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<document id="161C61724298639D9C7D70F749620153" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFEC6D5ED6DDDFC7BB9B044E" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nipponophloeostiba Watanabe 1962" docType="key" docVersion="3" lastPageNumber="257" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="256" updateTime="1710782037732" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="E23D5E5A055D1D796B00FD34E66A1E90">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<treatment id="A15C878DFFEC6D5ED6DDDFC7BB9B044E" LSID="urn:lsid:plazi:treatment:A15C878DFFEC6D5ED6DDDFC7BB9B044E" httpUri="http://treatment.plazi.org/id/A15C878DFFEC6D5ED6DDDFC7BB9B044E" lastPageId="6" lastPageNumber="257" pageId="5" pageNumber="256">
<subSubSection id="61EF6510FFEC6D5DD6DDDFC7BA5D014A" box="[151,1114,1775,1802]" pageId="5" pageNumber="256" type="nomenclature">
<paragraph id="294A369BFFEC6D5DD6DDDFC7BA5D014A" blockId="5.[151,1114,1775,1802]" box="[151,1114,1775,1802]" pageId="5" pageNumber="256">
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<emphasis id="1B81EA89FFEC6D5DD6DDDFC7BA5D014A" bold="true" box="[151,1114,1775,1802]" pageId="5" pageNumber="256">
Key to species of the genus
<taxonomicName id="EEF54D18FFEC6D5DD7A6DFD8BCD80149" ID-CoL="8XN7R" authorityName="Watanabe" authorityYear="1962" box="[492,735,1775,1802]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFEC6D5DD7A6DFD8BCD80149" bold="true" box="[492,735,1775,1802]" italics="true" pageId="5" pageNumber="256">Nipponophloeostiba</emphasis>
</taxonomicName>
(modified from Shavrin 2020)
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<subSubSection id="61EF6510FFEC6D5ED6DDDE02BB9B044E" lastPageId="6" lastPageNumber="257" pageId="5" pageNumber="256" type="key">
<keyStep id="92012E3EFFEC6D5DD6DDDE02BB9B01FC" pageId="5" pageNumber="256">
<paragraph id="294A369BFFEC6D5DD6DDDE02BB9B01C6" blockId="5.[151,1437,1845,2043]" pageId="5" pageNumber="256">
<keyLead id="920495AEFFEC6D5DD6DDDE02BB9B01C6" pageId="5" pageNumber="256">
1 Postocular portions of head distinctly widened lateroposteriad, with short posterior carina not reaching posterior margin of eye. Aedeagus as in
<figureCitation id="B1CE2A1EFFEC6D5DD733DE65BFEE012B" box="[377,489,1874,1896]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="5" pageNumber="256">
Figs.
<quantity id="EE0D9B7EFFEC6D5DD7E4DE65BFEE012B" box="[430,489,1874,1896]" metricMagnitude="0" metricUnit="m" metricValue="2.1971" metricValueMax="2.2098" metricValueMin="2.1844" pageId="5" pageNumber="256" unit="in" value="86.5" valueMax="87.0" valueMin="86.0">86–87</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFEC6D5DD442DE65BCA90124" author="Watanabe, Y." box="[520,686,1874,1896]" pageId="5" pageNumber="256" pagination="59 - 391" refId="ref36201" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990)</bibRefCitation>
. Head and pronotum reddish-brown, elytra and abdomen yellow-brown. Body length:
<quantity id="EE0D9B7EFFEC6D5DD761DE58BFBE01C6" box="[299,441,1903,1925]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="5" pageNumber="256" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.30–2.70 mm</quantity>
. South
<collectingCountry id="51E2760BFFEC6D5DD44ADE47BC3001C6" box="[512,567,1904,1925]" name="Japan" pageId="5" pageNumber="256">Japan</collectingCountry>
..............................................
<taxonomicName id="EEF54D18FFEC6D5DD23FDE47BB9B01C6" ID-CoL="47HB5" authority="Watanabe, 1962" authorityName="Watanabe" authorityYear="1962" box="[1141,1436,1903,1925]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="verrucifera">
<emphasis id="1B81EA89FFEC6D5DD23FDE47BAFA01C7" box="[1141,1277,1903,1925]" italics="true" pageId="5" pageNumber="256">N. verrucifera</emphasis>
<bibRefCitation id="4D644B6AFFEC6D5DD348DE58BB9B01C6" author="Watanabe, Y." box="[1282,1436,1903,1925]" pageId="5" pageNumber="256" pagination="77 - 80" refId="ref36155" refString="Watanabe, Y. (1962) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe, 1962</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="294A369BFFEC6D5DD6DDDEBABB9B01FC" blockId="5.[151,1437,1845,2043]" pageId="5" pageNumber="256">
<keyLead id="920495AEFFEC6D5DD6DDDEBABB9B01FC" pageId="5" pageNumber="256">- Postocular portions of head not or indistinctly widened lateroposteriad, with longer posterior carina reaching posterior margin of eye or basal margin of ptonotum. Coloration different...................................................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="92012E3EFFEC6D5ED6DDDEF0BB9B06A4" lastPageId="6" lastPageNumber="257" pageId="5" pageNumber="256">
<paragraph id="294A369BFFEC6D5ED6DDDEF0BB9B06EE" blockId="5.[151,1437,1845,2043]" lastBlockId="6.[151,1437,151,525]" lastPageId="6" lastPageNumber="257" pageId="5" pageNumber="256">
<keyLead id="920495AEFFEC6D5ED6DDDEF0BB9B06EE" lastPageId="6" lastPageNumber="257" pageId="5" pageNumber="256">
2 Head as broad as pronotum. Elytra moderately long, significantly longer than pronotum. Median lobe and parameres narrow as in
<figureCitation id="B1CE2A1EFFEC6D5DD6B1DED2BF3401B9" box="[251,307,2021,2043]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="5" pageNumber="256">
Fig.
<quantity id="EE0D9B7EFFEC6D5DD76DDED2BF3401B9" box="[295,307,2021,2042]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="256" unit="in" value="2.0">2</quantity>
</figureCitation>
in Shavrin (2020). Head and abdomen brown, pronotum reddish-brown. Body length:
<quantity id="EE0D9B7EFFEC6D5DD23EDED2BB0501B9" box="[1140,1282,2021,2043]" metricMagnitude="-3" metricUnit="m" metricValue="1.985" metricValueMax="2.22" metricValueMin="1.75" pageId="5" pageNumber="256" unit="mm" value="1.985" valueMax="2.22" valueMin="1.75">1.75–2.22 mm</quantity>
.
<collectingCountry id="51E2760BFFEC6D5DD346DED2BB6301B8" box="[1292,1380,2021,2043]" name="Malaysia" pageId="5" pageNumber="256">Malaysia</collectingCountry>
....................................................................................
<taxonomicName id="EEF54D18FFEF6D5ED2FAD9AFBB9B06EE" ID-CoL="8Y6P9" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[1200,1436,151,173]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="thayeri">
<emphasis id="1B81EA89FFEF6D5ED2FAD9AFBB1706EF" box="[1200,1296,151,173]" italics="true" pageId="6" pageNumber="257">N. thayeri</emphasis>
Shavrin, 2020
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="294A369BFFEF6D5ED6DDD982BB9B06A4" blockId="6.[151,1437,151,525]" pageId="6" pageNumber="257">
<keyLead id="920495AEFFEF6D5ED6DDD982BB9B06A4" pageId="6" pageNumber="257">- Head slightly narrower or broader than pronotum. Elytra shorter, slightly broader than pronotum. Median lobes and parameres different. Coloration different........................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="92012E3EFFEF6D5ED6DDD9C7BB9B071E" pageId="6" pageNumber="257">
<paragraph id="294A369BFFEF6D5ED6DDD9C7BB9B077C" blockId="6.[151,1437,151,525]" pageId="6" pageNumber="257">
<keyLead id="920495AEFFEF6D5ED6DDD9C7BB9B077C" pageId="6" pageNumber="257">
3 Head slightly narrower than pronotum.Aedeagus as in
<figureCitation id="B1CE2A1EFFEF6D5ED4A6D9D8BD280746" box="[748,815,239,261]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="6" pageNumber="257">
Fig.
<quantity id="EE0D9B7EFFEF6D5ED55DD9C7BD280746" box="[791,815,240,261]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="6" pageNumber="257" unit="in" value="39.0">39</quantity>
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in Shavrin (2020). Head, pronotum and abdomen reddish-brown, elytra yellow-brown, with darker parascutellar portion. Body length:
<quantity id="EE0D9B7EFFEF6D5ED530D83ABDD50761" box="[890,978,269,290]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="6" pageNumber="257" unit="mm" value="2.07">2.07 mm</quantity>
.
<collectingCountry id="51E2760BFFEF6D5ED591D83ABA0B0761" box="[987,1036,269,290]" name="Laos" pageId="6" pageNumber="257">Laos</collectingCountry>
,
<collectingCountry id="51E2760BFFEF6D5ED25CD83ABA740760" box="[1046,1139,269,291]" name="Indonesia" pageId="6" pageNumber="257">Indonesia</collectingCountry>
(
<collectingRegion id="EB31F879FFEF6D5ED2CAD83ABADE0760" box="[1152,1241,269,291]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="257">Sulawesi</collectingRegion>
)............................................................................................
<taxonomicName id="EEF54D18FFEF6D5ED2CFD81CBB9B077C" ID-CoL="8Y6P8" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[1157,1436,298,320]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="longilobata">
<emphasis id="1B81EA89FFEF6D5ED2CFD81CBB17077C" box="[1157,1296,298,320]" italics="true" pageId="6" pageNumber="257">N. longilobata</emphasis>
Shavrin, 2020
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="294A369BFFEF6D5ED6DDD87FBB9B071E" blockId="6.[151,1437,151,525]" box="[151,1436,327,349]" pageId="6" pageNumber="257">
<keyLead id="920495AEFFEF6D5ED6DDD87FBB9B071E" box="[151,1436,327,349]" pageId="6" pageNumber="257">- Head slightly broader than pronotum. Aedeagus and coloration different. Body larger............................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="92012E3EFFEF6D5ED6DDD852BB9B044E" pageId="6" pageNumber="257">
<paragraph id="294A369BFFEF6D5ED6DDD852BB9B07F6" blockId="6.[151,1437,151,525]" pageId="6" pageNumber="257">
<keyLead id="920495AEFFEF6D5ED6DDD852BB9B07F6" pageId="6" pageNumber="257">
4 Temples slightly shorter than longitudinal length of eyes. Elytra as broad as long. Body reddish-brown. Median lobe narrow; parameres strongly broadened in apical portions as in
<figureCitation id="B1CE2A1EFFEF6D5ED4A3D8B5BD2607D4" box="[745,801,386,408]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="6" pageNumber="257">
Fig.
<quantity id="EE0D9B7EFFEF6D5ED55FD8B5BD2607D4" box="[789,801,386,407]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="257" unit="in" value="3.0">3</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFEF6D5ED575D8B5BDD907D4" author="Shavrin, A. V." box="[831,990,386,408]" pageId="6" pageNumber="257" pagination="245 - 250" refId="ref35879" refString="Shavrin, A. V. (2023 a) Two new species and a new record of Omaliini McLeay, 1825 (Coleoptera: Staphylinidae: Omaliinae) from Indonesia. Zootaxa, 5258 (2), 245 - 250. https: // doi. org / 10.11646 / zootaxa. 5258.2.7" type="journal article" year="2023">Shavrin (2023a)</bibRefCitation>
. Habitus as in
<figureCitation id="B1CE2A1EFFEF6D5ED220D8B5BAA507D4" box="[1130,1186,386,408]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="6" pageNumber="257">
Fig.
<quantity id="EE0D9B7EFFEF6D5ED2DCD8B5BAA507D4" box="[1174,1186,386,407]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="257" unit="in" value="1.0">1</quantity>
</figureCitation>
in
<bibRefCitation id="4D644B6AFFEF6D5ED28BD8B5BB5807D4" author="Shavrin, A. V." box="[1217,1375,386,408]" pageId="6" pageNumber="257" pagination="245 - 250" refId="ref35879" refString="Shavrin, A. V. (2023 a) Two new species and a new record of Omaliini McLeay, 1825 (Coleoptera: Staphylinidae: Omaliinae) from Indonesia. Zootaxa, 5258 (2), 245 - 250. https: // doi. org / 10.11646 / zootaxa. 5258.2.7" type="journal article" year="2023">Shavrin (2023a)</bibRefCitation>
. Body length:
<quantity id="EE0D9B7EFFEF6D5ED761D897BF8407F6" box="[299,387,416,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="6" pageNumber="257" unit="mm" value="2.33">2.33 mm</quantity>
.
<collectingCountry id="51E2760BFFEF6D5ED7C6D8A8BFEE07F6" box="[396,489,415,437]" name="Indonesia" pageId="6" pageNumber="257">Indonesia</collectingCountry>
(
<collectingRegion id="EB31F879FFEF6D5ED7BCD8A8BC2707F6" box="[502,544,415,437]" country="Indonesia" name="Bali" pageId="6" pageNumber="257">Bali</collectingRegion>
)...................................................
<taxonomicName id="EEF54D18FFEF6D5ED2DFD897BB9B07F6" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1173,1436,415,437]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="explanata">
<emphasis id="1B81EA89FFEF6D5ED2DFD897BAA207F6" box="[1173,1189,416,437]" italics="true" pageId="6" pageNumber="257">N</emphasis>
.
<emphasis id="1B81EA89FFEF6D5ED2FBD8A8BB1707F7" box="[1201,1296,415,436]" italics="true" pageId="6" pageNumber="257">explanata</emphasis>
Shavrin, 2023
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="294A369BFFEF6D5ED6DDD88ABB9B044E" blockId="6.[151,1437,151,525]" pageId="6" pageNumber="257">
<keyLead id="920495AEFFEF6D5ED6DDD88ABB9B044E" pageId="6" pageNumber="257">
- Temples three times shorter than longitudinal length of eyes. Elytra slightly longer than broad. Head, pronotum and abdomen reddish-brown; elytra yellow-brown, with reddish apical portion. Median lobe wide; parameres slightly broadened in apical portions (
<figureCitation id="B1CE2A1EFFEF6D5ED774D8C0BF7E044E" box="[318,377,503,525]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="6" pageNumber="257">Fig. 7</figureCitation>
). Habitus as in
<figureCitation id="B1CE2A1EFFEF6D5ED444D8C0BC40044E" box="[526,583,503,525]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="6" pageNumber="257">Fig. 1</figureCitation>
. Body length:
<quantity id="EE0D9B7EFFEF6D5ED49FD8CFBD2A044E" box="[725,813,504,525]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" pageId="6" pageNumber="257" unit="mm" value="2.34">2.34 mm</quantity>
.
<collectingCountry id="51E2760BFFEF6D5ED57CD8CFBDE8044E" box="[822,1007,503,525]" name="Papua New Guinea" pageId="6" pageNumber="257">Papua New Guinea</collectingCountry>
.................
<taxonomicName id="EEF54D18FFEF6D5ED287D8CFBB67044F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1229,1376,503,525]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="hornabrooki" status="sp. n.">
<emphasis id="1B81EA89FFEF6D5ED287D8CFBADA044E" box="[1229,1245,504,525]" italics="true" pageId="6" pageNumber="257">N</emphasis>
.
<emphasis id="1B81EA89FFEF6D5ED2A3D8C0BB67044F" box="[1257,1376,503,524]" italics="true" pageId="6" pageNumber="257">hornabrooki</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="00B257F2FFEF6D5ED32CD8CFBB9B044E" box="[1382,1436,504,525]" pageId="6" pageNumber="257" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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