<documentid="3ECCF4F3158970FF66AAFAD987A643B2"ID-CLB-Dataset="299670"ID-DOI="10.5281/zenodo.11512126"ID-GBIF-Dataset="522b8de7-880d-40ca-803e-aa5a3a05c5f6"ID-ISSN="0068-547X"ID-Zenodo-Dep="11512126"IM.bibliography_approvedBy="carolina"IM.illustrations_approvedBy="carolina"IM.materialsCitations_approvedBy="carolina"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="carolina"IM.taxonomicNames_approvedBy="carolina"IM.treatmentCitations_approvedBy="carolina"IM.treatments_approvedBy="carolina"checkinTime="1717719049762"checkinUser="felipe"docAuthor="Iwamoto, Tomio, Nakayama, Naohide, Shao, Kwang-Tsao & Table, Hsuan-Ching Ho"docDate="2015"docId="039F87BDFFED1166AB64E67BFCF3FC39"docLanguage="en"docName="ProcCalifAcadSci.62.3.31-126.pdf"docOrigin="Proceedings of the California Academy of Sciences 62 (3)"docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article"docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B"docStyleName="ProcCalifAcadSci.2015-.journal_article"docStyleVersion="2"docTitle="Malacocephalus nipponensis Gilbert and Hubbs 1916"docType="treatment"docVersion="8"lastPageNumber="87"masterDocId="FFA6FFC5FFDA115EAB1FE201FF8EFFEA"masterDocTitle="Synopsis of the Grenadier Fishes (Gadiformes; Teleostei) of Taiwan"masterLastPageNumber="126"masterPageNumber="31"pageNumber="86"updateTime="1720664413385"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0">
<mods:affiliationid="1B88C680467D4F9FB3245020CC970A2B">Section of Ichthyology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA;</mods:affiliation>
<mods:affiliationid="404DFD09E97954679B4FBCE3C24A8B31">Research Center for Biodiversity, Academia Sinica No. 128, Sec. 2, Academia Road, Nankang, Taipei 115, Taiwan;</mods:affiliation>
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<bibRefCitationid="EFA74B5AFFED1169AF47E6E1FEDAFAF9"author="MASUDA, H. & K. AMAOKA & C. ARAGA & T. UYENO & T. YOSHINO"pageId="55"pageNumber="86"refId="ref80275"refString="MASUDA, H., K. AMAOKA, C. ARAGA, T. UYENO, AND T. YOSHINO. 1984. The fishes of the Japanese Archipelago. Tokai University Press, Tokyo, Japan. 437 pp., 370 pls."type="book"year="1984">Masuda et al., 1984:94</bibRefCitation>
<bibRefCitationid="EFA74B5AFFED1169A9A1E6FCFC38FAF9"author="OKAMURA O & AND T. & KITAJIMA, EDS"box="[702,950,1277,1299]"pageId="55"pageNumber="86"refId="ref80944"refString="OKAMURA O., AND T. KITAJIMA, EDS. 1984. Fishes of the Okinawa Trough and the Adjacent Waters. I. The Intensive Research of Unexploited Fishery Resources on Continental Slopes. Japan Fisheries Resource Conservation Association, Tokyo, Japan. 414 pp."type="book"year="1984">Okamura et al., 1984:144</bibRefCitation>
<bibRefCitationid="EFA74B5AFFED1169A8DDE71DFB1BFADB"author="SHEN, S. - C. & S. - C. LEE & K. - T. SHAO & H. - C. MOK & C. - H. CHEN & C. - C. CHEN & C. - S. TZENG & EDS"box="[962,1173,1308,1330]"pageId="55"pageNumber="86"refId="ref81759"refString="SHEN, S. - C., S. - C. LEE, K. - T. SHAO, H. - C. MOK, C. - H. CHEN, C. - C. CHEN, C. - S. TZENG, EDS. 1993. Fishes of Taiwan. National Taiwan University, Taipei, Taiwan. 960 pp. (In Chinese.)"type="book"year="1993">Shen et al., 1993:172</bibRefCitation>
(descr.).—
<bibRefCitationid="EFA74B5AFFED1169AA09E73BFE61FAA5"author="CHIOU, M. & SHAO, AND & T. IWAMOTO"box="[278,495,1338,1360]"pageId="55"pageNumber="86"pagination="35 - 50"refId="ref77534"refString="CHIOU, M. - L, K. - T. SHAO, AND T. IWAMOTO. 2004 b. A new species, Caelorinchus sheni, and 19 new records of grenadiers (Pisces: Gadiformes: Macrouridae) from Taiwan. Zoological Studies 43: 35 - 50."type="journal article"year="2004">Chiou et al., 2004b:37</bibRefCitation>
, 47 (in key, list).—
<bibRefCitationid="EFA74B5AFFED1169A9AAE73BFCD6FAA5"author="SHAO, K. - T. & T. IWAMOTO & H. - C. HO & T. - Y. CHEN & C. - Y. CHEN"box="[693,856,1338,1360]"pageId="55"pageNumber="86"refId="ref81544"refString="SHAO, K. - T., T. IWAMOTO, H. - C. HO, T. - Y. CHEN, AND C. - Y. CHEN. 2008. Species composition and distribution pattern of grenadiers (family Bathygadidae, Macrouridae and Macrouroididae) from Taiwan. Pages 17 - 29 in A. Orlov and T. Iwamoto, eds., Grenadiers of the World Oceans: Biology, Stock Assessment, and Fisheries. American Fisheries Society, Symposium 63, Bethesda, Maryland, USA. 484 pp."type="book"year="2008">Shao et al., 2008</bibRefCitation>
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,
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:
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9 (8–10); inner GR-I 2–4+7–9; pyl. caeca 107–130. Snout 23–28%; orbit 29–37%; suborbital 11–14%; interorbital 26–31%; orbit-preopercle 42–50%; upper jaw 45–56%; barbel 18–22%. Snout bluntly pointed; mouth large, upper jaw extends to below hind margin of orbit; orbit large, greater than snout length; suborbital region vertical and smoothly curved; gill openings wide, extend forward to under posterior margin of orbit. Scales small, beset with fine, slender spinules; scales uniformly and smoothly cover head and body; no coarsely modified scales on ridges of head or tip of snout, lower branchiostegal rays scaled, but gular membrane naked. Light organ with anterior dermal window relatively small and round, situated between or slightly anterior to
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bases; periproct enclosing posterior dermal window and urogenital openings located between
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fins, far removed from A origin. Coloration gray to swarthy with silvery sides, blackish ventrally on head and over abdomen; fins dark dusky to black. Attains>
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<bibRefCitationid="EFA74B5AFFE21166AF08E0BAFF5FFD19"author="OKAMURA O & AND T. & KITAJIMA, EDS"pageId="56"pageNumber="87"refId="ref80944"refString="OKAMURA O., AND T. KITAJIMA, EDS. 1984. Fishes of the Okinawa Trough and the Adjacent Waters. I. The Intensive Research of Unexploited Fishery Resources on Continental Slopes. Japan Fisheries Resource Conservation Association, Tokyo, Japan. 414 pp."type="book"year="1984">Okamura et al. 1984</bibRefCitation>
) considered it a valid species based on the absence of scales on the gular membrane (usually present in
<bibRefCitationid="EFA74B5AFFE21166AB64E11AFED7FCD9"author="IWAMOTO, T."box="[123,345,795,819]"pageId="56"pageNumber="87"pagination="135 - 179"refId="ref78785"refString="IWAMOTO, T. 1979. Eastern Pacific macrourine grenadiers with seven branchiostegal rays (Pisces: Macrouridae). Proceedings of the California Academy of Sciences, ser. 4, 42 (5): 135 - 179."type="journal article"year="1979">Iwamoto (1979:149)</bibRefCitation>
<taxonomicNameid="4C364D28FFE21166A969E11DFC95FCD9"authorityName="Gilbert and Hubbs"authorityYear="1916"box="[630,795,796,819]"family="Macrouridae"genus="Malacocephalus"kingdom="Animalia"order="Gadiformes"pageId="56"pageNumber="87"phylum="Chordata"rank="species"species="nipponensis">M. nipponensis</taxonomicName>
<bibRefCitationid="EFA74B5AFFE21166AF1AE11AFB1BFCD9"author="GILBERT, C. H."box="[1029,1173,795,819]"pageId="56"pageNumber="87"refId="ref77969"refString="GILBERT, C. H. 1905. The deep-sea fishes of the Hawaiian Islands. Pages 575 - 713, figs. 230 - 276, pls. 66 - 101 in D. S. Jordan and B. W. Evermann, eds., The Aquatic Resources of the Hawaiian Islands. Bulletin of the United States Fish Commission for 1903, 22 (pt. 2, sect. 2)."type="book"year="1905">
eventually prove to be the same. Although specimens of this species are available in many collections around the world, no one has yet to comprehensively study these collections. A molecular study may prove the easiest and the most-effective approach to resolving this question of how many species are involved in this clade. We have taken a conservative approach and treat these northwestern Pacific specimens as
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