 1966. Some new Miocene Gliridae (Rodentia, Mammalia) from the Calatayud area (prov. Zaragoza, Spain), I. Proceedings of the Koninklijke Nederlandse Akademie Van Wetenschappen Series B 69: 58 - 71." type="journal article" year="1966">De Bruijn, 1966: 7</bibRefCitation>
):
</emphasis>
‘
<taxonomicName id="4C7C4D1E1752AB64FDEC60B19CDD1B83" authorityName="MUIRHEAD" authorityYear="1819" box="[632,726,1235,1257]" class="Mammalia" family="Gliridae" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="645" phylum="Chordata" rank="family">Gliridae</taxonomicName>
of intermediate size. Occlusal surface of cheek teeth concave. Crowns of cheek teeth low. P4/p4 small relative to M1/m1 and rounded; M1/m1 smaller than M2/m2. The Spanish representatives of the genus
<taxonomicName id="4C7C4D1E1752AB64FF04610E9F4B1AEB" authorityName="De Bruijn" authorityYear="1966" box="[144,320,1388,1409]" class="Mammalia" family="Gliridae" genus="Pseudodryomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="645" phylum="Chordata" rank="genus">
<emphasis id="B908EA8F1752AB64FF04610E9F4B1AEB" box="[144,320,1388,1409]" italics="true" pageId="23" pageNumber="645">Pseudodryomys</emphasis>
</taxonomicName>
are characterized by a gradual simplification of the dental pattern through time’.
</paragraph>
<paragraph id="8BC3369D1752AB64FF0461A99DC31E3C" blockId="23.[144,759,1483,1903]" lastBlockId="23.[806,1421,198,342]" pageId="23" pageNumber="645">
<emphasis id="B908EA8F1752AB64FF0461A99F7C1A8A" box="[144,375,1483,1504]" italics="true" pageId="23" pageNumber="645">Emended diagnosis:</emphasis>
Medium to large-sized
<taxonomicName id="4C7C4D1E1752AB64FD0661A99CF91A8B" authorityName="MUIRHEAD" authorityYear="1819" box="[658,754,1483,1505]" class="Mammalia" family="Gliridae" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="645" phylum="Chordata" rank="family">Gliridae</taxonomicName>
, which have cheek teeth with a concave occlusal surface. The dental pattern is formed by the four main ridges and a variable number of extra ridges, usually between two and eight, including centrolophs and centrolophids. The ridges are relatively high, thick, and slanting and the valleys are narrow. In P4, M1, and M2 the trigon is very asymmetric forming a composite Y-shape. The upper molars generally have both centrolophs, the anterior one usually being longer. The lingual parts of the centrolophs may be joined to each other and/or also to the metaloph. The P4 has an oval outline; it is not very simplified and may present one or even two short centrolophs. Lower molars have a relatively long centrolophid and in m1 and m2 there is at least one extra ridge in the posterior valley. The mesolophid of the m3 is not reduced and labially is not connected to the posterolophid.
</paragraph>
<paragraph id="8BC3369D1752AB64FCB265E29B241EDE" blockId="23.[806,1421,384,436]" pageId="23" pageNumber="645">
<emphasis id="B908EA8F1752AB64FCB265E29A661EFF" box="[806,1133,384,405]" italics="true" pageId="23" pageNumber="645">Stratigraphical distribution:</emphasis>
Local zones X to E (
<collectionCode id="ED6DAE581752AB64FAF565E29B861EFF" box="[1377,1421,384,405]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="23" pageNumber="645">MN</collectionCode>
1, Lower Miocene –
<collectionCode id="ED6DAE581752AB64FB8165FD9A4B1EDE" box="[1045,1088,415,436]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="23" pageNumber="645">MN</collectionCode>
5, Middle Miocene).
</paragraph>
<paragraph id="8BC3369D1752AB64FCB265BC9D8B1D78" blockId="23.[806,1421,478,530]" pageId="23" pageNumber="645">
<emphasis id="B908EA8F1752AB64FCB265BC9A591E99" box="[806,1106,478,499]" italics="true" pageId="23" pageNumber="645">Geographical distribution:</emphasis>
Central and south-western Europe.
</paragraph>
<paragraph id="8BC3369D1752AB64FCB2665E9B861DE5" blockId="23.[806,1421,572,747]" pageId="23" pageNumber="645">
<emphasis id="B908EA8F1752AB64FCB2665E9DEF1D3B" box="[806,996,572,593]" italics="true" pageId="23" pageNumber="645">Species included</emphasis>
<taxonomicName id="4C7C4D1E1752AB64FCB266399AF51D1B" authority="De Bruijn, 1966" authorityName="De Bruijn" authorityYear="1966" box="[806,1278,603,625]" class="Mammalia" family="Gliridae" genus="Pseudodryomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="645" phylum="Chordata" rank="species" species="ibericus">
<emphasis id="B908EA8F1752AB64FCB266399A321D1A" box="[806,1081,603,624]" italics="true" pageId="23" pageNumber="645">Pseudodryomys ibericus</emphasis>
<bibRefCitation id="EFED4B6C1752AB64FBD766399AF51D1B" author="De Bruijn H" box="[1091,1278,603,625]" pageId="23" pageNumber="645" pagination="58 - 71" refId="ref17196" refString="De Bruijn H. 1966. Some new Miocene Gliridae (Rodentia, Mammalia) from the Calatayud area (prov. Zaragoza, Spain), I. Proceedings of the Koninklijke Nederlandse Akademie Van Wetenschappen Series B 69: 58 - 71." type="journal article" year="1966">De Bruijn, 1966</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C7C4D1E1752AB64FCB2661B9A081DE5" baseAuthorityName="Garcia Moreno" baseAuthorityYear="1986" box="[806,1027,633,655]" class="Mammalia" family="Gliridae" genus="Pseudodryomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="645" phylum="Chordata" rank="species" species="rex">
<emphasis id="B908EA8F1752AB64FCB2661B9A081DE5" box="[806,1027,633,655]" italics="true" pageId="23" pageNumber="645">Pseudodryomys rex</emphasis>
</taxonomicName>
(García Moreno,
<quantity id="4C849B781752AB64FB47661B9B201DE5" box="[1235,1323,633,655]" metricMagnitude="1" metricUnit="m" metricValue="5.04444" pageId="23" pageNumber="645" unit="in" value="1986.0">1986 in</quantity>
<bibRefCitation id="EFED4B6C1752AB64FAA2661A9B861DE5" author="Alvarez Sierra MA &amp; Garcia-Moreno E" box="[1334,1421,632,655]" pageId="23" pageNumber="645" pagination="145 - 189" refId="ref15557" refString="Alvarez Sierra MA, Garcia-Moreno E. 1986. New Gliridae and Cricetidae from the Middle and Upper Miocene of the Duero basin, Spain. Studia Geologica Salmanticensia 22: 145 - 189." type="journal article" year="1986">Álvarez</bibRefCitation>
</paragraph>
<paragraph id="8BC3369D1752AB64FCAB66FA9B861DA6" blockId="23.[806,1421,572,747]" pageId="23" pageNumber="645">
Sierra &amp; García-Moreno, 1986)
<taxonomicName id="4C7C4D1E1752AB64FCB266D49B0C1DA6" authority="Agusti, 1993" authorityName="Agusti" authorityYear="1993" box="[806,1287,694,716]" class="Mammalia" family="Gliridae" genus="Pseudodryomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="645" phylum="Chordata" rank="species" species="granatensis">
<emphasis id="B908EA8F1752AB64FCB266D49A6D1DA6" box="[806,1126,694,716]" italics="true" pageId="23" pageNumber="645">Pseudodryomys granatensis</emphasis>
Agustí,
<quantity id="4C849B781752AB64FB4466D59B0C1DA6" box="[1232,1287,695,716]" metricMagnitude="1" metricUnit="m" metricValue="5.06222" pageId="23" pageNumber="645" unit="in" value="1993.0">1993</quantity>
</taxonomicName>
in Martín-
</paragraph>
<paragraph id="8BC3369D1752AB64FCAB66B79A131D80" blockId="23.[806,1421,572,747]" box="[831,1048,725,747]" pageId="23" pageNumber="645">
Suárez
<emphasis id="B908EA8F1752AB64FC0D66B49DDA1D80" box="[921,977,725,747]" italics="true" pageId="23" pageNumber="645">et al.</emphasis>
, 1993
</paragraph>
</subSubSection>
</treatment>
</document>
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