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189 lines
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<document id="942E6577EBCD850AA68BA37ADA730154" ID-DOI="10.1649/984.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5370732" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630269392519" checkinUser="valdenar" docAuthor="Philips, T. Keith & Bell, Karen L." docDate="2008" docId="03D687D5C277A538581EFDDDFD52FB07" docLanguage="en" docName="984.1.pdf" docOrigin="The Coleopterists Bulletin 62 (1)" docSource="http://dx.doi.org/10.1649/984.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Attavicinus Philips and Bell 2008, new genus" docType="treatment" docVersion="3" lastPageNumber="81" masterDocId="FFEFFFADC275A53C5974FFD1FFC5FFE2" masterDocTitle="Attavicinus, a New Generic Name for the Myrmecophilous Dung Beetle Liatongus monstrosus (Scarabaeidae: Scarabaeinae)" masterLastPageNumber="81" masterPageNumber="67" pageNumber="67" updateTime="1699205956304" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="BE8413E9603A3BFFE4A055885DC5AFEC">Attavicinus, a New Generic Name for the Myrmecophilous Dung Beetle Liatongus monstrosus (Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="5AA81C23BEA868E17FA7118D544FFC61">Philips, T. Keith</mods:namePart>
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<mods:roleTerm id="FF06841CCAFDBBBC454A6D132971FA4A">Author</mods:roleTerm>
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<mods:namePart id="0DE1C2D27AE814C47F47B46E693309E1">Bell, Karen L.</mods:namePart>
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<mods:typeOfResource id="2F11A7C34618AABD5C9295CAA97F9BF5">text</mods:typeOfResource>
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<mods:title id="B4651AA41D17B049BC2C9C6D90A7CC51">The Coleopterists Bulletin</mods:title>
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<mods:part id="C022F238DA1679494A1B880BCDB03C66">
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<mods:date id="68B48B77E3024AE82D2520D623FFE45D">2008</mods:date>
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<mods:number id="7D0FC66FDEC6E470D9E08B3F4F98EB18">2008-03-31</mods:number>
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<mods:detail id="51FC8A681655F8829B2E553153F400D0" type="volume">
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<mods:number id="2E1AF32F332376E7E8555CAB28308646">62</mods:number>
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<mods:number id="9892B0E4CDC3DE996FADE6DB01128B44">1</mods:number>
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<mods:url id="15417A8133A4526E0947D7BDDE7206A9">http://dx.doi.org/10.1649/984.1</mods:url>
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<mods:classification id="5EB5A9A12CA8AF7DE8537C3CD4995425">journal article</mods:classification>
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<mods:identifier id="CBDEFB690B9EC47EB49E6F18E287B16D" type="DOI">10.1649/984.1</mods:identifier>
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<mods:identifier id="329B69081BD0D777CB7E6D88E1F7A3FA" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="318EAD538FCA503D7EE525776265A0B2" type="Zenodo-Dep">5370732</mods:identifier>
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</mods:mods>
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<treatment id="03D687D5C277A538581EFDDDFD52FB07" LSID="urn:lsid:plazi:treatment:03D687D5C277A538581EFDDDFD52FB07" httpUri="http://treatment.plazi.org/id/03D687D5C277A538581EFDDDFD52FB07" lastPageId="4" pageId="2">
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<subSubSection id="C3656548C277A53E581EFDDDFD32FDC0" box="[362,759,524,546]" pageId="2" type="nomenclature">
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<paragraph id="8BC036C3C277A53E581EFDDDFD32FDC0" blockId="2.[362,759,524,573]" box="[362,759,524,546]" pageId="2">
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<heading id="D08881AFC277A53E581EFDDDFD32FDC0" box="[362,759,524,546]" centered="true" fontSize="9" level="2" pageId="2" reason="2">
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<emphasis id="B90BEAD1C277A53E581EFDDDFD32FDC0" bold="true" box="[362,759,524,546]" pageId="2">
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<taxonomicName id="4C7F4D40C277A53E581EFDDDFD41FDC0" ID-CoL="35F4" authority="Philips and Bell" authorityName="Philips and Bell" authorityYear="2008" box="[362,644,524,546]" class="Insecta" family="Scarabaeidae" genus="Attavicinus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90BEAD1C277A53E581EFDDDFE1FFDC0" bold="true" box="[362,474,524,546]" italics="true" pageId="2">Attavicinus</emphasis>
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Philips and Bell
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23857AAC277A53E5BE5FDDDFD32FDC0" box="[657,759,524,546]" pageId="2" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656548C277A53E5880FDF6FDABFDDF" box="[500,622,551,573]" pageId="2" type="description">
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<paragraph id="8BC036C3C277A53E5880FDF6FDABFDDF" blockId="2.[362,759,524,573]" box="[500,622,551,573]" pageId="2">
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(
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<figureCitation id="13442A46C277A53E588FFDF6FDA3FDDF" box="[507,614,551,573]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Fig" captionStartId-0="3.[196,227,1288,1307]" captionStartId-1="4.[167,198,853,872]" captionStartId-2="5.[196,235,977,996]" captionStartId-3="5.[196,235,1526,1545]" captionStartId-4="6.[167,206,1363,1382]" captionStartId-5="7.[196,227,763,782]" captionTargetBox-0="[200,980,195,1265]" captionTargetBox-1="[228,894,195,829]" captionTargetBox-2="[176,1003,194,951]" captionTargetBox-3="[176,992,1128,1492]" captionTargetBox-4="[156,974,208,1341]" captionTargetBox-5="[176,1003,194,741]" captionTargetId-0="figure-105@3.[199,980,194,1265]" captionTargetId-1="figure-210@4.[226,895,194,830]" captionTargetId-2="figure-1@5.[176,1003,194,955]" captionTargetId-3="figure-44@5.[176,1003,1121,1504]" captionTargetId-4="figure-0@6.[147,974,194,1341]" captionTargetId-5="figure-313@7.[176,1003,194,741]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="Fig. 1. Dorsal habitus of Attavicinus monstrosus, female." captionText-1="Fig. 2. Dorsal habitus of Attavicinus monstrosus, male, anterior half." captionText-2="Figs. 3–8. Attavicinus monstrosus head structures. Scale bars equal 0.5 mm. 3) Antenna; 4) Epipharynx, ventral; 5) Mandible, ventral; 6) Mandible, dorsal; 7) Maxillae, dorsal and ventral views (scale bar 5 0.5 mm); 8) Labium, ventral." captionText-3="Figs. 9–10. Attavicinus monstrosus elytron and wing. Scale bars equal 2.0 mm. 9) Elytron; 10) Wing." captionText-4="Figs. 11–15. Attavicinus monstrosus structural features. Scale bars equal 0.5 mm. 11) Pygidium and propygidium, male; 12) Pygidium and propygidium, female; 13) Male genitalia, lateral view; 14) Mesoscutum-scutellum; 15) Ventral surface of mesoventrite, metaventrite and leg bases with right metaleg removed." captionText-5="Fig. 16. Distribution records of Attavicinus monstrosus." figureDoi-0="http://doi.org/10.5281/zenodo.5370734" figureDoi-1="http://doi.org/10.5281/zenodo.5370738" figureDoi-2="http://doi.org/10.5281/zenodo.5370740" figureDoi-3="http://doi.org/10.5281/zenodo.5370742" figureDoi-4="http://doi.org/10.5281/zenodo.5370744" figureDoi-5="http://doi.org/10.5281/zenodo.5370746" httpUri-0="https://zenodo.org/record/5370734/files/figure.png" httpUri-1="https://zenodo.org/record/5370738/files/figure.png" httpUri-2="https://zenodo.org/record/5370740/files/figure.png" httpUri-3="https://zenodo.org/record/5370742/files/figure.png" httpUri-4="https://zenodo.org/record/5370744/files/figure.png" httpUri-5="https://zenodo.org/record/5370746/files/figure.png" pageId="2">Figs. 1–16</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656548C277A53E59DEFD89FD51FD8C" box="[170,660,600,622]" pageId="2" type="type_taxon">
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<paragraph id="8BC036C3C277A53E59DEFD89FD51FD8C" blockId="2.[146,975,600,1570]" box="[170,660,600,622]" pageId="2">
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<emphasis id="B90BEAD1C277A53E59DEFD89FEEDFD8C" bold="true" box="[170,296,600,622]" pageId="2">
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<typeStatus id="54C48861C277A53E59DEFD89FF19FD8C" box="[170,220,600,622]" pageId="2">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C7F4D40C277A53E5843FD89FD55FD8C" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[311,656,600,622]" class="Insecta" family="Scarabaeidae" genus="Oniticellus" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="montrosus">
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<emphasis id="B90BEAD1C277A53E5843FD89FDCAFD8F" box="[311,527,600,621]" italics="true" pageId="2">Oniticellus montrosus</emphasis>
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<bibRefCitation id="EFEE4B32C277A53E5B63FD89FD55FD8C" author="Bates" box="[535,656,600,622]" pageId="2" refId="ref6592" refString="Bates, H. W. 1887. Biologia Centrali-Americana, Insecta, Coleoptera, Copridae, Aphodiidae, Orphnidae, Hybosoridae, Geotrupidae, Trogidae, Aclopidae, Chasmatopteridae, Melolothidae, vol 2, pt. 2." type="journal volume" year="1887">Bates, 1887</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656548C277A53859DEFDA5FE98FBA6" lastPageId="4" pageId="2" type="description">
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<paragraph id="8BC036C3C277A53E59DEFDA5FCF4FD68" blockId="2.[146,975,600,1570]" box="[170,817,628,650]" pageId="2">
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<emphasis id="B90BEAD1C277A53E59DEFDA5FEFDFD68" bold="true" box="[170,312,628,650]" pageId="2">Redescription.</emphasis>
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<emphasis id="B90BEAD1C277A53E5834FDA5FE52FD6B" box="[320,407,628,649]" italics="true" pageId="2">Habitus.</emphasis>
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Body robust (
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<figureCitation id="13442A46C277A53E5B59FDA5FD4BFD68" box="[557,654,628,650]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[196,227,1288,1307]" captionStartId-1="4.[167,198,853,872]" captionTargetBox-0="[200,980,195,1265]" captionTargetBox-1="[228,894,195,829]" captionTargetId-0="figure-105@3.[199,980,194,1265]" captionTargetId-1="figure-210@4.[226,895,194,830]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1. Dorsal habitus of Attavicinus monstrosus, female." captionText-1="Fig. 2. Dorsal habitus of Attavicinus monstrosus, male, anterior half." figureDoi-0="http://doi.org/10.5281/zenodo.5370734" figureDoi-1="http://doi.org/10.5281/zenodo.5370738" httpUri-0="https://zenodo.org/record/5370734/files/figure.png" httpUri-1="https://zenodo.org/record/5370738/files/figure.png" pageId="2">Figs. 1–2</figureCitation>
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), dark in color.
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</paragraph>
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<paragraph id="8BC036C3C277A53E59DEFD5EFC0CFBD8" blockId="2.[146,975,600,1570]" pageId="2">
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Head with clypeus broadly rounded; external genal margin broadly rounded, posteriorly abruptly angled inward at eye; eye small; female with more pronounced armature consisting of anterior transverse ridge and basal protuberances, male with only basal protuberance. Antennae (
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<figureCitation id="13442A46C277A53E5A6FFD31FC9DFD14" box="[795,856,736,758]" captionStart="Figs" captionStartId="5.[196,235,977,996]" captionTargetBox="[176,1003,194,951]" captionTargetId="figure-1@5.[176,1003,194,955]" captionTargetPageId="5" captionText="Figs. 3–8. Attavicinus monstrosus head structures. Scale bars equal 0.5 mm. 3) Antenna; 4) Epipharynx, ventral; 5) Mandible, ventral; 6) Mandible, dorsal; 7) Maxillae, dorsal and ventral views (scale bar 5 0.5 mm); 8) Labium, ventral." figureDoi="http://doi.org/10.5281/zenodo.5370740" httpUri="https://zenodo.org/record/5370740/files/figure.png" pageId="2">Fig. 3</figureCitation>
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) with eight segments, last three lamellate, vesicle or cavity present in antennomeres 7 and 8. Epipharynx (
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<figureCitation id="13442A46C277A53E586DFCC7FE92FCCE" box="[281,343,790,812]" captionStart="Figs" captionStartId="5.[196,235,977,996]" captionTargetBox="[176,1003,194,951]" captionTargetId="figure-1@5.[176,1003,194,955]" captionTargetPageId="5" captionText="Figs. 3–8. Attavicinus monstrosus head structures. Scale bars equal 0.5 mm. 3) Antenna; 4) Epipharynx, ventral; 5) Mandible, ventral; 6) Mandible, dorsal; 7) Maxillae, dorsal and ventral views (scale bar 5 0.5 mm); 8) Labium, ventral." figureDoi="http://doi.org/10.5281/zenodo.5370740" httpUri="https://zenodo.org/record/5370740/files/figure.png" pageId="2">Fig. 4</figureCitation>
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) with apex truncate, anterior edge medially with distinct setal tuft, second tuft just posterior with relatively few coarse setae, lateral (setal) combs with setal insertion points forming a curved line, remaining ventral surface with relatively evenly scattered fine setae. Mandibles (
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<figureCitation id="13442A46C277A53E5B9BFCB6FC8AFC9F" box="[751,847,871,893]" captionStart="Figs" captionStartId="5.[196,235,977,996]" captionTargetBox="[176,1003,194,951]" captionTargetId="figure-1@5.[176,1003,194,955]" captionTargetPageId="5" captionText="Figs. 3–8. Attavicinus monstrosus head structures. Scale bars equal 0.5 mm. 3) Antenna; 4) Epipharynx, ventral; 5) Mandible, ventral; 6) Mandible, dorsal; 7) Maxillae, dorsal and ventral views (scale bar 5 0.5 mm); 8) Labium, ventral." figureDoi="http://doi.org/10.5281/zenodo.5370740" httpUri="https://zenodo.org/record/5370740/files/figure.png" pageId="2">Fig. 5, 6</figureCitation>
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) with apex relatively sharply acute. Maxillary palp (
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<figureCitation id="13442A46C277A53E5B4CFC53FDB0FC7A" box="[568,629,898,920]" captionStart="Figs" captionStartId="5.[196,235,977,996]" captionTargetBox="[176,1003,194,951]" captionTargetId="figure-1@5.[176,1003,194,955]" captionTargetPageId="5" captionText="Figs. 3–8. Attavicinus monstrosus head structures. Scale bars equal 0.5 mm. 3) Antenna; 4) Epipharynx, ventral; 5) Mandible, ventral; 6) Mandible, dorsal; 7) Maxillae, dorsal and ventral views (scale bar 5 0.5 mm); 8) Labium, ventral." figureDoi="http://doi.org/10.5281/zenodo.5370740" httpUri="https://zenodo.org/record/5370740/files/figure.png" pageId="2">Fig. 7</figureCitation>
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) four-segmented, first palpomere shortest, fourth nearly as long as previous three segments combined; galea broad, wider than long, scoop-like, lacinia much smaller. Labial palp (
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<figureCitation id="13442A46C277A53E5A37FC69FC45FC2C" box="[835,896,952,974]" captionStart="Figs" captionStartId="5.[196,235,977,996]" captionTargetBox="[176,1003,194,951]" captionTargetId="figure-1@5.[176,1003,194,955]" captionTargetPageId="5" captionText="Figs. 3–8. Attavicinus monstrosus head structures. Scale bars equal 0.5 mm. 3) Antenna; 4) Epipharynx, ventral; 5) Mandible, ventral; 6) Mandible, dorsal; 7) Maxillae, dorsal and ventral views (scale bar 5 0.5 mm); 8) Labium, ventral." figureDoi="http://doi.org/10.5281/zenodo.5370740" httpUri="https://zenodo.org/record/5370740/files/figure.png" pageId="2">Fig. 8</figureCitation>
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) threesegmented, second palpomere enlarged, third strongly reduced, barely visible; mentum transverse, anteriorly with shallow notch sharply emarginate medially, laterally with small notch, surface hirsute, medially smooth and impunctate; anterior declivous region broad, triangular. Eye completely divided by canthus.
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</paragraph>
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<paragraph id="8BC036C3C277A53E59DEFB91FD14FB3F" blockId="2.[146,975,600,1570]" pageId="2">
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Pronotum (
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<figureCitation id="13442A46C277A53E5855FB91FE45FBB4" box="[289,384,1088,1110]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[196,227,1288,1307]" captionStartId-1="4.[167,198,853,872]" captionTargetBox-0="[200,980,195,1265]" captionTargetBox-1="[228,894,195,829]" captionTargetId-0="figure-105@3.[199,980,194,1265]" captionTargetId-1="figure-210@4.[226,895,194,830]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1. Dorsal habitus of Attavicinus monstrosus, female." captionText-1="Fig. 2. Dorsal habitus of Attavicinus monstrosus, male, anterior half." figureDoi-0="http://doi.org/10.5281/zenodo.5370734" figureDoi-1="http://doi.org/10.5281/zenodo.5370738" httpUri-0="https://zenodo.org/record/5370734/files/figure.png" httpUri-1="https://zenodo.org/record/5370738/files/figure.png" pageId="2">Figs. 1–2</figureCitation>
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) laterally straight-edged, broadening anteriorly, protruding ridges on either side of middle, more pronounced sculpture on female compared to male. Depressed at middle, small slight oval depression at posterior margin opposite scutellum, laterally with shiny smooth depression at anterolateral angles. Scutellum (
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<figureCitation id="13442A46C277A53E58E9FB7DFE22FB20" box="[413,487,1196,1218]" captionStart="Figs" captionStartId="6.[167,206,1363,1382]" captionTargetBox="[156,974,208,1341]" captionTargetId="figure-0@6.[147,974,194,1341]" captionTargetPageId="6" captionText="Figs. 11–15. Attavicinus monstrosus structural features. Scale bars equal 0.5 mm. 11) Pygidium and propygidium, male; 12) Pygidium and propygidium, female; 13) Male genitalia, lateral view; 14) Mesoscutum-scutellum; 15) Ventral surface of mesoventrite, metaventrite and leg bases with right metaleg removed." figureDoi="http://doi.org/10.5281/zenodo.5370744" httpUri="https://zenodo.org/record/5370744/files/figure.png" pageId="2">Fig. 14</figureCitation>
|
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|
|
) with visible portion triangular in shape, short and broad, apex narrowly rounded, parabolic in shape.
|
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|
|
</paragraph>
|
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|
|
<paragraph id="8BC036C3C277A53E59DEFB32FD0CFA0E" blockId="2.[146,975,600,1570]" pageId="2">
|
|||
|
|
Elytra (
|
|||
|
|
<figureCitation id="13442A46C277A53E598FFB32FEFDFB1B" box="[251,312,1251,1273]" captionStart="Figs" captionStartId="5.[196,235,1526,1545]" captionTargetBox="[176,992,1128,1492]" captionTargetId="figure-44@5.[176,1003,1121,1504]" captionTargetPageId="5" captionText="Figs. 9–10. Attavicinus monstrosus elytron and wing. Scale bars equal 2.0 mm. 9) Elytron; 10) Wing." figureDoi="http://doi.org/10.5281/zenodo.5370742" httpUri="https://zenodo.org/record/5370742/files/figure.png" pageId="2">Fig. 9</figureCitation>
|
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|
|
) with surface uneven, raised towards anterior on sides. Suture depressed anteriorly and raised posteriorly. Each elytron wih seven visible striae, each consisting of a single shallow groove. Hind wings (
|
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|
|
<figureCitation id="13442A46C277A53E5BA0FAC8FCDBFACD" box="[724,798,1305,1327]" captionStart="Figs" captionStartId="5.[196,235,1526,1545]" captionTargetBox="[176,992,1128,1492]" captionTargetId="figure-44@5.[176,1003,1121,1504]" captionTargetPageId="5" captionText="Figs. 9–10. Attavicinus monstrosus elytron and wing. Scale bars equal 2.0 mm. 9) Elytron; 10) Wing." figureDoi="http://doi.org/10.5281/zenodo.5370742" httpUri="https://zenodo.org/record/5370742/files/figure.png" pageId="2">Fig. 10</figureCitation>
|
|||
|
|
) fully developed, pigmented throughout; AA vein lacking remnant cross vein proximally, MP vein present, long in length, pigment patch near apical 1/3 present and distal edge emarginated and not strongly defined, anal region large, well developed, lacking posterior notch, jugal and AP veins both present, MP vein near posterior margin of wing distinctly, strongly curved, RA vein greatly extended along apical margin, ending near posterior margin, AA vein posteriorly forming cell partially closed, extending on both sides at proximal apex near middle.
|
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|
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</paragraph>
|
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|
|
<paragraph id="8BC036C3C277A53F59DEFA20FC29FAAF" blockId="2.[146,975,600,1570]" lastBlockId="3.[175,1004,1335,1570]" lastPageId="3" pageId="2">Legs robust and stout; tarsi distinct, strongly developed, each with five short tarsomeres. Protibiae with four teeth, apical three strongly developed, basal tooth distinct but weakly projecting. Meso- and metatibia gradually expanding in basal</paragraph>
|
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|
|
<caption id="DF00664BC276A53F59B0FAD9FD19FAF9" ID-DOI="http://doi.org/10.5281/zenodo.5370734" ID-Zenodo-Dep="5370734" box="[196,732,1288,1307]" httpUri="https://zenodo.org/record/5370734/files/figure.png" pageId="3" startId="3.[196,227,1288,1307]" targetBox="[200,980,195,1265]" targetPageId="3">
|
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|
|
<paragraph id="8BC036C3C276A53F59B0FAD9FD19FAF9" blockId="3.[196,732,1288,1307]" box="[196,732,1288,1307]" pageId="3">
|
|||
|
|
<emphasis id="B90BEAD1C276A53F59B0FAD9FEC7FAF9" bold="true" box="[196,258,1288,1307]" pageId="3">Fig. 1.</emphasis>
|
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|
|
Dorsal habitus of
|
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|
|
<taxonomicName id="4C7F4D40C276A53F58B4FAD9FD49FAF9" authorityName="Philips & Bell" authorityYear="2008" baseAuthorityName=": Halffter and Matthews" baseAuthorityYear="2008" box="[448,652,1288,1307]" class="Insecta" family="Scarabaeidae" genus="Attavicinus" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="monstrosus">
|
|||
|
|
<emphasis id="B90BEAD1C276A53F58B4FAD9FD49FAF9" box="[448,652,1288,1307]" italics="true" pageId="3">Attavicinus monstrosus</emphasis>
|
|||
|
|
</taxonomicName>
|
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|
|
, female.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC036C3C276A53F59DBFA80FDC9FA60" blockId="3.[175,1004,1335,1570]" pageId="3">
|
|||
|
|
<emphasis id="B90BEAD1C276A53F59DBFA80FF02FA85" bold="true" box="[175,199,1361,1383]" pageId="3">L</emphasis>
|
|||
|
|
, greatly expanded at apical
|
|||
|
|
<emphasis id="B90BEAD1C276A53F5B75FA80FDDCFA85" bold="true" box="[513,537,1361,1383]" pageId="3">J</emphasis>
|
|||
|
|
, mesotibial apex with two spurs, pro- and metatibia each with a single spur.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC036C3C276A53859B3FA59FCA3FC16" blockId="3.[175,1004,1335,1570]" lastBlockId="4.[146,976,937,1253]" lastPageId="4" pageId="3">
|
|||
|
|
Ventral surface with prosternum nearly equal in width to procoxal width, distinctly longer towards lateral margin; mesosternum (
|
|||
|
|
<figureCitation id="13442A46C276A53F5B93FA73FCF5FA5A" box="[743,816,1442,1464]" captionStart="Figs" captionStartId="6.[167,206,1363,1382]" captionTargetBox="[156,974,208,1341]" captionTargetId="figure-0@6.[147,974,194,1341]" captionTargetPageId="6" captionText="Figs. 11–15. Attavicinus monstrosus structural features. Scale bars equal 0.5 mm. 11) Pygidium and propygidium, male; 12) Pygidium and propygidium, female; 13) Male genitalia, lateral view; 14) Mesoscutum-scutellum; 15) Ventral surface of mesoventrite, metaventrite and leg bases with right metaleg removed." figureDoi="http://doi.org/10.5281/zenodo.5370744" httpUri="https://zenodo.org/record/5370744/files/figure.png" pageId="3">Fig. 15</figureCitation>
|
|||
|
|
) with small lanceshaped ventral projection on anterior margin at middle, hidden in intact specimens; metepisternum subtriangular, broad anteriorly, narrowing to a point posteriorly. Abdominal ventrites 3–5 similar in length, shorter in middle and longer laterally and curved; ventrite 2 is 1.5 times the length of ventrites 3–5 at the 70 middle, similar length laterally and straight; ventrite 1 longest. Propygidium (
|
|||
|
|
<figureCitation id="13442A46C271A53859EEFC12FED3FC3B" box="[154,278,963,985]" captionStart="Figs" captionStartId="6.[167,206,1363,1382]" captionTargetBox="[156,974,208,1341]" captionTargetId="figure-0@6.[147,974,194,1341]" captionTargetPageId="6" captionText="Figs. 11–15. Attavicinus monstrosus structural features. Scale bars equal 0.5 mm. 11) Pygidium and propygidium, male; 12) Pygidium and propygidium, female; 13) Male genitalia, lateral view; 14) Mesoscutum-scutellum; 15) Ventral surface of mesoventrite, metaventrite and leg bases with right metaleg removed." figureDoi="http://doi.org/10.5281/zenodo.5370744" httpUri="https://zenodo.org/record/5370744/files/figure.png" pageId="4">Figs. 11, 12</figureCitation>
|
|||
|
|
) indistinct from pygidium, shorter than ventrites 1–5, with slight transverse groove anteriorly; pygidium (
|
|||
|
|
<figureCitation id="13442A46C271A5385B45FC0FFD6EFC16" box="[561,683,990,1012]" captionStart="Figs" captionStartId="6.[167,206,1363,1382]" captionTargetBox="[156,974,208,1341]" captionTargetId="figure-0@6.[147,974,194,1341]" captionTargetPageId="6" captionText="Figs. 11–15. Attavicinus monstrosus structural features. Scale bars equal 0.5 mm. 11) Pygidium and propygidium, male; 12) Pygidium and propygidium, female; 13) Male genitalia, lateral view; 14) Mesoscutum-scutellum; 15) Ventral surface of mesoventrite, metaventrite and leg bases with right metaleg removed." figureDoi="http://doi.org/10.5281/zenodo.5370744" httpUri="https://zenodo.org/record/5370744/files/figure.png" pageId="4">Figs. 11, 12</figureCitation>
|
|||
|
|
) broadly pointed.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF00664BC271A53859D3FC84FCEBFC8A" ID-DOI="http://doi.org/10.5281/zenodo.5370738" ID-Zenodo-Dep="5370738" box="[167,814,853,872]" httpUri="https://zenodo.org/record/5370738/files/figure.png" pageId="4" startId="4.[167,198,853,872]" targetBox="[228,894,195,829]" targetPageId="4">
|
|||
|
|
<paragraph id="8BC036C3C271A53859D3FC84FCEBFC8A" blockId="4.[167,814,853,872]" box="[167,814,853,872]" pageId="4">
|
|||
|
|
<emphasis id="B90BEAD1C271A53859D3FC84FF20FC8A" bold="true" box="[167,229,853,872]" pageId="4">Fig. 2.</emphasis>
|
|||
|
|
Dorsal habitus of
|
|||
|
|
<taxonomicName id="4C7F4D40C271A53858D7FC84FDAAFC8A" authorityName="Philips & Bell" authorityYear="2008" baseAuthorityName=": Halffter and Matthews" baseAuthorityYear="2008" box="[419,623,853,872]" class="Insecta" family="Scarabaeidae" genus="Attavicinus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="monstrosus">
|
|||
|
|
<emphasis id="B90BEAD1C271A53858D7FC84FDAAFC8A" box="[419,623,853,872]" italics="true" pageId="4">Attavicinus monstrosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male, anterior half.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC036C3C271A53859DEFC28FE98FBA6" blockId="4.[146,976,937,1253]" pageId="4">
|
|||
|
|
Male genitalia (
|
|||
|
|
<figureCitation id="13442A46C271A5385826FC28FE5EFBED" box="[338,411,1017,1039]" captionStart="Figs" captionStartId="6.[167,206,1363,1382]" captionTargetBox="[156,974,208,1341]" captionTargetId="figure-0@6.[147,974,194,1341]" captionTargetPageId="6" captionText="Figs. 11–15. Attavicinus monstrosus structural features. Scale bars equal 0.5 mm. 11) Pygidium and propygidium, male; 12) Pygidium and propygidium, female; 13) Male genitalia, lateral view; 14) Mesoscutum-scutellum; 15) Ventral surface of mesoventrite, metaventrite and leg bases with right metaleg removed." figureDoi="http://doi.org/10.5281/zenodo.5370744" httpUri="https://zenodo.org/record/5370744/files/figure.png" pageId="4">Fig. 13</figureCitation>
|
|||
|
|
) broad; parameres less than half the length of basal piece, apex blunt, ventromedially with small tooth, paramere position relative to basal piece oblique.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3656548C271A53859DEFB9BFC15FB77" pageId="4" type="etymology">
|
|||
|
|
<paragraph id="8BC036C3C271A53859DEFB9BFC15FB77" blockId="4.[146,976,937,1253]" pageId="4">
|
|||
|
|
<emphasis id="B90BEAD1C271A53859DEFB9BFED8FB82" bold="true" box="[170,285,1098,1120]" pageId="4">Etymology.</emphasis>
|
|||
|
|
This new generic name is derived from the host ant genus
|
|||
|
|
<taxonomicName id="4C7F4D40C271A5385AD6FB9BFC0AFBBD" authorityName="Fabricius" authorityYear="1804" box="[930,975,1098,1119]" class="Insecta" family="Formicidae" genus="Atta" kingdom="Animalia" order="Hymenoptera" pageId="4" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B90BEAD1C271A5385AD6FB9BFC0AFBBD" box="[930,975,1098,1119]" italics="true" pageId="4">Atta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
together with the Latin ‘‘vicini’’ that means neighboring or near, in reference to the association with the debris piles of leaf-cutter ants. The gender is masculine.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3656548C271A53859DEFB4BFD52FB07" pageId="4" type="discussion">
|
|||
|
|
<paragraph id="8BC036C3C271A53859DEFB4BFD52FB07" blockId="4.[146,976,937,1253]" pageId="4">
|
|||
|
|
<emphasis id="B90BEAD1C271A53859DEFB4BFEDFFB52" bold="true" box="[170,282,1178,1200]" pageId="4">Comments.</emphasis>
|
|||
|
|
<materialsCitation id="3B173C9EC271A5385852FB4BFD56FB07" country="Mexico" location="Jalisco" pageId="4" specimenCount="1" stateProvince="Jalisco">
|
|||
|
|
This genus includes a single species found only in
|
|||
|
|
<collectingRegion id="49BBF821C271A5385A2CFB4BFC59FB52" box="[856,924,1178,1200]" country="Mexico" name="Jalisco" pageId="4">Jalisco</collectingRegion>
|
|||
|
|
and
|
|||
|
|
<collectingRegion id="49BBF821C271A53859E6FB64FEC2FB29" box="[146,263,1205,1227]" country="Mexico" name="Michoacan de Ocampo" pageId="4">Michoacán</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F3687653C271A5385860FB64FEA5FB29" box="[276,352,1205,1227]" name="Mexico" pageId="4">Mexico</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="13442A46C271A5385805FB64FE7BFB29" box="[369,446,1205,1227]" captionStart="Fig" captionStartId="7.[196,227,763,782]" captionTargetBox="[176,1003,194,741]" captionTargetId="figure-313@7.[176,1003,194,741]" captionTargetPageId="7" captionText="Fig. 16. Distribution records of Attavicinus monstrosus." figureDoi="http://doi.org/10.5281/zenodo.5370746" httpUri="https://zenodo.org/record/5370746/files/figure.png" pageId="4">Fig. 16</figureCitation>
|
|||
|
|
). Adults and larvae can be found within or under attine ant nest debris piles where breeding occurs
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|