171 lines
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171 lines
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<document id="54D24B82828BCA544FF904B39D1EE9CD" ID-CLB-Dataset="30325" ID-DOI="10.11646/zootaxa.4405.1.1" ID-GBIF-Dataset="55df350b-41c6-4f2e-9a18-c0be6d47498b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3097993" ID-ZooBank="D33BF380-5AF7-4645-86C7-9981C528EAF0" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1522988054869" checkinUser="plazi" docAuthor="Alitto, Renata A. S., Bueno, Maristela L., Guilherme, Pablo D. B., Domenico, Maikon Di, Christensen, Ana Beardsley & Borges, Michela" docDate="2018" docId="03C3B82F9201C95507C8FD86FDFD3A68" docLanguage="en" docName="zootaxa.4405.1.1.pdf.imf" docOrigin="Zootaxa 4405 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hemipholis Lyman 1865" docType="treatment" docVersion="10" lastPageNumber="46" masterDocId="FFFAC057922CC978075FFFCEFF883E24" masterDocTitle="Shallow-water brittle stars (Echinodermata: Ophiuroidea) from Araçá Bay (Southeastern Brazil), with spatial distribution considerations" masterLastPageNumber="66" masterPageNumber="1" pageNumber="46" updateTime="1698535905832" updateUser="ExternalLinkService">
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<mods:title id="7C05358CC3A2D9AC418D6B455B77AD9C">Shallow-water brittle stars (Echinodermata: Ophiuroidea) from Araçá Bay (Southeastern Brazil), with spatial distribution considerations</mods:title>
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<mods:namePart id="F27547121C4CE45E9D5A8B4AE3936D92">Alitto, Renata A. S.</mods:namePart>
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<mods:namePart id="1C25A34F070591616EF3F70D8CCF0FFC">Bueno, Maristela L.</mods:namePart>
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<mods:namePart id="02B1703E6FEA7809A8776A78833209DF">Guilherme, Pablo D. B.</mods:namePart>
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<mods:namePart id="CD675CF7C2CBF600F966CF131D1F5023">Domenico, Maikon Di</mods:namePart>
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<mods:namePart id="E2AB17CEC39FF7375CB4B249816A6CAB">Christensen, Ana Beardsley</mods:namePart>
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<mods:namePart id="06677D772C049EDA73CE04AD62768461">Borges, Michela</mods:namePart>
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<mods:title id="F56F01248DB2513D2EA4A2344E4BCE49">Zootaxa</mods:title>
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<mods:date id="47049A2F808402882D3216F13DD71851">2018</mods:date>
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<mods:number id="FDEECDABF9FC9ED440B5238013B4FB99">2018-04-06</mods:number>
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<mods:number id="3A4A0A0D4C20FE2AE21F115F6402179A">4405</mods:number>
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<mods:identifier id="F5BC04BFAA9F73DA2D2A911C93B1E543" type="DOI">10.11646/zootaxa.4405.1.1</mods:identifier>
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<treatment id="03C3B82F9201C95507C8FD86FDFD3A68" ID-DOI="http://doi.org/10.5281/zenodo.5985160" ID-GBIF-Taxon="143478636" ID-Zenodo-Dep="5985160" LSID="urn:lsid:plazi:treatment:03C3B82F9201C95507C8FD86FDFD3A68" httpUri="http://treatment.plazi.org/id/03C3B82F9201C95507C8FD86FDFD3A68" lastPageNumber="46" pageId="45" pageNumber="46">
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<subSubSection id="C3705AB29201C95507C8FD86FDAB3C46" box="[151,547,583,610]" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="8BD509399201C95507C8FD86FDAB3C46" blockId="45.[151,547,583,610]" box="[151,547,583,610]" pageId="45" pageNumber="46">
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<heading id="D09DBE559201C95507C8FD86FDAB3C46" bold="true" box="[151,547,583,610]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
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<emphasis id="B91ED52B9201C95507C8FD86FDAB3C46" bold="true" box="[151,547,583,610]" pageId="45" pageNumber="46">
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Genus
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<taxonomicName id="4C6A72BA9201C95507B1FD89FDAB3C46" authority="Lyman, 1865" authorityName="Lyman" authorityYear="1865" box="[238,547,583,610]" class="Ophiuroidea" family="Ophiactidae" genus="Hemipholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="45" pageNumber="46" phylum="Echinodermata" rank="genus">
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<emphasis id="B91ED52B9201C95507B1FD89FEF23C45" bold="true" box="[238,378,583,609]" italics="true" pageId="45" pageNumber="46">Hemipholis</emphasis>
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<bibRefCitation id="EFFB74C89201C95506DEFD86FDAB3C46" author="Lyman, T." bookContentInfo="203 pp." box="[385,547,584,610]" journalOrPublisher="University Press: Welch, Bigelow, & Company, Cambridge" pageId="45" pageNumber="46" refId="ref50591" refString="Lyman, T. (1865) Ophiuridae and Astrophytidae. University Press: Welch, Bigelow, & Company, Cambridge, 203 pp." title="Ophiuridae and Astrophytidae" type="book" year="1865">Lyman, 1865</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3705AB29201C95507C8FD40FBB93C80" box="[151,1073,654,676]" pageId="45" pageNumber="46" type="materials_examined">
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<paragraph id="8BD509399201C95507C8FD40FBB93C80" blockId="45.[151,1073,654,676]" box="[151,1073,654,676]" pageId="45" pageNumber="46">
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<emphasis id="B91ED52B9201C95507C8FD40FE9C3C80" bold="true" box="[151,276,654,676]" pageId="45" pageNumber="46">Type taxon.</emphasis>
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<taxonomicName id="4C6A72BA9201C9550644FD40FDF63C80" authority="(Bosc, 1802)" baseAuthorityName="Bosc" baseAuthorityYear="1802" box="[283,638,654,676]" class="Ophiuroidea" family="Ophiactidae" genus="Hemipholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="45" pageNumber="46" phylum="Echinodermata" rank="species" species="cordifera">
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<emphasis id="B91ED52B9201C9550644FD40FE7B3C80" box="[283,499,654,676]" italics="true" pageId="45" pageNumber="46">Hemipholis cordifera</emphasis>
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(
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<bibRefCitation id="EFFB74C89201C955055EFD40FDF03C80" author="Bosc, L. A. G." bookContentInfo="300 pp." box="[513,632,654,676]" journalOrPublisher="Chez Deterville, de l'imprimerie de Guilleminet, Paris" pageId="45" pageNumber="46" refId="ref45280" refString="Bosc, L. A. G. (1802) Histoire naturelle des vers, contenant leur description et leurs moeurs; avec figures dessinees d'apres nature. Chez Deterville, de l'imprimerie de Guilleminet, Paris, 300 pp." title="Histoire naturelle des vers, contenant leur description et leurs moeurs; avec figures dessinees d'apres nature" type="book" year="1802">Bosc, 1802</bibRefCitation>
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)
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</taxonomicName>
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, originally described as
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<taxonomicName id="4C6A72BA9201C955042AFD41FBA33C80" authorityName="Bosc" authorityYear="1802" box="[885,1067,654,676]" class="Asteroidea" family="Asteriidae" genus="Asterias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Forcipulatida" pageId="45" pageNumber="46" phylum="Echinodermata" rank="species" species="cordifera">
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<emphasis id="B91ED52B9201C955042AFD41FBA33C80" box="[885,1067,654,676]" italics="true" pageId="45" pageNumber="46">Asterias cordifera</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3705AB29201C95507C8FD02FE483D74" pageId="45" pageNumber="46" type="diagnosis">
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<paragraph id="8BD509399201C95507C8FD02FE483D74" blockId="45.[151,1437,716,1100]" pageId="45" pageNumber="46">
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<emphasis id="B91ED52B9201C95507C8FD02FE993CC1" bold="true" box="[151,273,716,741]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
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Disc regularly covered with scales dorsally and naked ventrally, with soft interradial recesses. Absence of bursae. Radial shields partially joined. One lateral oral papilla. Adoral shields all in contact, forming a continuous ring around the mouth. One tentacle scale. Three pointed arm spines (
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<bibRefCitation id="EFFB74C89201C9550340FCDAFB383D09" author="Lyman, T." bookContentInfo="203 pp." box="[1055,1200,788,813]" journalOrPublisher="University Press: Welch, Bigelow, & Company, Cambridge" pageId="45" pageNumber="46" refId="ref50591" refString="Lyman, T. (1865) Ophiuridae and Astrophytidae. University Press: Welch, Bigelow, & Company, Cambridge, 203 pp." title="Ophiuridae and Astrophytidae" type="book" year="1865">Lyman 1865</bibRefCitation>
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;
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<bibRefCitation id="EFFB74C89201C95503E3FCDAFAA13D09" author="Fell, H. B." box="[1212,1321,788,813]" journalOrPublisher="Zoology Publications from Victoria University of Wellington" pageId="45" pageNumber="46" pagination="1 - 44" part="26" refId="ref47348" refString="Fell, H. B. (1960) Synoptic keys to the genera of Ophiuroidea. Zoology Publications from Victoria University of Wellington, 26, 1 - 44." title="Synoptic keys to the genera of Ophiuroidea" type="journal article" year="1960">Fell 1960</bibRefCitation>
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;
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<bibRefCitation id="EFFB74C89201C955026AFCDAFF5B3D74" author="Tommasi, L. R." journalOrPublisher="Contribuicoes Avulsas do Instituto Oceanografico, Universidade de Sao Paulo" pageId="45" pageNumber="46" pagination="1 - 146" part="20" refId="ref54902" refString="Tommasi, L. R. (1970) Os ofiuroides recentes do Brasil e de regioes vizinhas. Contribuicoes Avulsas do Instituto Oceanografico, Universidade de Sao Paulo, 20, 1 - 146." title="Os ofiuroides recentes do Brasil e de regioes vizinhas" type="journal article" year="1970">Tommasi 1970</bibRefCitation>
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;
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<bibRefCitation id="EFFB74C89201C9550782FCF9FE3B3D74" author="Albuquerque, M. N." bookContentInfo="409 pp." box="[221,435,823,848]" journalOrPublisher="Universidade de Sao Paulo, Sao Paulo" pageId="45" pageNumber="46" refId="ref43538" refString="Albuquerque, M. N. (1986) Ophiuroidea Gray, 1840 (Echinodermata) da plataforma continental do norte e nordeste brasileiro. Phd thesis, Universidade de Sao Paulo, Sao Paulo, 409 pp." title="Ophiuroidea Gray, 1840 (Echinodermata) da plataforma continental do norte e nordeste brasileiro" type="book" year="1986">Albuquerque 1986</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3705AB29201C9550798FC92FDFD3A68" pageId="45" pageNumber="46" type="discussion">
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<paragraph id="8BD509399201C9550798FC92FDFD3A68" blockId="45.[151,1437,716,1100]" pageId="45" pageNumber="46">
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<emphasis id="B91ED52B9201C9550798FC92FEC73D51" bold="true" box="[199,335,860,885]" pageId="45" pageNumber="46">Comments.</emphasis>
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<taxonomicName id="4C6A72BA9201C9550606FC92FE523D50" authorityName="Lyman" authorityYear="1865" box="[345,474,860,884]" class="Ophiuroidea" family="Ophiactidae" genus="Hemipholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="45" pageNumber="46" phylum="Echinodermata" rank="genus">
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<emphasis id="B91ED52B9201C9550606FC92FE523D50" box="[345,474,860,884]" italics="true" pageId="45" pageNumber="46">Hemipholis</emphasis>
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</taxonomicName>
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shares features with
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<emphasis id="B91ED52B9201C955058EFC92FCCD3D50" box="[721,837,860,884]" italics="true" pageId="45" pageNumber="46">
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<taxonomicName id="4C6A72BA9201C955058EFC92FCC93D50" authorityName="Lutken" authorityYear="1856" box="[721,833,860,884]" class="Ophiuroidea" family="Ophiotrichidae" genus="Ophiactis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="45" pageNumber="46" phylum="Echinodermata" rank="genus">Ophiactis</taxonomicName>
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,
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</emphasis>
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such as a ring of contiguous adoral shields and the presence of hemoglobin containing coelomocytes in its water vascular system (
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<bibRefCitation id="EFFB74C89201C9550355FCB1FA993DBC" author="Christensen, A. B. & Christensen, E. F. & Weisrock, D. W." box="[1034,1297,895,920]" journalOrPublisher="Manine Biology" pageId="45" pageNumber="46" pagination="755 - 763" part="154" refId="ref45851" refString="Christensen, A. B., Christensen, E. F. & Weisrock, D. W. (2008) Population genetic structure of North American Ophiactis spp. brittle stars possessing hemoglobin. Manine Biology, 154, 755 - 763. https: // doi. org / 10.1007 / s 00227 - 008 - 0968 - 1" title="Population genetic structure of North American Ophiactis spp. brittle stars possessing hemoglobin" type="journal article" year="2008">
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Christensen
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<emphasis id="B91ED52B9201C95503C8FC4FFB403DBC" box="[1175,1224,896,920]" italics="true" pageId="45" pageNumber="46">et al</emphasis>
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. 2008
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</bibRefCitation>
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), suggesting a close systematic relationship between both genera (
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<bibRefCitation id="EFFB74C89201C95505B7FC6AFC0F3D98" author="Hendler, G." box="[744,903,932,957]" journalOrPublisher="Zootaxa" pageId="45" pageNumber="46" pagination="44 - 52" part="3048" refId="ref48483" refString="Hendler, G. (2011) New light on the nomenclature, taxonomy, and biology of Hemipholis species (Echinodermata: Ophiuroidea: Ophiactidae). Zootaxa, 3048, 44 - 52." title="New light on the nomenclature, taxonomy, and biology of Hemipholis species (Echinodermata: Ophiuroidea: Ophiactidae)" type="journal article" year="2011">Hendler 2011</bibRefCitation>
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).
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<taxonomicName id="4C6A72BA9201C95504C6FC6AFB923D98" authorityName="Lyman" authorityYear="1865" box="[921,1050,932,956]" class="Ophiuroidea" family="Ophiactidae" genus="Hemipholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="45" pageNumber="46" phylum="Echinodermata" rank="genus">
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<emphasis id="B91ED52B9201C95504C6FC6AFB923D98" box="[921,1050,932,956]" italics="true" pageId="45" pageNumber="46">Hemipholis</emphasis>
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</taxonomicName>
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is well-studied and has repeatedly been cited in ecological, environmental, and biological studies owing to its abundance, broad geographic range, and its remarkable morphological and physiological specializations (
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<bibRefCitation id="EFFB74C89201C95504CAFC22FBF93A21" author="Hendler, G. & Miller, J. E. & Pawson, D. L. & Kier, P. M." bookContentInfo="390 pp." box="[917,1137,1004,1029]" journalOrPublisher="Smithsonian Institution Press, Washington" pageId="45" pageNumber="46" refId="ref48691" refString="Hendler, G., Miller, J. E., Pawson, D. L. & Kier, P. M. (1995) Sea stars, sea urchins, and allies: echinoderms of Florida and the Caribbean. Smithsonian Institution Press, Washington, 390 pp. https: // doi. org / 10.1017 / s 0025315400031568" title="Sea stars, sea urchins, and allies: echinoderms of Florida and the Caribbean" type="book" year="1995">
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Hendler
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<emphasis id="B91ED52B9201C95504A7FC23FBA03A20" box="[1016,1064,1004,1028]" italics="true" pageId="45" pageNumber="46">et al</emphasis>
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. 1995
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</bibRefCitation>
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;
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<bibRefCitation id="EFFB74C89201C9550323FC22FA913A20" author="Hendler, G." box="[1148,1305,1004,1029]" journalOrPublisher="Zootaxa" pageId="45" pageNumber="46" pagination="44 - 52" part="3048" refId="ref48483" refString="Hendler, G. (2011) New light on the nomenclature, taxonomy, and biology of Hemipholis species (Echinodermata: Ophiuroidea: Ophiactidae). Zootaxa, 3048, 44 - 52." title="New light on the nomenclature, taxonomy, and biology of Hemipholis species (Echinodermata: Ophiuroidea: Ophiactidae)" type="journal article" year="2011">Hendler 2011</bibRefCitation>
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). Currently, two species are accepted in
|
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<taxonomicName id="4C6A72BA9201C9550680FBDEFCCB3A0C" authority="(Stohr et al. 2016)" authorityName="Stohr et al." authorityYear="2016" box="[479,835,1039,1064]" class="Ophiuroidea" family="Ophiactidae" genus="Hemipholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="45" pageNumber="46" phylum="Echinodermata" rank="genus">
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<emphasis id="B91ED52B9201C9550680FBDEFDE83A0C" box="[479,608,1040,1064]" italics="true" pageId="45" pageNumber="46">Hemipholis</emphasis>
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(
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<bibRefCitation id="EFFB74C89201C955052EFBC1FCB33A0C" author="Stohr, S. & Martynov, A." box="[625,827,1039,1064]" journalOrPublisher="PLoS ONE" pageId="45" pageNumber="46" pagination="0164562" part="11" refId="ref54292" refString="Stohr, S. & Martynov, A. (2016) Paedomorphosis as an Evolutionary Driving Force: Insights from Deep-Sea Brittle Stars. PLoS ONE, 11, e 0164562. https: // doi. org / 10.1371 / journal. pone. 0164562" title="Paedomorphosis as an Evolutionary Driving Force: Insights from Deep-Sea Brittle Stars" type="journal article" year="2016">
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Stöhr
|
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<emphasis id="B91ED52B9201C95505E5FBDFFD663A0C" box="[698,750,1040,1064]" italics="true" pageId="45" pageNumber="46">et al</emphasis>
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. 2016
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</bibRefCitation>
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)
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</taxonomicName>
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, with one recorded from Brazil (
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<bibRefCitation id="EFFB74C89201C955039BFBC1FF593A68" author="Barboza, C. A. M. & Borges, M." journalOrPublisher="Zootaxa" pageId="45" pageNumber="46" pagination="1 - 21" part="3447" refId="ref44247" refString="Barboza, C. A. M. & Borges, M. (2012) A checklist of the extant species of ophiuroids (Echinodermata: Ophiuroidea) from Brazilian waters. Zootaxa, 3447, 1 - 21." title="A checklist of the extant species of ophiuroids (Echinodermata: Ophiuroidea) from Brazilian waters" type="journal article" year="2012">Barboza & Borges 2012</bibRefCitation>
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):
|
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<taxonomicName id="4C6A72BA9201C95507B8FBFAFDF93A68" authority="(Bosc, 1802)" baseAuthorityName="Bosc" baseAuthorityYear="1802" box="[231,625,1076,1100]" class="Ophiuroidea" family="Ophiactidae" genus="Hemipholis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="45" pageNumber="46" phylum="Echinodermata" rank="species" species="cordifera">
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<emphasis id="B91ED52B9201C95507B8FBFAFE5E3A68" box="[231,470,1076,1100]" italics="true" pageId="45" pageNumber="46">Hemipholis cordifera</emphasis>
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(
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<bibRefCitation id="EFFB74C89201C95506BAFBFAFDE03A68" author="Bosc, L. A. G." bookContentInfo="300 pp." box="[485,616,1076,1100]" journalOrPublisher="Chez Deterville, de l'imprimerie de Guilleminet, Paris" pageId="45" pageNumber="46" refId="ref45280" refString="Bosc, L. A. G. (1802) Histoire naturelle des vers, contenant leur description et leurs moeurs; avec figures dessinees d'apres nature. Chez Deterville, de l'imprimerie de Guilleminet, Paris, 300 pp." title="Histoire naturelle des vers, contenant leur description et leurs moeurs; avec figures dessinees d'apres nature" type="book" year="1802">Bosc, 1802</bibRefCitation>
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)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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