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<document id="C0F78CCB3F11175BBE9CD2B7A5A13F2C" ID-CLB-Dataset="280650" ID-DOI="10.15407/zoo2023.01.019" ID-GBIF-Dataset="c28542b3-7e6f-4bac-af73-52113ff5d050" ID-ISSN="2707-7268" ID-Zenodo-Dep="10263894" ID-ZooBank="1F6FA9AB-88F0-4C1B-8895-6629C23C646B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1681820915329" checkinUser="felipe" docAuthor="Dzeverin, I." docDate="2023" docId="811E87B32A08FF92FF4EFC296E5CFBD1" docLanguage="en" docName="Zoodiversity.57.1.19-28.pdf" docOrigin="Zoodiversity 57 (1)" docSource="http://dx.doi.org/10.15407/zoo2023.01.019" docStyle="DocumentStyle:65E6F982F72656C606C1639D3996B172.2:Zoodiversity.2020-.journal_article" docStyleId="65E6F982F72656C606C1639D3996B172" docStyleName="Zoodiversity.2020-.journal_article" docStyleVersion="2" docTitle="Myotis aurascens Kusjakin 1935" docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="7D27FFCB2A09FF90FFADFF916C06FF8D" masterDocTitle="Synonymisation Of Myotis Aurascens With M. Davidii (Chiroptera, Vespertilionidae) Is Premature" masterLastPageNumber="28" masterPageNumber="19" pageNumber="20" updateTime="1701796663683" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="A85EA228D7F49FB9DBA323A7BD0C1F81">Synonymisation Of Myotis Aurascens With M. Davidii (Chiroptera, Vespertilionidae) Is Premature</mods:title>
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<subSubSection id="41AD652E2A08FF91FF4EFC296844FC5F" box="[227,1090,952,979]" pageId="1" pageNumber="20" type="nomenclature">
<paragraph id="090836A52A08FF91FF4EFC296844FC5F" blockId="1.[174,1233,952,1859]" box="[227,1090,952,979]" pageId="1" pageNumber="20">
<bibRefCitation id="6D264B542A08FF91FF4EFC296DC2FC5F" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[227,452,952,979]" pageId="1" pageNumber="20" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al. (2012)</bibRefCitation>
base the hypothesis of conspecificity of
<taxonomicName id="CEB74D262A08FF91FC0AFC286844FC5F" ID-CoL="8LTYX" authorityName="Kusjakin" authorityYear="1935" box="[935,1090,953,978]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="aurascens">
<emphasis id="3BC3EAB72A08FF91FC0AFC286844FC5F" box="[935,1090,953,978]" italics="true" pageId="1" pageNumber="20">M. aurascens</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="41AD652E2A08FF92FBE6FC296E5CFBD1" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="discussion">
<paragraph id="090836A52A08FF91FBE6FC296D82FB28" blockId="1.[174,1233,952,1859]" pageId="1" pageNumber="20">
and
<taxonomicName id="CEB74D262A08FF91FB2DFC286CE4FC78" baseAuthorityName="Peters" baseAuthorityYear="1869" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A08FF91FB2DFC286CE4FC78" italics="true" pageId="1" pageNumber="20">M. davidii</emphasis>
</taxonomicName>
on (
<emphasis id="3BC3EAB72A08FF91FEB0FC4D6D23FC78" box="[285,293,988,1013]" italics="true" pageId="1" pageNumber="20">i</emphasis>
) morphometric data (
<bibRefCitation id="6D264B542A08FF91FD9DFC4A6F45FC78" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[560,835,987,1014]" pageId="1" pageNumber="20" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al., 2012: 329</bibRefCitation>
, table 16) and scatter plots based on them (
<bibRefCitation id="6D264B542A08FF91FE8DFC6F6E2AFB94" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[288,556,1022,1049]" pageId="1" pageNumber="20" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al., 2012: 330</bibRefCitation>
, figs 95, 96); (
<emphasis id="3BC3EAB72A08FF91FD62FC6E6ED9FB95" box="[719,735,1023,1048]" italics="true" pageId="1" pageNumber="20">ii</emphasis>
) outlines of skulls and the elements of the dental system of the specimens of several whiskered
<taxonomicName id="CEB74D262A08FF91FCB8FBB26F62FBB1" authorityName="Kaup" authorityYear="1829" box="[789,868,1059,1084]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="3BC3EAB72A08FF91FCB8FBB26F62FBB1" box="[789,868,1059,1084]" italics="true" pageId="1" pageNumber="20">Myotis</emphasis>
</taxonomicName>
OTUs (
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, 333, figs 97, 98); (
<emphasis id="3BC3EAB72A08FF91FE2FFBD76D9CFBD2" box="[386,410,1094,1119]" italics="true" pageId="1" pageNumber="20">iii</emphasis>
) the phylogeny reconstruction based on the cytochrome
<emphasis id="3BC3EAB72A08FF91FBE9FBD46854FBD2" box="[1092,1106,1093,1119]" italics="true" pageId="1" pageNumber="20">b</emphasis>
sequences (
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, table 17) and a tree built on the basis of these data (
<bibRefCitation id="6D264B542A08FF91FB91FBF96D26FB28" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." pageId="1" pageNumber="20" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al., 2012: 333</bibRefCitation>
, fig. 99).
</paragraph>
<paragraph id="090836A52A08FF91FF4EFB3F68BBFA82" blockId="1.[174,1233,952,1859]" pageId="1" pageNumber="20">
<bibRefCitation id="6D264B542A08FF91FF4EFB3F6DF7FB44" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[227,497,1198,1225]" pageId="1" pageNumber="20" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al. (2012: 329</bibRefCitation>
, table 16) provided the morphometric data of
<specimenCount id="1FB1FD2C2A08FF91FBB9FB3F68CAFB45" box="[1044,1228,1198,1224]" pageId="1" pageNumber="20" type="generic">nine specimens</specimenCount>
:
<specimenCount id="1FB1FD2C2A08FF91FF03FB436D01FB61" box="[174,263,1234,1260]" pageId="1" pageNumber="20" type="generic">
four
<taxonomicName id="CEB74D262A08FF91FF4BFB426D01FB61" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[230,263,1235,1260]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" sensu="stricto" species="mystacinus">
<emphasis id="3BC3EAB72A08FF91FF4BFB426D01FB61" box="[230,263,1235,1260]" italics="true" pageId="1" pageNumber="20">M.</emphasis>
</taxonomicName>
</specimenCount>
mystacinus
<taxonomicNameLabel id="20F057CC2A08FF91FE39FB436DD4FB61" box="[404,466,1234,1260]" pageId="1" pageNumber="20" sensu="stricto">s. str.</taxonomicNameLabel>
,
<specimenCount id="1FB1FD2C2A08FF91FE70FB436E30FB61" box="[477,566,1234,1260]" pageId="1" pageNumber="20" type="generic">
four
<taxonomicName id="CEB74D262A08FF91FDB8FB426E30FB61" baseAuthorityName="Peters" baseAuthorityYear="1869" box="[533,566,1235,1260]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A08FF91FDB8FB426E30FB61" box="[533,566,1235,1260]" italics="true" pageId="1" pageNumber="20">M.</emphasis>
</taxonomicName>
</specimenCount>
davidii from
<collectingCountry id="71A076352A08FF91FD7EFB436F03FB61" box="[723,773,1234,1260]" name="Iran" pageId="1" pageNumber="20">Iran</collectingCountry>
(i. e.,
<taxonomicName id="CEB74D262A08FF91FCE4FB426FE6FB61" authorityName="Kusjakin" authorityYear="1935" box="[841,992,1235,1260]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="aurascens">
<emphasis id="3BC3EAB72A08FF91FCE4FB426FE6FB61" box="[841,992,1235,1260]" italics="true" pageId="1" pageNumber="20">M. aurascens</emphasis>
</taxonomicName>
in the previous classification by
<bibRefCitation id="6D264B542A08FF91FEE9FB646E81FA82" author="Benda, P. &amp; Tsytsulina, K. A." box="[324,647,1269,1295]" pageId="1" pageNumber="20" pagination="331 - 398" refId="ref5672" refString="Benda, P. &amp; Tsytsulina, K. A. 2000. Taxonomic revision of Myotis mystacinus group (Mammalia: Chiroptera) in the western Palearctic. Acta Societatis Zoologicae Bohemicae, 64 (4), 331 - 398." type="journal article" year="2000">Benda &amp; Tsytsulina (2000))</bibRefCitation>
and a
<typeStatus id="D60C88072A08FF91FD78FB646F01FA82" box="[725,775,1269,1295]" pageId="1" pageNumber="20">type</typeStatus>
specimen of
<taxonomicName id="CEB74D262A08FF91FC0FFB67681CFA82" baseAuthorityName="Peters" baseAuthorityYear="1869" box="[930,1050,1269,1295]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A08FF91FC0FFB67681CFA82" box="[930,1050,1269,1295]" italics="true" pageId="1" pageNumber="20">M. davidii</emphasis>
</taxonomicName>
from Beijing.
</paragraph>
<paragraph id="090836A52A08FF91FF4EFA896F81FAF4" blockId="1.[174,1233,952,1859]" pageId="1" pageNumber="20">
The scatter plots given by
<bibRefCitation id="6D264B542A08FF91FD8DFA896F32FABF" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[544,820,1304,1331]" pageId="1" pageNumber="20" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al. (2012: 330</bibRefCitation>
, figs 95, 96) show that the Iranian specimens are markedly different from
<taxonomicName id="CEB74D262A08FF91FDD1FAAD6F20FAD8" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[636,806,1340,1365]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" sensu="stricto" species="mystacinus">
<emphasis id="3BC3EAB72A08FF91FDD1FAAD6F20FAD8" box="[636,806,1340,1365]" italics="true" pageId="1" pageNumber="20">M. mystacinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20F057CC2A08FF91FC81FAAD6F6DFADB" box="[812,875,1340,1366]" pageId="1" pageNumber="20" sensu="stricto">s. str.</taxonomicNameLabel>
, and that the
<typeStatus id="D60C88072A08FF91FBA7FAAD683AFADB" box="[1034,1084,1340,1366]" pageId="1" pageNumber="20">type</typeStatus>
specimen of
<taxonomicName id="CEB74D262A08FF91FF03FACE6D20FAF5" baseAuthorityName="Peters" baseAuthorityYear="1869" box="[174,294,1374,1400]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A08FF91FF03FACE6D20FAF5" box="[174,294,1374,1400]" italics="true" pageId="1" pageNumber="20">M. davidii</emphasis>
</taxonomicName>
is much closer to them than to
<taxonomicName id="CEB74D262A08FF91FD34FACE6F45FAF5" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[665,835,1375,1400]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" sensu="stricto" species="mystacinus">
<emphasis id="3BC3EAB72A08FF91FD34FACE6F45FAF5" box="[665,835,1375,1400]" italics="true" pageId="1" pageNumber="20">M. mystacinus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="090836A52A08FF91FF4EFA136D63F955" blockId="1.[174,1233,952,1859]" pageId="1" pageNumber="20">
At the same time, the
<typeStatus id="D60C88072A08FF91FE4DFA136E14FA11" box="[480,530,1410,1436]" pageId="1" pageNumber="20">type</typeStatus>
specimen of
<taxonomicName id="CEB74D262A08FF91FD06FA126F25FA11" baseAuthorityName="Peters" baseAuthorityYear="1869" box="[683,803,1410,1436]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A08FF91FD06FA126F25FA11" box="[683,803,1410,1436]" italics="true" pageId="1" pageNumber="20">M. davidii</emphasis>
</taxonomicName>
in some characters falls into the area of Iranian specimens on the plot (
<figureCitation id="918C2A202A08FF91FDECFA346E78FA32" box="[577,638,1445,1471]" captionStart="Fig" captionStartId="4.[166,199,1203,1225]" captionTargetBox="[173,1208,176,1183]" captionTargetId="graphics-277@4.[775,1208,688,1121]" captionTargetPageId="4" captionText="Fig. 1. Bivariate plots of the Myotis samples listed by Benda et al. (2012) to demonstrate the correspondence of Iranian steppe whiskered bats with M. mystacinus s. str. and M. davidii. LCr is the greatest length of skull; LMd is condylar length of mandible; CM3 is length of upper toothrow between C and M3 (incl.); CM is length 3 of lower toothrow between C and M3 (incl.); LaI is width of interorbital constriction; LaN is neurocranium width; CC is rostral width between canines (incl.); M3M3 is rostral width between the third upper molars (incl.). The measurements are in mm. All measurements are from Benda et al. (2012: 329, table 16). Fig. 1, A partly repeats Fig. 95 of Benda et al. (2012: 330), however Benda et al. also added the data given by DeBlase (1980) to this figure as well as to fig. 96. M. mystacinus s. str. specimens are shown in green; M. aurascens specimens from Iran attributed by Benda et al. (2012) to M. davidii are shown in yellow; type specimen of M. davidii is shown in black. See Benda et al. (2012: 329, table 16) for more details about these specimens. The figure was obtained using R software (R Core Team, 2021)." figureDoi="http://doi.org/10.5281/zenodo.10263898" httpUri="https://zenodo.org/record/10263898/files/figure.png" pageId="1" pageNumber="20">fig. 1</figureCitation>
, AD in the present paper). In other characters it does not fall into this area, but is at a place close to this area, being smaller in most characters than Iranian specimens (
<figureCitation id="918C2A202A08FF91FDAAFA7A6E42F988" box="[519,580,1515,1541]" captionStart="Fig" captionStartId="4.[166,199,1203,1225]" captionTargetBox="[173,1208,176,1183]" captionTargetId="graphics-277@4.[775,1208,688,1121]" captionTargetPageId="4" captionText="Fig. 1. Bivariate plots of the Myotis samples listed by Benda et al. (2012) to demonstrate the correspondence of Iranian steppe whiskered bats with M. mystacinus s. str. and M. davidii. LCr is the greatest length of skull; LMd is condylar length of mandible; CM3 is length of upper toothrow between C and M3 (incl.); CM is length 3 of lower toothrow between C and M3 (incl.); LaI is width of interorbital constriction; LaN is neurocranium width; CC is rostral width between canines (incl.); M3M3 is rostral width between the third upper molars (incl.). The measurements are in mm. All measurements are from Benda et al. (2012: 329, table 16). Fig. 1, A partly repeats Fig. 95 of Benda et al. (2012: 330), however Benda et al. also added the data given by DeBlase (1980) to this figure as well as to fig. 96. M. mystacinus s. str. specimens are shown in green; M. aurascens specimens from Iran attributed by Benda et al. (2012) to M. davidii are shown in yellow; type specimen of M. davidii is shown in black. See Benda et al. (2012: 329, table 16) for more details about these specimens. The figure was obtained using R software (R Core Team, 2021)." figureDoi="http://doi.org/10.5281/zenodo.10263898" httpUri="https://zenodo.org/record/10263898/files/figure.png" pageId="1" pageNumber="20">fig. 1</figureCitation>
, A, C). In some additional characters,
<taxonomicName id="CEB74D262A08FF91FBBCFA7D688DF988" baseAuthorityName="Peters" baseAuthorityYear="1869" box="[1041,1163,1515,1541]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A08FF91FBBCFA7D688DF988" box="[1041,1163,1515,1541]" italics="true" pageId="1" pageNumber="20">M. davidii</emphasis>
</taxonomicName>
is intermediate between the Iranian sample and
<taxonomicName id="CEB74D262A08FF91FD00F99E6F50F9A5" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[685,854,1551,1576]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" sensu="stricto" species="mystacinus">
<emphasis id="3BC3EAB72A08FF91FD00F99E6F50F9A5" box="[685,854,1551,1576]" italics="true" pageId="1" pageNumber="20">M. mystacinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20F057CC2A08FF91FCF6F99E6F9EF9A4" box="[859,920,1551,1577]" pageId="1" pageNumber="20" sensu="stricto">s. str.</taxonomicNameLabel>
sample (
<figureCitation id="918C2A202A08FF91FC52F99F683CF9A4" box="[1023,1082,1550,1577]" captionStart="Fig" captionStartId="4.[166,199,1203,1225]" captionTargetBox="[173,1208,176,1183]" captionTargetId="graphics-277@4.[775,1208,688,1121]" captionTargetPageId="4" captionText="Fig. 1. Bivariate plots of the Myotis samples listed by Benda et al. (2012) to demonstrate the correspondence of Iranian steppe whiskered bats with M. mystacinus s. str. and M. davidii. LCr is the greatest length of skull; LMd is condylar length of mandible; CM3 is length of upper toothrow between C and M3 (incl.); CM is length 3 of lower toothrow between C and M3 (incl.); LaI is width of interorbital constriction; LaN is neurocranium width; CC is rostral width between canines (incl.); M3M3 is rostral width between the third upper molars (incl.). The measurements are in mm. All measurements are from Benda et al. (2012: 329, table 16). Fig. 1, A partly repeats Fig. 95 of Benda et al. (2012: 330), however Benda et al. also added the data given by DeBlase (1980) to this figure as well as to fig. 96. M. mystacinus s. str. specimens are shown in green; M. aurascens specimens from Iran attributed by Benda et al. (2012) to M. davidii are shown in yellow; type specimen of M. davidii is shown in black. See Benda et al. (2012: 329, table 16) for more details about these specimens. The figure was obtained using R software (R Core Team, 2021)." figureDoi="http://doi.org/10.5281/zenodo.10263898" httpUri="https://zenodo.org/record/10263898/files/figure.png" pageId="1" pageNumber="20">fig. 1</figureCitation>
, B, D). Since variation patterns seem to be different for different characters, skull shape can differ in the studied OTUs. These differences do not refute the conspecificity of Iranian and Chinese steppe whiskered bats (differences are expected for populations living in geographically remote and ecologically different localities), but also they cannot be evidence in support of this hypothesis.
</paragraph>
<paragraph id="090836A52A08FF92FF4EF9736F75FE9D" blockId="1.[174,1233,952,1859]" lastBlockId="2.[166,1225,175,1117]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
The name-bearing
<typeStatus id="D60C88072A08FF91FE6AF9736DFFF971" box="[455,505,1762,1788]" pageId="1" pageNumber="20">type</typeStatus>
specimen of
<taxonomicName id="CEB74D262A08FF91FD36F9736F77F971" box="[667,881,1762,1788]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A08FF91FD36F9736F77F971" box="[667,881,1762,1788]" italics="true" pageId="1" pageNumber="20">Vespertilio davidii</emphasis>
</taxonomicName>
has not been compared with the
<typeStatus id="D60C88072A08FF91FF74F8946D0DF892" box="[217,267,1797,1823]" pageId="1" pageNumber="20">type</typeStatus>
specimens of
<taxonomicName id="CEB74D262A08FF91FE02F8976E26F892" authorityName="Kusjakin" authorityYear="1935" box="[431,544,1798,1823]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="aurascens">
<emphasis id="3BC3EAB72A08FF91FE02F8976E26F892" box="[431,544,1798,1823]" italics="true" pageId="1" pageNumber="20">aurascens</emphasis>
</taxonomicName>
,
<taxonomicName id="CEB74D262A08FF91FD80F8976E7CF892" box="[557,634,1798,1823]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="popovi">
<emphasis id="3BC3EAB72A08FF91FD80F8976E7CF892" box="[557,634,1798,1823]" italics="true" pageId="1" pageNumber="20">popovi</emphasis>
</taxonomicName>
,
<taxonomicName id="CEB74D262A08FF91FD2AF8946EFFF892" authorityName="Kuzyakin" authorityYear="1934" box="[647,761,1797,1823]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="1" pageNumber="20" phylum="Chordata" rank="species" species="sogdianus">
<emphasis id="3BC3EAB72A08FF91FD2AF8946EFFF892" box="[647,761,1797,1823]" italics="true" pageId="1" pageNumber="20">sogdianus</emphasis>
</taxonomicName>
, and a number of other OTUs (possibly, subspecific) of the steppe whiskered bat (while some other
<typeStatus id="D60C88072A08FF91FCF5F8B86F8CF8CE" box="[856,906,1833,1859]" pageId="1" pageNumber="20">type</typeStatus>
specimens are shown in the plots). In addition, samples of only
<specimenCount id="1FB1FD2C2A0BFF92FDFDFF3E6F03FF44" box="[592,773,175,201]" pageId="2" pageNumber="21" type="generic">four specimens</specimenCount>
are insufficient when such a complex group of taxa as the
<taxonomicName id="CEB74D262A0BFF92FE0EFF426E56FF61" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[419,592,211,236]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="mystacinus">
<emphasis id="3BC3EAB72A0BFF92FE0EFF426E56FF61" box="[419,592,211,236]" italics="true" pageId="2" pageNumber="21">M. mystacinus</emphasis>
</taxonomicName>
complex, which comprises a number of OTUs with considerable between- and within-group variation is studied.
</paragraph>
<paragraph id="090836A52A0BFF92FF76FE88680AFEF4" blockId="2.[166,1225,175,1117]" pageId="2" pageNumber="21">
At most we can say from the morphometric data provided by
<bibRefCitation id="6D264B542A0BFF92FC1BFE886895FEBE" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[950,1171,281,307]" pageId="2" pageNumber="21" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al. (2012)</bibRefCitation>
that the studied Iranian steppe whiskered bats are more similar to the
<taxonomicName id="CEB74D262A0BFF92FC0FFEAC681CFEDB" box="[930,1050,316,342]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A0BFF92FC0FFEAC681CFEDB" box="[930,1050,316,342]" italics="true" pageId="2" pageNumber="21">M. davidii</emphasis>
</taxonomicName>
<typeStatus id="D60C88072A0BFF92FB8CFEAC6855FEDA" box="[1057,1107,317,343]" pageId="2" pageNumber="21">type</typeStatus>
specimen than to the studied
<taxonomicName id="CEB74D262A0BFF92FE27FEF16E35FEF4" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[394,563,352,377]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" sensu="stricto" species="mystacinus">
<emphasis id="3BC3EAB72A0BFF92FE27FEF16E35FEF4" box="[394,563,352,377]" italics="true" pageId="2" pageNumber="21">M. mystacinus</emphasis>
</taxonomicName>
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sample in a number of characters.
</paragraph>
<paragraph id="090836A52A0BFF92FF76FE136D51FD54" blockId="2.[166,1225,175,1117]" pageId="2" pageNumber="21">
Almost the same can be said about the drawings of
<taxonomicName id="CEB74D262A0BFF92FCE9FE126F95FE11" authorityName="Kaup" authorityYear="1829" box="[836,915,387,412]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="genus">
<emphasis id="3BC3EAB72A0BFF92FCE9FE126F95FE11" box="[836,915,387,412]" italics="true" pageId="2" pageNumber="21">Myotis</emphasis>
</taxonomicName>
skulls (
<bibRefCitation id="6D264B542A0BFF92FC43FE136CD5FE4D" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." pageId="2" pageNumber="21" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al., 2012: 332</bibRefCitation>
, fig. 97) and teeth (
<bibRefCitation id="6D264B542A0BFF92FE11FE376ECAFE4D" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[444,716,422,448]" pageId="2" pageNumber="21" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al., 2012: 333</bibRefCitation>
, fig. 98).
<bibRefCitation id="6D264B542A0BFF92FC91FE37681BFE4D" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[828,1053,422,448]" pageId="2" pageNumber="21" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al. (2012)</bibRefCitation>
demonstrated four skull and teeth outlines: for
<taxonomicName id="CEB74D262A0BFF92FD98FE5B6EE7FE6E" baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[565,737,458,483]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" sensu="stricto" species="mystacinus">
<emphasis id="3BC3EAB72A0BFF92FD98FE5B6EE7FE6E" box="[565,737,458,483]" italics="true" pageId="2" pageNumber="21">M. mystacinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20F057CC2A0BFF92FD47FE586F2AFE6E" box="[746,812,457,483]" pageId="2" pageNumber="21" sensu="stricto">s. str.</taxonomicNameLabel>
, the
<typeStatus id="D60C88072A0BFF92FCC5FE586F9CFE6E" box="[872,922,457,483]" pageId="2" pageNumber="21">type</typeStatus>
specimen of
<taxonomicName id="CEB74D262A0BFF92FB90FE5B6CCDFD8B" authorityName="Benda, Reiter &amp; Vallo" authorityYear="2012" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="hyrcanicus">
<emphasis id="3BC3EAB72A0BFF92FB90FE5B6CCDFD8B" italics="true" pageId="2" pageNumber="21">M. hyrcanicus</emphasis>
</taxonomicName>
, the steppe bat from
<collectingCountry id="71A076352A0BFF92FE79FE7C6E00FD8A" box="[468,518,493,519]" name="Iran" pageId="2" pageNumber="21">Iran</collectingCountry>
(
<taxonomicName id="CEB74D262A0BFF92FDB7FE7C6E90FD8B" box="[538,662,492,518]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A0BFF92FDB7FE7C6E90FD8B" box="[538,662,492,518]" italics="true" pageId="2" pageNumber="21">M. davidii</emphasis>
</taxonomicName>
according to the classification of Benda et al.;
<taxonomicName id="CEB74D262A0BFF92FF0BFD816D38FDA4" authorityName="Kusjakin" authorityYear="1935" box="[166,318,528,553]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="aurascens">
<emphasis id="3BC3EAB72A0BFF92FF0BFD816D38FDA4" box="[166,318,528,553]" italics="true" pageId="2" pageNumber="21">M. aurascens</emphasis>
</taxonomicName>
according to the previous classification), and the
<typeStatus id="D60C88072A0BFF92FCD2FD816FB7FDA7" box="[895,945,528,554]" pageId="2" pageNumber="21">type</typeStatus>
specimen of
<taxonomicName id="CEB74D262A0BFF92FBE7FD8168C4FDA4" box="[1098,1218,527,553]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A0BFF92FBE7FD8168C4FDA4" box="[1098,1218,527,553]" italics="true" pageId="2" pageNumber="21">M. davidii</emphasis>
</taxonomicName>
. The sample is hardly sufficient: the skull is a diverse and variable structure, both divergent and convergent evolutionary shifts are possible in its size and shape, at least it is needed to compare the skulls of a larger number of OTUs and take into account the within-group variation, particularly geographical, in skull and teeth size and shape for each of the OTUs under analysis.
</paragraph>
<paragraph id="090836A52A0BFF92FF76FD736E5CFBD1" blockId="2.[166,1225,175,1117]" pageId="2" pageNumber="21">
<location id="0C68607E2A0BFF92FF76FD736D0FFD71" LSID="urn:lsid:plazi:treatment:811E87B32A08FF92FF4EFC296E5CFBD1:0C68607E2A0BFF92FF76FD736D0FFD71" box="[219,265,738,764]" country="Kazakhstan" county="Eastern" municipality="Western" name="The" pageId="2" pageNumber="21">The</location>
results of
<location id="0C68607E2A0BFF92FE20FD736DF3FD71" LSID="urn:lsid:plazi:treatment:811E87B32A08FF92FF4EFC296E5CFBD1:0C68607E2A0BFF92FE20FD736DF3FD71" box="[397,501,738,764]" country="Kazakhstan" county="Eastern" municipality="Western" name="Bayesian" pageId="2" pageNumber="21">Bayesian</location>
inference analysis among the haplotypes within the
<taxonomicName id="CEB74D262A0BFF92FBD4FD726D2EFC92" baseAuthorityName="Kuhl" baseAuthorityYear="1817" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="mystacinus">
<emphasis id="3BC3EAB72A0BFF92FBD4FD726D2EFC92" italics="true" pageId="2" pageNumber="21">Myotis mystacinus</emphasis>
</taxonomicName>
morpho-group (complete cytochrome b sequences) provided by
<bibRefCitation id="6D264B542A0BFF92FB96FC976CEAFCCE" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." pageId="2" pageNumber="21" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al. (2012</bibRefCitation>
: fig. 99) show that the three haplotypes from the specimens collected in
<location id="0C68607E2A0BFF92FB93FCB86891FCCE" LSID="urn:lsid:plazi:treatment:811E87B32A08FF92FF4EFC296E5CFBD1:0C68607E2A0BFF92FB93FCB86891FCCE" box="[1086,1175,809,835]" country="Kazakhstan" county="Eastern" municipality="Western" name="Eastern" pageId="2" pageNumber="21">Eastern</location>
<collectingCountry id="71A076352A0BFF92FB30FCB86D16FCEB" name="Kazakhstan" pageId="2" pageNumber="21">Kazakhstan</collectingCountry>
,
<location id="0C68607E2A0BFF92FEB2FCDC6D55FCEA" LSID="urn:lsid:plazi:treatment:811E87B32A08FF92FF4EFC296E5CFBD1:0C68607E2A0BFF92FEB2FCDC6D55FCEA" box="[287,339,845,871]" country="Kazakhstan" county="Eastern" municipality="Western" name="Tiva" pageId="2" pageNumber="21">Tıva</location>
and
<collectingCountry id="71A076352A0BFF92FE38FCDD6E2DFCEA" box="[405,555,844,871]" name="South Korea" pageId="2" pageNumber="21">South Korea</collectingCountry>
form a clade, separate from the other terminal taxa of
<taxonomicName id="CEB74D262A0BFF92FF0BFCE16EF7FC04" authority="sensu Benda &amp; Tsytsulina (2000)" authorityName="sensu Benda &amp; Tsytsulina" authorityYear="2000" box="[166,753,879,906]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="aurascens">
<emphasis id="3BC3EAB72A0BFF92FF0BFCE16D6AFC04" box="[166,364,880,905]" italics="true" pageId="2" pageNumber="21">Myotis aurascens</emphasis>
sensu
<bibRefCitation id="6D264B542A0BFF92FE14FCFE6EF7FC04" author="Benda, P. &amp; Tsytsulina, K. A." box="[441,753,879,905]" pageId="2" pageNumber="21" pagination="331 - 398" refId="ref5672" refString="Benda, P. &amp; Tsytsulina, K. A. 2000. Taxonomic revision of Myotis mystacinus group (Mammalia: Chiroptera) in the western Palearctic. Acta Societatis Zoologicae Bohemicae, 64 (4), 331 - 398." type="journal article" year="2000">Benda &amp; Tsytsulina (2000)</bibRefCitation>
</taxonomicName>
, but only with pairwise genetic distances varying between 3.66.6, which is less than half the distances between
<location id="0C68607E2A0BFF92FC4CFC03683EFC21" LSID="urn:lsid:plazi:treatment:811E87B32A08FF92FF4EFC296E5CFBD1:0C68607E2A0BFF92FC4CFC03683EFC21" box="[993,1080,914,940]" country="Kazakhstan" county="Eastern" municipality="Western" name="Iranian" pageId="2" pageNumber="21">Iranian</location>
<taxonomicName id="CEB74D262A0BFF92FB93FC026CC9FC42" baseAuthorityName="Kuhl" baseAuthorityYear="1817" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="mystacinus">
<emphasis id="3BC3EAB72A0BFF92FB93FC026CC9FC42" italics="true" pageId="2" pageNumber="21">M. mystacinus</emphasis>
</taxonomicName>
and
<location id="0C68607E2A0BFF92FEA6FC276D64FC5D" LSID="urn:lsid:plazi:treatment:811E87B32A08FF92FF4EFC296E5CFBD1:0C68607E2A0BFF92FEA6FC276D64FC5D" box="[267,354,950,976]" country="Kazakhstan" county="Eastern" municipality="Western" name="Iranian" pageId="2" pageNumber="21">Iranian</location>
<taxonomicName id="CEB74D262A0BFF92FEC4FC276F6FFC5D" authority="sensu Benda &amp; Tsytsulina" authorityName="sensu Benda &amp; Tsytsulina" authorityYear="2000" box="[361,873,950,976]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="aurascens">
<emphasis id="3BC3EAB72A0BFF92FEC4FC276E36FC42" box="[361,560,950,975]" italics="true" pageId="2" pageNumber="21">Myotis aurascens</emphasis>
sensu Benda &amp; Tsytsulina
</taxonomicName>
(
<bibRefCitation id="6D264B542A0BFF92FCD5FC276883FC5D" author="Benda, P &amp; Faizolahi, K. &amp; Andreas, M. &amp; Obuch, J. &amp; Reiter, A. &amp; Sevcik, M. &amp; Uhrin, M. &amp; Vallo, P. &amp; Ashrafi, S." box="[888,1157,950,976]" pageId="2" pageNumber="21" pagination="163 - 582" refId="ref5594" refString="Benda, P, Faizolahi, K., Andreas, M., Obuch, J., Reiter, A., Sevcik, M., Uhrin, M., Vallo, P. &amp; Ashrafi, S. 2012. Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 10. Bat fauna of Iran. Acta Societatis Zoologicae Bohemicae, 76, 163 - 582." type="journal article" year="2012">Benda et al., 2012: 337</bibRefCitation>
, table 18).
<location id="0C68607E2A0BFF92FF74FC486D0BFC7E" LSID="urn:lsid:plazi:treatment:811E87B32A08FF92FF4EFC296E5CFBD1:0C68607E2A0BFF92FF74FC486D0BFC7E" box="[217,269,985,1011]" country="Kazakhstan" county="Eastern" municipality="Western" name="This" pageId="2" pageNumber="21">This</location>
shows that differences between the
<collectingMunicipality id="E96CACDF2A0BFF92FD1DFC486F12FC7E" box="[688,788,985,1011]" pageId="2" pageNumber="21">Western</collectingMunicipality>
and the
<collectingCounty id="E0694E292A0BFF92FCD5FC486FD7FC7E" box="[888,977,985,1011]" pageId="2" pageNumber="21">Eastern</collectingCounty>
(to which the
<typeStatus id="D60C88072A0BFF92FBD4FC4868ADFC7E" box="[1145,1195,985,1011]" pageId="2" pageNumber="21">type</typeStatus>
of
<taxonomicName id="CEB74D262A0BFF92FF0BFC6C6D19FB9B" box="[166,287,1020,1046]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="davidii">
<emphasis id="3BC3EAB72A0BFF92FF0BFC6C6D19FB9B" box="[166,287,1020,1046]" italics="true" pageId="2" pageNumber="21">M. davidii</emphasis>
</taxonomicName>
is believed to belong) populations of
<taxonomicName id="CEB74D262A0BFF92FD7AFC6C6F98FB9B" authorityName="sensu Benda &amp; Tsytsulina" authorityYear="2000" box="[727,926,1021,1046]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="2" pageNumber="21" phylum="Chordata" rank="species" species="aurascens">
<emphasis id="3BC3EAB72A0BFF92FD7AFC6C6F98FB9B" box="[727,926,1021,1046]" italics="true" pageId="2" pageNumber="21">Myotis aurascens</emphasis>
</taxonomicName>
are of at most subspecies level. Later, such a subdivision was generally confirmed and substantiated by
<bibRefCitation id="6D264B542A0BFF92FB91FB8E6D0BFBD0" author="Coraman, E &amp; Dundarova, H. &amp; Dietz, C. &amp; Mayer, F." pageId="2" pageNumber="21" pagination="1 - 13" refId="ref5715" refString="Coraman, E, Dundarova, H., Dietz, C. &amp; Mayer, F. 2020. Patterns of mtDNA introgression suggest population replacement in Palaearctic whiskered bat species. Royal Society Open Science, 7: article No. 191805, 1 - 13, http: // dx. doi. org / 10.1098 / rsos. 191805" type="journal article" year="2020">Çoraman et al. (2020</bibRefCitation>
; see below for more details).
</paragraph>
</subSubSection>
</treatment>
</document>