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<document id="C806362CC8E3F95AC8D1668D6C3E6175" ID-DOI="10.5281/zenodo.174076" ID-GBIF-Dataset="50e26ef8-c2f5-47ff-a6bd-b6b34323800b" ID-ISSN="1175­5326" ID-Zenodo-Dep="174076" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459789464809" checkinUser="plazi" docAuthor="Schwendinger, Peter J." docDate="2006" docId="03A5879D753CFF90FE985F7AD8492A03" docLanguage="en" docName="zt01325p266.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Martensiellus" docType="treatment" docVersion="9" lastPageNumber="258" masterDocId="FF9CFFE5753FFF93FF905E66DD6C2F5D" masterDocTitle="A taxonomic revision of the family Oncopodidae VI. Martensiellus, a new genus from Borneo, and the discovery of a tarsal pore organ in Oncopodidae (Opiliones: Laniatores)" masterLastPageNumber="266" masterPageNumber="255" pageNumber="258" updateTime="1698219045570" updateUser="plazi">
<mods:mods id="BDA5A5F9278DAF12BF82F0A533FD35EC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6082B595B7EC97E1792D628421E5DFB0">A taxonomic revision of the family Oncopodidae VI. Martensiellus, a new genus from Borneo, and the discovery of a tarsal pore organ in Oncopodidae (Opiliones: Laniatores)</mods:title>
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<mods:namePart id="47769D71F89A00615DB95D24A7B7A1EC">Schwendinger, Peter J.</mods:namePart>
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<mods:typeOfResource id="8B650020C001DEEB249681EE23BA5939">text</mods:typeOfResource>
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<mods:date id="38BA18E578153B9836B0AE83EF66C2BA">2006</mods:date>
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<mods:classification id="3C59B3B8719A7A68812A3FBD2BCBE1CD">journal article</mods:classification>
<mods:identifier id="5A30C4620DA1437A83825DC6584DF13C" type="DOI">10.5281/zenodo.174076</mods:identifier>
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<treatment id="03A5879D753CFF90FE985F7AD8492A03" ID-DOI="http://doi.org/10.5281/zenodo.6260940" ID-GBIF-Taxon="119349888" ID-Zenodo-Dep="6260940" LSID="urn:lsid:plazi:treatment:03A5879D753CFF90FE985F7AD8492A03" httpUri="http://treatment.plazi.org/id/03A5879D753CFF90FE985F7AD8492A03" lastPageNumber="258" pageId="3" pageNumber="258">
<subSubSection id="C3166500753CFF90FE985F7ADC872E68" box="[264,491,283,310]" pageId="3" pageNumber="258" type="nomenclature">
<paragraph id="8BB3368B753CFF90FE985F7ADC872E68" blockId="3.[264,491,283,310]" box="[264,491,283,310]" pageId="3" pageNumber="258">
<heading id="D0FB81E7753CFF90FE985F7ADC872E68" bold="true" box="[264,491,283,310]" fontSize="11" level="1" pageId="3" pageNumber="258" reason="1">
<emphasis id="B978EA99753CFF90FE985F7ADC872E68" bold="true" box="[264,491,283,310]" pageId="3" pageNumber="258">
<taxonomicName id="4C0C4D08753CFF90FE985F7ADCC22E6B" ID-CoL="7PNJ7" box="[264,430,284,310]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FE985F7ADCC22E6B" bold="true" box="[264,430,284,310]" italics="true" pageId="3" pageNumber="258">Martensiellus</emphasis>
</taxonomicName>
n. g.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3166500753CFF90FE985F3DDE412E28" box="[264,813,347,374]" pageId="3" pageNumber="258" type="materials_examined">
<paragraph id="8BB3368B753CFF90FE985F3DDE412E28" blockId="3.[264,1324,347,1374]" box="[264,813,347,374]" pageId="3" pageNumber="258">
<emphasis id="B978EA99753CFF90FE985F3DDCCC2E28" bold="true" box="[264,416,347,373]" pageId="3" pageNumber="258">
<typeStatus id="54B78829753CFF90FE985F3DDC2F2E28" box="[264,323,347,373]" pageId="3" pageNumber="258">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C0C4D08753CFF90FE225F3ADF8F2E28" box="[434,739,348,373]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="tenuipalpus" status="sp. nov.">
<emphasis id="B978EA99753CFF90FE225F3ADF8F2E28" box="[434,739,348,373]" italics="true" pageId="3" pageNumber="258">Martensiellus tenuipalpus</emphasis>
</taxonomicName>
<emphasis id="B978EA99753CFF90FD7A5F3DDE412E28" bold="true" box="[746,813,347,373]" pageId="3" pageNumber="258">
<taxonomicNameLabel id="A24B57E2753CFF90FD7A5F3DDE412E28" box="[746,813,347,373]" pageId="3" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3166500753CFF90FEA85FE5D8702EC3" box="[312,1308,387,414]" pageId="3" pageNumber="258" type="etymology">
<paragraph id="8BB3368B753CFF90FEA85FE5D8702EC3" blockId="3.[264,1324,347,1374]" box="[312,1308,387,414]" pageId="3" pageNumber="258">
<emphasis id="B978EA99753CFF90FEA85FE5DCD12EC0" bold="true" box="[312,445,387,413]" pageId="3" pageNumber="258">Etymology</emphasis>
. The generic name (male in gender) is derived from the name “Martens”.
</paragraph>
</subSubSection>
<subSubSection id="C3166500753CFF90FEA85FCDD8492A03" pageId="3" pageNumber="258" type="diagnosis">
<paragraph id="8BB3368B753CFF90FEA85FCDDEE02BCB" blockId="3.[264,1324,347,1374]" pageId="3" pageNumber="258">
<emphasis id="B978EA99753CFF90FEA85FCDDCDC2E98" bold="true" box="[312,432,427,453]" pageId="3" pageNumber="258">Diagnosis</emphasis>
. The new genus is characterized by the following combination of characters: 1. The penis carries an expandable, distad­directed glans in which the median plate lies dorsally of the lateral sclerites (
<figureCitation id="13372A0E753CFF90FD7B5F9ADE1F2D4B" box="[747,883,508,534]" captionStart="FIGURES 11 – 17" captionStartId="6.[264,383,1444,1468]" captionTargetBox="[278,1309,534,1410]" captionTargetId="figure@6.[264,1323,518,1422]" captionTargetPageId="6" captionText="FIGURES 11 – 17. Martensiellus tenuipalpus n. g., n. sp., penis (11 – 14 holotype, 15 – 17 paratype). 11 – 12. Total penis (11 dorsal view, 12 lateral view). 13 – 15, 17. Apex of penis in resting position (13, 15 dorsal view, 14, 17 lateral view). 16. Apex of penis with expanded glans, dorsal view. Abbreviations: d. m. distal margin of truncus penis, l. s. lateral sclerite of glans, m. p. median plate of glans, m. t. membranous tubes of glans, s stylus. Scale bars: 0.5 mm (11 – 12), 0.1 mm (13 – 17)." httpUri="https://zenodo.org/record/174079/files/figure.png" pageId="3" pageNumber="258">Figs 11–17</figureCitation>
). The same is found in
<taxonomicName id="4C0C4D08753CFF90FB195F9ADF552D63" authority="Martens &amp; Schwendinger" authorityName="Martens &amp; Schwendinger" class="Arachnida" family="Sandokanidae" genus="Biantoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FB195F9AD8462D48" box="[1161,1322,508,533]" italics="true" pageId="3" pageNumber="258">Biantoncopus</emphasis>
Martens &amp; Schwendinger
</taxonomicName>
, whereas in the distad­directed glans of
<taxonomicName id="4C0C4D08753CFF90FB8C5C42DCA22D3B" authority="Martens &amp; Schwendinger" authorityName="Martens &amp; Schwendinger" class="Arachnida" family="Sandokanidae" genus="Palaeoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FB8C5C42D9AE2D60" box="[1052,1218,548,573]" italics="true" pageId="3" pageNumber="258">Palaeoncopus</emphasis>
Martens &amp; Schwendinger
</taxonomicName>
the plate lies ventral to the sclerites. 2. A dorsal pore is present on tarsi of legs I and II, as is also the case in
<taxonomicName id="4C0C4D08753CFF90FD275C12DE312DD0" box="[695,861,628,653]" class="Arachnida" family="Sandokanidae" genus="Palaeoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FD275C12DE312DD0" box="[695,861,628,653]" italics="true" pageId="3" pageNumber="258">Palaeoncopus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0C4D08753CFF90FC0C5C12D9572DD0" box="[924,1083,628,653]" class="Arachnida" family="Sandokanidae" genus="Caenoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FC0C5C12D9572DD0" box="[924,1083,628,653]" italics="true" pageId="3" pageNumber="258">Caenoncopus</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C0C4D08753CFF90FBE05C12D9B42DD0" box="[1136,1240,628,653]" class="Insecta" family="Geometridae" genus="Oncopus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FBE05C12D9B42DD0" box="[1136,1240,628,653]" italics="true" pageId="3" pageNumber="258">Oncopus</emphasis>
</taxonomicName>
dorsal pores are present on all leg tarsi. In
<taxonomicName id="4C0C4D08753CFF90FD285CFADEE32DEB" authority="Thorell" authorityName="Thorell" box="[696,911,668,694]" class="Arachnida" family="Sandokanidae" genus="Gnomulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FD285CFADE5D2DE8" box="[696,817,668,693]" italics="true" pageId="3" pageNumber="258">Gnomulus</emphasis>
Thorell
</taxonomicName>
and
<taxonomicName id="4C0C4D08753CFF90FC5C5CFAD9012DE8" box="[972,1133,668,693]" class="Arachnida" family="Sandokanidae" genus="Biantoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FC5C5CFAD9012DE8" box="[972,1133,668,693]" italics="true" pageId="3" pageNumber="258">Biantoncopus</emphasis>
</taxonomicName>
tarsal pores are absent. 3. The tarsal formula is 1­1­2­2 (as in
<taxonomicName id="4C0C4D08753CFF90FCE35CA2D9CD2D80" box="[883,1185,708,733]" class="Arachnida" family="Sandokanidae" genus="Caenoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="cuspidatus">
<emphasis id="B978EA99753CFF90FCE35CA2D9CD2D80" box="[883,1185,708,733]" italics="true" pageId="3" pageNumber="258">Caenoncopus cuspidatus</emphasis>
</taxonomicName>
), all tarsi/ tarsomeres are large. In lateral view the undivided tarsi of legs I and II in
<taxonomicName id="4C0C4D08753CFF90FBFA5C8AD8672C58" box="[1130,1291,748,773]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FBFA5C8AD8672C58" box="[1130,1291,748,773]" italics="true" pageId="3" pageNumber="258">Martensiellus</emphasis>
</taxonomicName>
<emphasis id="B978EA99753CFF90FA835C8ADC722C73" bold="true" pageId="3" pageNumber="258">n. g.</emphasis>
are ovoid, clearly longer than deep (
<figureCitation id="13372A0E753CFF90FD495D72DE292C73" box="[729,837,788,814]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="3" pageNumber="258">Figs 6–7</figureCitation>
), whereas in
<taxonomicName id="4C0C4D08753CFF90FC755D72D9E92C70" box="[997,1157,788,813]" class="Arachnida" family="Sandokanidae" genus="Caenoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FC755D72D9E92C70" box="[997,1157,788,813]" italics="true" pageId="3" pageNumber="258">Caenoncopus</emphasis>
</taxonomicName>
they both are semiglobular and clearly deeper than long (see
<bibRefCitation id="EF9D4B7A753CFF90FCA15D5AD9C82C0B" author="Martens" box="[817,1188,828,854]" pageId="3" pageNumber="266" refString="Martens, J. &amp; Schwendinger, P. J. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de Zoologie, 105 (3), 499 - 555." type="journal article" year="1998">Martens &amp; Schwendinger 1998</bibRefCitation>
: fig. 4c). In
<taxonomicName id="4C0C4D08753CFF90FE985D02DC1C2C20" box="[264,368,868,893]" class="Insecta" family="Geometridae" genus="Oncopus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FE985D02DC1C2C20" box="[264,368,868,893]" italics="true" pageId="3" pageNumber="258">Oncopus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9D4B7A753CFF90FE3F5D02DF202C23" author="Thorell" box="[431,588,868,894]" pageId="3" pageNumber="266" refString="Thorell, T. (1891) Opilioni nuovi o poco cognosciuti dell'Archipelago Malese. Annali del Museo Civico di Storia Naturale di Genova, ser. 2, 10, 669 - 770, plates VIII, IX." type="journal article" year="1891">Thorell 1891</bibRefCitation>
: figs 34–35) and, less distinctly so, in
<taxonomicName id="4C0C4D08753CFF90FB9D5D02D9D82C20" box="[1037,1204,868,893]" class="Arachnida" family="Sandokanidae" genus="Palaeoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FB9D5D02D9D82C20" box="[1037,1204,868,893]" italics="true" pageId="3" pageNumber="258">Palaeoncopus</emphasis>
</taxonomicName>
the tarsus of leg I is slightly longer than deep and the tarsus of leg II slightly deeper than long. Tarsomeres of legs III and IV in
<taxonomicName id="4C0C4D08753CFF90FD195DD2DE462C90" box="[649,810,948,973]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FD195DD2DE462C90" box="[649,810,948,973]" italics="true" pageId="3" pageNumber="258">Martensiellus</emphasis>
</taxonomicName>
<emphasis id="B978EA99753CFF90FCA15DD2DE0B2C93" bold="true" box="[817,871,948,974]" pageId="3" pageNumber="258">n. g.</emphasis>
are fairly long and large, only slightly less deep than the corresponding metatarsi in their proximal portion (
<figureCitation id="13372A0E753CFF90FBC25DBAD9AD2CAB" box="[1106,1217,988,1014]" captionStart="FIGURES 4 – 10" captionStartId="5.[264,383,1011,1035]" captionTargetBox="[275,1314,521,997]" captionTargetId="figure@5.[264,1323,503,1005]" captionTargetPageId="5" captionText="FIGURES 4 – 10. Martensiellus tenuipalpus n. g., n. sp., male (4 – 9 holotype, 10 paratype). 4. Chelicera, lateral view. 5. Palpus, lateral view. 6. Distal parts of legs, lateral view (6 leg I, 7 leg II, 8 leg III, 9 leg IV). 10. Palpus and anterior part of body, lateral view. Scale bars: 1 mm (4 – 9 same scale)." httpUri="https://zenodo.org/record/174078/files/figure.png" pageId="3" pageNumber="258">Figs 8–9</figureCitation>
). This is also the case in
<taxonomicName id="4C0C4D08753CFF90FE5E5A62DF2A2B40" box="[462,582,1028,1053]" class="Arachnida" family="Sandokanidae" genus="Gnomulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FE5E5A62DF2A2B40" box="[462,582,1028,1053]" italics="true" pageId="3" pageNumber="258">Gnomulus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9D4B7A753CFF90FD1A5A62DE412B43" author="Silhavy" box="[650,813,1028,1054]" pageId="3" pageNumber="266" refString="Silhavy, V. (1962) Un opilion nouveau, Pelitnus drescoi n. sp. (Fam. Oncopodidae). Bulletin du Museum National d'Histoire Naturelle, ser. 2, 34 (6), 464 - 466." type="journal article" year="1962">Silhavy 1962</bibRefCitation>
: fig. 4) and
<taxonomicName id="4C0C4D08753CFF90FC545A62D9092B40" box="[964,1125,1028,1053]" class="Arachnida" family="Sandokanidae" genus="Biantoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FC545A62D9092B40" box="[964,1125,1028,1053]" italics="true" pageId="3" pageNumber="258">Biantoncopus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9D4B7A753CFF90FB395A62DC972B1B" author="Martens" pageId="3" pageNumber="266" refString="Martens, J. &amp; Schwendinger, P. J. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de Zoologie, 105 (3), 499 - 555." type="journal article" year="1998">Martens &amp; Schwendinger 1998</bibRefCitation>
: fig. 69), whereas in
<taxonomicName id="4C0C4D08753CFF90FC9B5A4ADEC72B18" box="[779,939,1068,1093]" class="Arachnida" family="Sandokanidae" genus="Caenoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FC9B5A4ADEC72B18" box="[779,939,1068,1093]" italics="true" pageId="3" pageNumber="258">Caenoncopus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF9D4B7A753CFF90FC635A4ADC2B2B33" author="Martens" pageId="3" pageNumber="266" refString="Martens, J. &amp; Schwendinger, P. J. (1998) A taxonomic revision of the family Oncopodidae I. New genera and new species of Gnomulus Thorell (Opiliones, Laniatores). Revue suisse de Zoologie, 105 (3), 499 - 555." type="journal article" year="1998">Martens &amp; Schwendinger 1998</bibRefCitation>
: figs 4b, d) and
<taxonomicName id="4C0C4D08753CFF90FD985A32DFC22B30" box="[520,686,1108,1133]" class="Arachnida" family="Sandokanidae" genus="Palaeoncopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FD985A32DFC22B30" box="[520,686,1108,1133]" italics="true" pageId="3" pageNumber="258">Palaeoncopus</emphasis>
</taxonomicName>
the tarsomeres of legs III and IV are short and small, only about half as deep as the corresponding metatarsi.
</paragraph>
<paragraph id="8BB3368B753CFF90FEA85AC2D8492A03" blockId="3.[264,1324,347,1374]" pageId="3" pageNumber="258">
<emphasis id="B978EA99753CFF90FEA85AC2DFB32BE3" bold="true" box="[312,735,1188,1214]" pageId="3" pageNumber="258">Species account and distribution</emphasis>
. At the moment only one species from central
<collectingCountry id="F31B761B753CFF90FE985AAADC012BBB" box="[264,365,1228,1254]" name="Malaysia" pageId="3" pageNumber="258">Sarawak</collectingCountry>
can be unequivocally attributed to the new genus. Females of two different species from eastern
<collectingCountry id="F31B761B753CFF90FD945A92DF052A53" box="[516,617,1268,1294]" name="Malaysia" pageId="3" pageNumber="258">Sarawak</collectingCountry>
and southern Kalimantan (see paragraph “
<taxonomicName id="4C0C4D08753CFF90FBF95A92D8662A50" box="[1129,1290,1268,1293]" class="Arachnida" family="Sandokanidae" genus="Martensiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B978EA99753CFF90FBF95A92D8662A50" box="[1129,1290,1268,1293]" italics="true" pageId="3" pageNumber="258">Martensiellus</emphasis>
</taxonomicName>
<emphasis id="B978EA99753CFF90FA835A92DC352A6B" bold="true" pageId="3" pageNumber="258">n. g. spp.</emphasis>
”) presumably also belong there, but generic placement without males is uncertain. Several more species of this new genus are expected to occur in different parts of Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>
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