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<document id="3684A46695CBD157E3555507913538DD" ID-CLB-Dataset="8579" ID-DOI="10.11646/zootaxa.4850.1.1" ID-GBIF-Dataset="686337a2-1528-43d0-ae32-68ca023d8df3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4407072" ID-ZooBank="18AA0411-0E54-4922-84C7-608EAC68D281" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609428297648" checkinUser="plazi" docAuthor="Guéorguiev, Borislav &amp; Liang, Hongbin" docDate="2020" docId="03BC5E5B2960FF9DFF4BFF23ED45FE74" docLanguage="en" docName="zootaxa.4850.1.1.pdf" docOrigin="Zootaxa 4850 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pseudoodes rambouseki Guéorguiev &amp; Liang 2020, comb. n." docType="treatment" docVersion="9" lastPageNumber="35" masterDocId="FF8526232940FFBFFFDCFFB4EC2EFFFD" masterDocTitle="Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species" masterLastPageNumber="89" masterPageNumber="1" pageNumber="33" updateTime="1698852686469" updateUser="ExternalLinkService">
<mods:mods id="DF7BF440DBF67C9D76B3F3525DD52467" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9CFF364B51ABED0EDDB0A2425E79A793">Revision of the Palaearctic and Oriental representatives of Lachnocrepis LeConte and Oodes Bonelli (Coleoptera: Carabidae), with special account on Chinese species</mods:title>
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<mods:name id="6A9B72A12BD5FFE693D5B4865D1D75EB" type="personal">
<mods:role id="05F7F53F85D3BA271678F392986ED4CE">
<mods:roleTerm id="B880E2BA5538BEEE8F11ED9115DDFF91">Author</mods:roleTerm>
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<mods:namePart id="65CEA978C6192F5BA77B7CBE0C8A175C">Guéorguiev, Borislav</mods:namePart>
<mods:nameIdentifier id="0F6331DAA8394C8564CE3A3EBE6FCB8C" type="ORCID">0000-0002-8532-0657</mods:nameIdentifier>
<mods:nameIdentifier id="1487966338F8D270AF62304CB53868AB" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E5534F23C11C1E2165C8E920F1742F12" type="personal">
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<mods:namePart id="2641280736672158E3FFA69D2CB0E5DC">Liang, Hongbin</mods:namePart>
<mods:nameIdentifier id="FEC05EC1758A10CA3ADEFCED1B73767B" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
<mods:nameIdentifier id="AFE19B89397A7D85E9CA4C138374A58A" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:typeOfResource id="B8F04F1B633A8BEC8178BD076E3D8D2D">text</mods:typeOfResource>
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<mods:title id="06FEB021DF68AC3E5E563CC45BE183FD">Zootaxa</mods:title>
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<mods:part id="64E86B246162E17838C3B5ACF5ECACF5">
<mods:date id="EC9F9B359881DF517438E7CABEE38933">2020</mods:date>
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<mods:number id="6F1675118D5E49F703AA377FD19FD2F1">2020-09-08</mods:number>
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<mods:detail id="3886733271AE5B40C58CB6F3C95D73C5" type="volume">
<mods:number id="F89D3F2F4FCEA42436980B507F063F1F">4850</mods:number>
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<mods:classification id="D710C10955C7D8AF7A745A67917CECD3">journal article</mods:classification>
<mods:identifier id="9455D44E3F3F6B62AC4DE4DC74AA9550" type="CLB-Dataset">8579</mods:identifier>
<mods:identifier id="68EDEA7DF5EA40EEB132F949AF7017E6" type="DOI">10.11646/zootaxa.4850.1.1</mods:identifier>
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<mods:identifier id="5C3326A514F6874FE232A21DC05706B5" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BC5E5B2960FF9DFF4BFF23ED45FE74" ID-DOI="http://doi.org/10.5281/zenodo.4480007" ID-GBIF-Taxon="177139750" ID-Zenodo-Dep="4480007" LSID="urn:lsid:plazi:treatment:03BC5E5B2960FF9DFF4BFF23ED45FE74" httpUri="http://treatment.plazi.org/id/03BC5E5B2960FF9DFF4BFF23ED45FE74" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
<subSubSection id="C30FBCC62960FF9FFF4BFF23EF30FF4C" box="[151,798,151,178]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BAAEF4D2960FF9FFF4BFF23EF30FF4C" blockId="32.[151,798,151,213]" box="[151,798,151,178]" pageId="32" pageNumber="33">
<emphasis id="B961335F2960FF9FFF4BFF23EF30FF4C" bold="true" box="[151,798,151,178]" pageId="32" pageNumber="33">
9.
<taxonomicName id="4C1594CE2960FF9FFF69FF23EE8BFF4C" ID-CoL="8MGH5" authority="(Jedlicka, 1931)" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[181,677,151,178]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rambouseki" status="comb. nov.">
<emphasis id="B961335F2960FF9FFF69FF23EDF6FF4F" bold="true" box="[181,472,151,178]" italics="true" pageId="32" pageNumber="33">Pseudoodes rambouseki</emphasis>
(
<bibRefCitation id="EF8492BC2960FF9FFE3BFF23EEB2FF4C" author="Jedlicka, A." box="[487,668,151,177]" pageId="32" pageNumber="33" pagination="21 - 30" refId="ref62314" refString="Jedlicka, A. (1931) Novi Carabidi ze Secuanu v Cine. (Col.) Neue Carabiden aus China-Szetschuan. Casopis Ceskoslovenske Spolecnosti Entomologicke, 28, 21 - 30." type="journal article" year="1931">Jedlička, 1931</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2528E242960FF9FFD6FFF23EF30FF4C" box="[691,798,151,177]" pageId="32" pageNumber="33" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62960FF9FFF4BFF0FEDA5FF28" box="[151,395,187,213]" pageId="32" pageNumber="33" type="description">
<paragraph id="8BAAEF4D2960FF9FFF4BFF0FEDA5FF28" blockId="32.[151,798,151,213]" box="[151,395,187,213]" pageId="32" pageNumber="33">
(
<figureCitation id="132EF3C82960FF9FFF43FF0FECDDFF28" box="[159,243,187,213]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Figs 12</figureCitation>
A–J,
<tableCitation id="C697DAF62960FF9FFEF2FF0FEDACFF28" box="[302,386,187,213]" captionStart="TABLE 2" captionStartId="36.[151,239,646,672]" captionText="TABLE 2. Morphometric data for species of the genus Pseudoodes gen. n." pageId="32" pageNumber="33">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62960FF9FFF4BFEB4E888FEE5" box="[151,1190,256,280]" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="8BAAEF4D2960FF9FFF4BFEB4E888FEE5" blockId="32.[151,1190,256,280]" box="[151,1190,256,280]" pageId="32" pageNumber="33">
<taxonomicName id="4C1594CE2960FF9FFF4BFEB4EE15FEE5" ID-CoL="8LGGX" authority="Jedlicka, 1931: 21" authorityName="Jedlicka. Consists" authorityPageNumber="21" authorityYear="1931" box="[151,571,256,280]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2960FF9FFF4BFEB4ED52FEE5" box="[151,380,256,280]" italics="true" pageId="32" pageNumber="33">Holosoma rambouseki</emphasis>
<bibRefCitation id="EF8492BC2960FF9FFE5EFEB4EE15FEE5" author="Jedlicka, A." box="[386,571,256,280]" pageId="32" pageNumber="33" pagination="21 - 30" refId="ref62314" refString="Jedlicka, A. (1931) Novi Carabidi ze Secuanu v Cine. (Col.) Neue Carabiden aus China-Szetschuan. Casopis Ceskoslovenske Spolecnosti Entomologicke, 28, 21 - 30." type="journal article" year="1931">Jedlička, 1931: 21</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54AE51EF2960FF9FFD95FEB4EE58FEE5" box="[585,630,256,280]" pageId="32" pageNumber="33">type</typeStatus>
locality: ‘
<collectingCountry id="F302AFDD2960FF9FFD05FEB4EF34FEE5" box="[729,794,256,280]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
mer.: Prov. Szetschuan: Chung-king’).
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62960FF9FFF4BFE89ED3DFEA9" box="[151,275,317,341]" pageId="32" pageNumber="33" type="multiple">
<paragraph id="8BAAEF4D2960FF9FFF4BFE89ED3DFEA9" blockId="32.[151,1437,317,402]" box="[151,275,317,341]" pageId="32" pageNumber="33">
<heading id="D0E258212960FF9FFF4BFE89ED3DFEA9" bold="true" box="[151,275,317,341]" fontSize="10" level="2" pageId="32" pageNumber="33" reason="2">
<emphasis id="B961335F2960FF9FFF4BFE89ED23FEA8" bold="true" box="[151,269,317,341]" pageId="32" pageNumber="33">References</emphasis>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62960FF9FFF4BFEE8EDE0FE6F" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="8BAAEF4D2960FF9FFF4BFEE8EDE0FE6F" blockId="32.[151,1437,317,402]" pageId="32" pageNumber="33">
<taxonomicName id="4C1594CE2960FF9FFF4BFEE8EE11FE8E" ID-CoL="8LGGX" authority=": Csiki 1933: 1675" authorityName="Csiki" authorityPageNumber="1675" authorityYear="1933" box="[151,575,347,372]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2960FF9FFF4BFEE8ED53FE89" box="[151,381,348,372]" italics="true" pageId="32" pageNumber="33">Holosoma rambouseki</emphasis>
:
<bibRefCitation id="EF8492BC2960FF9FFE57FEE8EE11FE8E" author="Csiki, E." box="[395,575,347,372]" pageId="32" pageNumber="33" pagination="1599 - 1933" refId="ref60647" refString="Csiki, E. (1933) Carabidae: Harpalinae VIII. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum catalogus. Pars 126. W. Junk, Berlin, pp. 1599 - 1933." type="book chapter" year="1933">Csiki 1933: 1675</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EF8492BC2960FF9FFD95FEE8EEF5FE89" author="Wu, C. F." box="[585,731,348,372]" pageId="32" pageNumber="33" refId="ref64510" refString="Wu, C. F. (1937) Catalogus insectorum sinensium (Catalogue of Chinese insects). Vol. III. The Fan Memorial Institute of Biology, Peiping (Beijing), 1312 pp." type="book" year="1937">Wu 1937: 148</bibRefCitation>
(‘Szechnan’);
<bibRefCitation id="EF8492BC2960FF9FFCAFFEE8E802FE89" author="Lorenz, W." box="[883,1068,348,372]" pageId="32" pageNumber="33" refId="ref63123" refString="Lorenz, W. (1998) Systematic list of extant ground beetles of the world (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 1 st Edition. Published by the Author, Tutzing, [ii] + 502 pp." type="book" year="1998">Lorenz 1998: 304</bibRefCitation>
;
<bibRefCitation id="EF8492BC2960FF9FFBEBFEE8E926FE89" author="Bousquet, Y." box="[1079,1288,348,372]" pageId="32" pageNumber="33" pagination="1851" refId="ref60278" refString="Bousquet, Y. (2003) Tribe Oodini LaFerte-Senectere, 1851. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 444 - 445." type="book chapter" year="2003">Bousquet 2003: 444</bibRefCitation>
;
<bibRefCitation id="EF8492BC2960FF9FFACFFEE8ECDEFE6F" author="Lorenz, W." pageId="32" pageNumber="33" refId="ref63176" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world (Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, [ii] + 530 pp." type="book" year="2005">Lorenz 2005: 324</bibRefCitation>
;
<bibRefCitation id="EF8492BC2960FF9FFF27FECFEDE4FE6F" author="Bousquet, Y." box="[251,458,378,402]" pageId="32" pageNumber="33" pagination="1851" refId="ref60329" refString="Bousquet, Y. (2017) Tribe Oodini LaFerte-Senectere, 1851. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden and Boston, pp. 635 - 636." type="book chapter" year="2017">Bousquet 2017: 497</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62960FF9FFF4BFE75EE70FC0A" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BAAEF4D2960FF9FFF4BFE75EFBDFCE2" blockId="32.[151,1437,448,1987]" pageId="32" pageNumber="33">
<emphasis id="B961335F2960FF9FFF4BFE75ED6CFE27" bold="true" box="[151,322,448,475]" pageId="32" pageNumber="33">Type material</emphasis>
.
<taxonomicName id="4C1594CE2960FF9FFE88FE74EF03FE27" authority="Jedlicka. Consists" authorityName="Jedlicka. Consists" authorityYear="1931" box="[340,813,448,474]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2960FF9FFE88FE74EE7BFE27" box="[340,597,448,474]" italics="true" pageId="32" pageNumber="33">Holosoma rambouseki</emphasis>
Jedlička. Consists
</taxonomicName>
of
<specimenCount id="9D1324C42960FF9FFC86FE75EFBAFE26" box="[858,916,449,475]" pageId="32" pageNumber="33" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D1324C42960FF9FFC78FE75EFE9FE26" box="[932,967,449,475]" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
specimens, all preserved in NMPC. No specimen from the type series was explicitly designated as type or
<typeStatus id="54AE51EF2960FF9FFCA3FE50EFCFFE03" box="[895,993,484,510]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
in Jedlička’s work, thus all the samples are treated as
<typeStatus id="54AE51EF2960FF9FFEF0FDBDEDA0FDDE" box="[300,398,521,547]" pageId="32" pageNumber="33" type="syntype">syntypes</typeStatus>
(cf.Article 73.2.,
<bibRefCitation id="EF8492BC2960FF9FFD88FDBDEEF9FDDE" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[596,727,521,547]" pageId="32" pageNumber="33" refId="ref62087" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code on Zoological Nomenclature. 4 th Edition. Adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London. Available from: https: // www. iczn. org / the-code / the-international-code-of-zoological-nomenclature / the-codeonline / (accessed 24 December 2019)" type="url" year="1999">ICZN 1999</bibRefCitation>
). Of these specimens, a male is chosen for
<typeStatus id="54AE51EF2960FF9FFB74FDBCE921FDDF" box="[1192,1295,520,546]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
designation. It is labelled as follows: ‘CHUNG–KING Szetschuan [‘e’ almost illegible] Coll. Dr. Becker [w, p] // Rambouseki mihi det. Ing. Jedlička [w, h/p] // TYPE [r, with black frame, p] //
<typeStatus id="54AE51EF2960FF9FFCA6FDE5EFC6FD96" box="[890,1000,593,619]" pageId="32" pageNumber="33" type="paratype">Paratypus</typeStatus>
[r, p]’. The remaining three specimens are treated as
<typeStatus id="54AE51EF2960FF9FFEE2FDC0EDCAFD73" box="[318,484,628,654]" pageId="32" pageNumber="33" type="paralectotype">paralectotypes</typeStatus>
:
<specimenCount id="9D1324C42960FF9FFE2EFDC1EE38FD72" box="[498,534,629,655]" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
, ‘CHUNG–KING Szetschuan [‘e’ almost illegible] Coll. Dr. Becker [w, p] // TYPE [r, with black frame, p] //
<taxonomicName id="4C1594CE2960FF9FFE23FD2CEF2AFD4F" authority="Rambouseki" authorityName="Rambouseki" box="[511,772,664,690]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Holosoma Rambouseki</taxonomicName>
mihi DET.ING.JEDLIČKA [p, h/p]’;
<specimenCount id="9D1324C42960FF9FFB77FD2DE8E1FD4E" box="[1195,1231,665,691]" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
, ‘CHUNG–KING Szetschuan [‘e’ almost illegible] Coll. Dr. Becker [w, p] // TYPE [r, with black frame, p] // Rambouseki mihi det. Ing. Jedlička [w, h/p]’;
<specimenCount id="9D1324C42960FF9FFE7DFD55EDEAFD06" box="[417,452,737,763]" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
, ‘CHUNG–KING Szetschuan [‘e’ almost illegible] Coll. Dr. Becker [w, p] // TYPE [r, with black frame, p] // Rambouseki mihi det. Ing. Jedlička [w, h/p]’.
</paragraph>
<paragraph id="8BAAEF4D2960FF9FFF1BFC9CE941FC2E" blockId="32.[151,1437,448,1987]" pageId="32" pageNumber="33">
<emphasis id="B961335F2960FF9FFF1BFC9CEDD5FCBF" bold="true" box="[199,507,808,834]" pageId="32" pageNumber="33">Other material examined</emphasis>
.
<collectingCountry id="F302AFDD2960FF9FFDD0FC9DEE47FCBE" box="[524,617,809,835]" name="China" pageId="32" pageNumber="33">CHINA</collectingCountry>
:
<emphasis id="B961335F2960FF9FFDABFC9CEF92FCBF" bold="true" box="[631,956,808,834]" pageId="32" pageNumber="33">
<collectingRegion id="49D121AF2960FF9FFDABFC9CEF3CFCBF" box="[631,786,808,834]" country="China" name="Chongqing" pageId="32" pageNumber="33">Chongqing</collectingRegion>
Municipality
</emphasis>
:
<specimenCount id="9D1324C42960FF9FFC12FC9DE826FCBE" box="[974,1032,809,835]" pageId="32" pageNumber="33" type="male">4♂♂</specimenCount>
,
<specimenCount id="9D1324C42960FF9FFBCBFC9DE87FFCBE" box="[1047,1105,809,835]" pageId="32" pageNumber="33" type="female">2♀♀</specimenCount>
, ‘
<collectingCountry id="F302AFDD2960FF9FFBB9FC9DE8E9FCBE" box="[1125,1223,809,835]" name="China" pageId="32" pageNumber="33">CHINA</collectingCountry>
: SE
<collectingRegion id="49D121AF2960FF9FFADCFC9CE974FCBF" box="[1280,1370,808,834]" country="China" name="Sichuan" pageId="32" pageNumber="33">Sichuan</collectingRegion>
Jinfo Shan,
<geoCoordinate id="EE21898A2960FF9FFF03FCF9ED19FC9A" box="[223,311,845,871]" degrees="29" direction="north" minutes="01" orientation="latitude" pageId="32" pageNumber="33" precision="925" value="29.016666">29°01N</geoCoordinate>
<geoCoordinate id="EE21898A2960FF9FFE9CFCF9ED8BFC9A" box="[320,421,845,871]" degrees="107" direction="east" minutes="14" orientation="longitude" pageId="32" pageNumber="33" precision="925" value="107.23333">107°14E</geoCoordinate>
,
<quantity id="4CED42A82960FF9FFE6FFCF8EE7EFC9A" box="[435,592,844,871]" metricMagnitude="3" metricUnit="m" metricValue="1.825" metricValueMax="1.95" metricValueMin="1.7" pageId="32" pageNumber="33" unit="m" value="1825.0" valueMax="1950.0" valueMin="1700.0">1700–1950 m</quantity>
24–29.VI.98, J.Schneider // 1998
<collectingCountry id="F302AFDD2960FF9FFC00FCF8E831FC9B" box="[988,1055,844,870]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
Expedition J. Farkač, D. Král, J. Schneider &amp; A. Smetana’ (NHMB,
<collectionCode id="ED0477882960FF9FFDE5FCC5EE8FFC76" box="[569,673,881,907]" pageId="32" pageNumber="33">NMNHS</collectionCode>
, cDW);
<specimenCount id="9D1324C42960FF9FFCDBFCC5EF04FC76" box="[775,810,881,907]" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
,
<collectingCountry id="F302AFDD2960FF9FFCE0FCC5EFB6FC76" box="[828,920,881,907]" name="China" pageId="32" pageNumber="33">CHINA</collectingCountry>
,
<collectingRegion id="49D121AF2960FF9FFC7AFCC4E80BFC77" box="[934,1061,880,906]" country="China" name="Chongqing" pageId="32" pageNumber="33">Chongqing</collectingRegion>
, Nan–chuan, Jinfo Mt. N slope,
<quantity id="4CED42A82960FF9FFF4BFC21ECC6FC52" box="[151,232,917,943]" metricMagnitude="3" metricUnit="m" metricValue="1.713" pageId="32" pageNumber="33" unit="m" value="1713.0">1713m</quantity>
,
<date id="FFABC98D2960FF9FFF2EFC21EDAAFC52" box="[242,388,917,943]" pageId="32" pageNumber="33" value="2012-08-18">2012.VIII.18</date>
day, N
<geoCoordinate id="EE21898A2960FF9FFE0EFC21EE7CFC52" box="[466,594,917,943]" degrees="29.04043" direction="east" orientation="longitude" pageId="32" pageNumber="33" precision="1" value="29.04043">29.04043 E</geoCoordinate>
107.18410 // under stone, broadleaf forest, SHI Hongliang, HUANG Xinlei, LIU Yizhou leg. IOZ CAS (
<collectionCode id="ED0477882960FF9FFE13FC0DEE3DFC2E" box="[463,531,953,979]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="32" pageNumber="33">IZAS</collectionCode>
);
<specimenCount id="9D1324C42960FF9FFDFBFC0DEE65FC2E" box="[551,587,953,979]" pageId="32" pageNumber="33" type="male">1♂</specimenCount>
, ‘China—Chongqing Jinfo Shan 7.5.–
<date id="FFABC98D2960FF9FFC2BFC0CE844FC2E" box="[1015,1130,952,979]" pageId="32" pageNumber="33" value="2018-05-13">13.5.2018</date>
Lgt. E. Kučera’ (cEK).
</paragraph>
<paragraph id="8BAAEF4D2960FF9FFF1BFC69EE70FC0A" blockId="32.[151,1437,448,1987]" box="[199,606,988,1015]" pageId="32" pageNumber="33">
TME:
<specimenCount id="9D1324C42960FF9FFECDFC69EDB0FC0B" box="[273,414,988,1015]" pageId="32" pageNumber="33" type="generic">9 specimens</specimenCount>
. TGE:
<specimenCount id="9D1324C42960FF9FFE31FC69EE06FC0A" box="[493,552,989,1015]" pageId="32" pageNumber="33" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D1324C42960FF9FFDEFFC69EE76FC0A" box="[563,600,989,1015]" pageId="32" pageNumber="33" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62960FF9FFF1BFBB4EEC9FB7A" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BAAEF4D2960FF9FFF1BFBB4EEC9FB7A" blockId="32.[151,1437,448,1987]" pageId="32" pageNumber="33">
<emphasis id="B961335F2960FF9FFF1BFBB4ED14FBE7" bold="true" box="[199,314,1024,1050]" pageId="32" pageNumber="33">Diagnosis</emphasis>
. This species differs from other taxa of the “
<taxonomicName id="4C1594CE2960FF9FFCC3FBB4EF8CFBE7" authorityName="Jedlicka. Consists" authorityYear="1931" box="[799,930,1024,1050]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2960FF9FFCC3FBB4EF8CFBE7" box="[799,930,1024,1050]" italics="true" pageId="32" pageNumber="33">rambouseki</emphasis>
</taxonomicName>
” group except
<taxonomicName id="4C1594CE2960FF9FFB94FBB5E8D3FBE7" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" box="[1096,1277,1024,1051]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="emeishanicus">
<emphasis id="B961335F2960FF9FFB94FBB5E877FBE7" box="[1096,1113,1025,1050]" italics="true" pageId="32" pageNumber="33">P</emphasis>
.
<emphasis id="B961335F2960FF9FFBBAFBB4E8D3FBE7" box="[1126,1277,1024,1050]" italics="true" pageId="32" pageNumber="33">emeishanicus</emphasis>
</taxonomicName>
in its PW/HW &lt;2.10 (
<tableCitation id="C697DAF62960FF9FFF34FB90ED6FFBC2" box="[232,321,1060,1087]" captionStart="TABLE 2" captionStartId="36.[151,239,646,672]" captionText="TABLE 2. Morphometric data for species of the genus Pseudoodes gen. n." pageId="32" pageNumber="33">Table 2</tableCitation>
). It is readily distinguished from
<taxonomicName id="4C1594CE2960FF9FFD62FB91EF5BFBC3" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" box="[702,885,1060,1087]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="emeishanicus">
<emphasis id="B961335F2960FF9FFD62FB91EEE1FBC3" box="[702,719,1061,1086]" italics="true" pageId="32" pageNumber="33">P</emphasis>
.
<emphasis id="B961335F2960FF9FFD02FB90EF5BFBC3" box="[734,885,1060,1086]" italics="true" pageId="32" pageNumber="33">emeishanicus</emphasis>
</taxonomicName>
by its PW/PL ≤1.61. In addition,
<taxonomicName id="4C1594CE2960FF9FFB25FB91E9B3FBC3" authorityName="Jedlicka. Consists" authorityYear="1931" box="[1273,1437,1060,1087]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2960FF9FFB25FB91E924FBC3" box="[1273,1290,1061,1086]" italics="true" pageId="32" pageNumber="33">P</emphasis>
.
<emphasis id="B961335F2960FF9FFAC6FB90E9B3FBC3" box="[1306,1437,1060,1086]" italics="true" pageId="32" pageNumber="33">rambouseki</emphasis>
</taxonomicName>
differs from all members of the group in the male mesotibia strongly curved and dilated distally, highest W/Lp2, and apical lamella of median lobe bent to left (
<figureCitation id="132EF3C82960FF9FFDAEFBD8EEF5FB7A" box="[626,731,1132,1159]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Fig. 12D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62960FF9FFF1BFB24E9B2F83E" pageId="32" pageNumber="33" type="description">
<paragraph id="8BAAEF4D2960FF9FFF1BFB24E9B2F83E" blockId="32.[151,1437,448,1987]" pageId="32" pageNumber="33">
<emphasis id="B961335F2960FF9FFF1BFB24ED7FFB57" bold="true" box="[199,337,1168,1194]" pageId="32" pageNumber="33">Description</emphasis>
.
<emphasis id="B961335F2960FF9FFEBCFB24ED92FB56" bold="true" box="[352,444,1168,1195]" italics="true" pageId="32" pageNumber="33">Habitus</emphasis>
. Moderately large specimens (BL: about
<quantity id="4CED42A82960FF9FFC4EFB25E816FB56" box="[914,1080,1169,1195]" metricMagnitude="-2" metricUnit="m" metricValue="1.285" metricValueMax="1.33" metricValueMin="1.24" pageId="32" pageNumber="33" unit="mm" value="12.85" valueMax="13.3" valueMin="12.4">12.4–13.3 mm</quantity>
, BW:
<quantity id="4CED42A82960FF9FFB5CFB24E925FB56" box="[1152,1291,1168,1195]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.2" metricValueMin="4.7" pageId="32" pageNumber="33" unit="mm" value="4.95" valueMax="5.2" valueMin="4.7">4.7–5.2 mm</quantity>
), with ovate, subconvex body (
<figureCitation id="132EF3C82960FF9FFE82FB00EE23FB32" box="[350,525,1204,1231]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Figs 12A, B, E</figureCitation>
).
<emphasis id="B961335F2960FF9FFDFDFB00EF62FB32" bold="true" box="[545,844,1204,1231]" italics="true" pageId="32" pageNumber="33">Ratios and measurements</emphasis>
. See
<tableCitation id="C697DAF62960FF9FFC56FB00EFCFFB32" box="[906,993,1204,1231]" captionStart="TABLE 2" captionStartId="36.[151,239,646,672]" captionText="TABLE 2. Morphometric data for species of the genus Pseudoodes gen. n." pageId="32" pageNumber="33">Table 2</tableCitation>
.
<emphasis id="B961335F2960FF9FFC31FB00E884FB32" bold="true" box="[1005,1194,1204,1231]" italics="true" pageId="32" pageNumber="33">Color and luster</emphasis>
. Body dark brown to black; antennae, palpi, tibiae and tarsi rufopiceous. Integument moderately shiny, without iridescence.
<emphasis id="B961335F2960FF9FFADAFB6CE9BBFB0F" bold="true" box="[1286,1429,1240,1266]" italics="true" pageId="32" pageNumber="33">Punctuation</emphasis>
. Dorsal surface without punctuation; prosternum punctate at sides; proepisternum finely and shallowly punctate and striate; meso- and metepisternum, and metasternum at sides with dense and moderately strong punctuation; abdominal ventrites 1–5 punctate and finely rugose at sides, ventrite 6 densely punctate at apex.
<emphasis id="B961335F2960FF9FFB61FAF0E8D5FAA2" bold="true" box="[1213,1275,1348,1375]" italics="true" pageId="32" pageNumber="33">Head</emphasis>
. About half as wide as pronotum (
<tableCitation id="C697DAF62960FF9FFEB1FADCEDEAFA7E" box="[365,452,1384,1411]" captionStart="TABLE 2" captionStartId="36.[151,239,646,672]" captionText="TABLE 2. Morphometric data for species of the genus Pseudoodes gen. n." pageId="32" pageNumber="33">Table 2</tableCitation>
).
<collectingRegion id="49D121AF2960FF9FFE04FADDEE19FA7E" box="[472,567,1385,1411]" country="Myanmar" name="Chin" pageId="32" pageNumber="33">Mentum</collectingRegion>
tooth with apex more or less emarginate, with paramedial border finer than in other taxa of “
<taxonomicName id="4C1594CE2960FF9FFEEEFA38ED9BFA5B" authorityName="Jedlicka. Consists" authorityYear="1931" box="[306,437,1420,1446]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2960FF9FFEEEFA38ED9BFA5B" box="[306,437,1420,1446]" italics="true" pageId="32" pageNumber="33">rambouseki</emphasis>
</taxonomicName>
” group (
<figureCitation id="132EF3C82960FF9FFDC8FA38EE58FA5A" box="[532,630,1420,1447]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Fig. 12F</figureCitation>
).
<emphasis id="B961335F2960FF9FFD5BFA38EEF3FA5A" bold="true" box="[647,733,1420,1447]" italics="true" pageId="32" pageNumber="33">Thorax</emphasis>
. Pronotum with sides rounded toward posterior angles (PW/PB: 1.07–1.11); maximum width in posterior third; laterobasal impressions long, shallow, nearly reaching basal margin; base indistinctly sinuate; anterior angles rounded, rather projected. Prosternum with median longitudinal sulcus distinct; prosternal process elliptic, widely rounded apically (
<figureCitation id="132EF3C82960FF9FFC9FFA4CEF85F9EE" box="[835,939,1528,1555]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Fig. 12G</figureCitation>
). Metepisternum about one-and-a-half times wider than long (
<tableCitation id="C697DAF62960FF9FFE8EF9A8ED89F9CA" box="[338,423,1564,1591]" captionStart="TABLE 2" captionStartId="36.[151,239,646,672]" captionText="TABLE 2. Morphometric data for species of the genus Pseudoodes gen. n." pageId="32" pageNumber="33">Table 2</tableCitation>
), with lateral margin straight, its coadunation with epipleuron short, located anteriorly (
<figureCitation id="132EF3C82960FF9FFAB3F9A8ECE5F9A6" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Fig. 12H</figureCitation>
).
<emphasis id="B961335F2960FF9FFF3DF9F4ED1EF9A6" bold="true" box="[225,304,1600,1627]" pageId="32" pageNumber="33">
<emphasis id="B961335F2960FF9FFF3DF9F4ED07F9A6" bold="true" box="[225,297,1600,1627]" italics="true" pageId="32" pageNumber="33">Elytra</emphasis>
.
</emphasis>
Apical sinuation weak, poorly defined. Basal margin distinct laterally, forming distinct denticle at shoulder, disappearing medially at level between striae 2 and 3. Parascutellar striola rather reduced, punctiform; striae impunctate. Intervals 1–7 subconvex, interval 8 more convex and more raised than other intervals.
<emphasis id="B961335F2960FF9FFB3EF93DE936F95E" bold="true" box="[1250,1304,1673,1699]" italics="true" pageId="32" pageNumber="33">Legs</emphasis>
. Metacoxal basal sulcus barely reaching lateral third.Male mesotibia slightly concave on lateral margin,and significantly thickened apically along medial margin (
<figureCitation id="132EF3C82960FF9FFE3BF964EE62F916" box="[487,588,1744,1771]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Fig. 12B</figureCitation>
). Protarsomeres 1–3 of male strongly dilated, with second segment one and a half time wider than long (W/Lp2: 1.51–1.55).
<emphasis id="B961335F2960FF9FFD44F940EF13F8F2" bold="true" box="[664,829,1780,1807]" italics="true" pageId="32" pageNumber="33">Male genitalia</emphasis>
. Median lobe (
<figureCitation id="132EF3C82960FF9FFC3CF940E848F8F2" box="[992,1126,1780,1807]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Figs 12C, D</figureCitation>
) with basal bulb moderately long; angle between basal bulb and shaft acute; shaft long, swollen; apex short, curved ventrally; apical lamella more elongate than in other representatives of “
<taxonomicName id="4C1594CE2960FF9FFDBCF888EECDF8AB" authorityName="Jedlicka. Consists" authorityYear="1931" box="[608,739,1852,1878]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2960FF9FFDBCF888EECDF8AB" box="[608,739,1852,1878]" italics="true" pageId="32" pageNumber="33">rambouseki</emphasis>
</taxonomicName>
” group, left-orientated, rounded at tip.
<emphasis id="B961335F2960FF9FFB4DF888E97CF8AA" bold="true" box="[1169,1362,1852,1879]" italics="true" pageId="32" pageNumber="33">Female genitalia</emphasis>
. Basal gonocoxite with three lateroapical setae. Apical gonocoxite as in
<taxonomicName id="4C1594CE2960FF9FFC5FF8D5E874F887" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" box="[899,1114,1888,1915]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="leigongshanicus">
<emphasis id="B961335F2960FF9FFC5FF8D5EFBAF887" box="[899,916,1889,1914]" italics="true" pageId="32" pageNumber="33">P</emphasis>
.
<emphasis id="B961335F2960FF9FFC78F8D4E874F887" box="[932,1114,1888,1914]" italics="true" pageId="32" pageNumber="33">leigongshanicus</emphasis>
</taxonomicName>
(
<figureCitation id="132EF3C82960FF9FFBB7F8D4E8E4F886" box="[1131,1226,1888,1915]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Fig. 12I</figureCitation>
).
<collectingRegion id="49D121AF2960FF9FFB3CF8D5E90FF886" box="[1248,1313,1889,1915]" country="Turkey" name="Bursa" pageId="32" pageNumber="33">Bursa</collectingRegion>
copulatrix with small dorsal sclerite (not visible in
<figureCitation id="132EF3C82960FF9FFD88F830EE9EF862" box="[596,688,1924,1951]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Fig. 12J</figureCitation>
); spermatheca very long, slightly differentiated, with seminal canal narrow and receptaculum twisted apically; spermathecal gland connected in apical third of seminal canal (
<figureCitation id="132EF3C82960FF9FFAEDF81CE9A1F83E" box="[1329,1423,1960,1987]" captionStart="FIGURE 12" captionStartId="33.[151,250,1820,1844]" captionTargetBox="[261,1305,201,1783]" captionTargetId="figure-17@33.[244,1342,181,1796]" captionTargetPageId="33" captionText="FIGURE 12. Pseudoodes rambouseki (Jedlička, 1931) (A: lectotype of Holosoma rambouseki and its labels; B–D:topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I)." figureDoi="http://doi.org/10.5281/zenodo.4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="32" pageNumber="33">Fig. 12J</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF6ABFC52961FF9EFF4BF8A8EE62F814" ID-DOI="http://doi.org/10.5281/zenodo.4407096" ID-Zenodo-Dep="4407096" httpUri="https://zenodo.org/record/4407096/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1820,1844]" targetBox="[261,1305,201,1783]" targetPageId="33">
<paragraph id="8BAAEF4D2961FF9EFF4BF8A8EE62F814" blockId="33.[151,1437,1820,2025]" pageId="33" pageNumber="34">
<emphasis id="B961335F2961FF9EFF4BF8A8ED30F8C8" bold="true" box="[151,286,1820,1845]" pageId="33" pageNumber="34">FIGURE 12.</emphasis>
<taxonomicName id="4C1594CE2961FF9EFEFEF8A9EE95F8C8" authority="(Jedlicka, 1931)" authorityName="Guéorguiev &amp; Liang" authorityYear="2020" baseAuthorityName="Jedlicka" baseAuthorityYear="1931" box="[290,699,1821,1845]" class="Insecta" family="Carabidae" genus="Pseudoodes" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2961FF9EFEFEF8A9EE3BF8C8" box="[290,533,1821,1845]" italics="true" pageId="33" pageNumber="34">Pseudoodes rambouseki</emphasis>
(
<bibRefCitation id="EF8492BC2961FF9EFDFDF8A9EE9AF8C8" author="Jedlicka, A." box="[545,692,1821,1845]" pageId="33" pageNumber="34" pagination="21 - 30" refId="ref62314" refString="Jedlicka, A. (1931) Novi Carabidi ze Secuanu v Cine. (Col.) Neue Carabiden aus China-Szetschuan. Casopis Ceskoslovenske Spolecnosti Entomologicke, 28, 21 - 30." type="journal article" year="1931">Jedlička, 1931</bibRefCitation>
)
</taxonomicName>
(A: lectotype of
<taxonomicName id="4C1594CE2961FF9EFCBCF8A9E86DF8C8" authorityName="Jedlicka. Consists" authorityYear="1931" box="[864,1091,1821,1845]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="rambouseki">
<emphasis id="B961335F2961FF9EFCBCF8A9E86DF8C8" box="[864,1091,1821,1845]" italics="true" pageId="33" pageNumber="34">Holosoma rambouseki</emphasis>
</taxonomicName>
and its labels; B–D: topotype male specimen, China, Jinfo Shan, 29°01N 107°14E; E–H: topotype female specimen, China Jinfo Shan, N29.04043 E107.18410; I–J: topotype female specimen, SE Sichuan Jinfo Shan, 29°01N 107°14E). A–B, Е: habitus; C: median lobe of aedeagus, left lateral view; D: same, dorsal view; F: mentum; G: prosternal process; H: left metepisternum and metacoxa; I: left gonocoxite, ventral view; J: spermathecal complex and gonocoxites, ventral view. Scale lines = 2 mm (Figs A–B, E); = 1 mm (Fig. H); = 0.5 mm (Figs C–D, F–G, J); = 0.2 mm (Fig. I).
</paragraph>
</caption>
<subSubSection id="C30FBCC62962FF9DFF1BFF22E8FDFF28" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BAAEF4D2962FF9DFF1BFF22E8FDFF28" blockId="34.[151,1437,150,393]" pageId="34" pageNumber="35">
<emphasis id="B961335F2962FF9DFF1BFF22ED77FF4D" bold="true" box="[199,345,150,176]" pageId="34" pageNumber="35">Distribution</emphasis>
. This species is restricted to Mount Jinfo, Nanchuan District,
<collectingRegion id="49D121AF2962FF9DFBF0FF23E886FF4C" box="[1068,1192,151,177]" country="China" name="Chongqing" pageId="34" pageNumber="35">Chongqing</collectingRegion>
Municipality,
<collectingCountry id="F302AFDD2962FF9DFA8EFF23E9B7FF4C" box="[1362,1433,151,177]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
. Most probably, it may inhabit other parts of Dalou Mts., which extend also to
<collectingRegion id="49D121AF2962FF9DFBDCFF0FE8E1FF28" box="[1024,1231,187,213]" country="China" name="Guizhou" pageId="34" pageNumber="35">Guizhou Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30FBCC62962FF9DFF1BFF6AED45FE74" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BAAEF4D2962FF9DFF1BFF6AEF93FEE0" blockId="34.[151,1437,150,393]" pageId="34" pageNumber="35">
<emphasis id="B961335F2962FF9DFF1BFF6AED6DFF05" bold="true" box="[199,323,222,248]" pageId="34" pageNumber="35">Bionomics</emphasis>
. Almost nothing is known about the bionomics of this species. Label data indicate only that a single specimen was found under a stone, in a broadleaf forest, in the daytime.
</paragraph>
<paragraph id="8BAAEF4D2962FF9DFF1BFE93ED45FE74" blockId="34.[151,1437,150,393]" pageId="34" pageNumber="35">
<emphasis id="B961335F2962FF9DFF1BFE93ED24FEBC" bold="true" box="[199,266,295,321]" pageId="34" pageNumber="35">Notes</emphasis>
. In original description,
<bibRefCitation id="EF8492BC2962FF9DFDF7FE93EF27FEBC" author="Jedlicka, A." box="[555,777,295,321]" pageId="34" pageNumber="35" pagination="21 - 30" refId="ref62314" refString="Jedlicka, A. (1931) Novi Carabidi ze Secuanu v Cine. (Col.) Neue Carabiden aus China-Szetschuan. Casopis Ceskoslovenske Spolecnosti Entomologicke, 28, 21 - 30." type="journal article" year="1931">Jedlička (1931: 21)</bibRefCitation>
compared the species with
<taxonomicName id="4C1594CE2962FF9DFB91FE93E8DAFEBC" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[1101,1268,295,321]" class="Insecta" family="Carabidae" genus="Oodes" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="helopioides">
<emphasis id="B961335F2962FF9DFB91FE93E84FFEBC" box="[1101,1121,295,321]" italics="true" pageId="34" pageNumber="35">O</emphasis>
.
<emphasis id="B961335F2962FF9DFBAFFE93E8DAFEBC" box="[1139,1268,295,321]" italics="true" pageId="34" pageNumber="35">helopioides</emphasis>
</taxonomicName>
. Nevertheless, he placed it in another genus,
<taxonomicName id="4C1594CE2962FF9DFE24FEFFEF09FE98" authority="Semenov, 1889" authorityName="Semenov" authorityYear="1889" box="[504,807,331,357]" class="Insecta" family="Carabidae" genus="Holosoma" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B961335F2962FF9DFE24FEFFEE45FE98" box="[504,619,331,357]" italics="true" pageId="34" pageNumber="35">Holosoma</emphasis>
<bibRefCitation id="EF8492BC2962FF9DFDA9FEFFEF09FE98" author="Semenov, A. P." box="[629,807,331,357]" pageId="34" pageNumber="35" pagination="291 - 294" refId="ref64165" refString="Semenov, A. P. (1889) Chlaenius lepidus et Oodes integer, especes nouvelles de l'Asie russe. Horae Societatis Entomologicae Rossicae, 23, 291 - 294." type="journal article" year="1889">Semenov, 1889</bibRefCitation>
</taxonomicName>
, which at that time was classified in tribe
<taxonomicName id="4C1594CE2962FF9DFACBFEFFE94BFE98" authorityName=", Bousquet" authorityYear="1996" box="[1303,1381,331,357]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="tribe" tribe="Oodini">Oodini</taxonomicName>
(see
<bibRefCitation id="EF8492BC2962FF9DFF4BFEDBED71FE74" author="Gueorguiev, B. V." box="[151,351,367,393]" pageId="34" pageNumber="35" pagination="87 - 104" refId="ref61520" refString="Gueorguiev, B. V. (2014) Two new related oodine genera in the Oriental Region, with remarks on the systematic position of the genera Hololeius and Holosoma (Coleoptera, Carabidae). Deutsche Entomologische Zeitschrift, 61, 87 - 104. https: // doi. org / 10.3897 / dez. 61.7754" type="journal article" year="2014">Guéorguiev 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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