303 lines
38 KiB
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303 lines
38 KiB
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<document id="C0252F29260DA94B6522B4FC5A900110" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFC06D73D6DDDE10BD6D03DE" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba rufula Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="294" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="292" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="057E25FEE95C9FBA8026437CD11D5F15">
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<mods:title id="CB2A2967065A3A5F85D39F761ED3C784">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<mods:name id="0597006DAC95ABEEADE53ADA24A82397" type="personal">
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<mods:roleTerm id="188781E1928F8EF95E5837AF29CD3BF8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="16B29097D912BD724BFB92FD7A27E284">Shavrin, Alexey V.</mods:namePart>
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<mods:typeOfResource id="5C9196D62EA836DCD08A0796529814EE">text</mods:typeOfResource>
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<mods:title id="808239517A7FED31CEEB4AA918C3B3A6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="05037D4B988EF55AC9D865AE5CA3FF8E">
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<mods:date id="C8C4E849F9DEB9D792A980C028D68F63">2024</mods:date>
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<mods:detail id="9B6DF2F52D0BDB94AC5B098FA0B6EE86" type="pubDate">
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<mods:number id="5C744BD7530247832FE56D496676F76E">2024-03-14</mods:number>
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<mods:number id="D6010E7DA62C49165AF62D391D5C7858">5424</mods:number>
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</mods:detail>
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<mods:detail id="5E4F76BF18330C33F74ADA86CEA7C7C0" type="issue">
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<mods:number id="01D6346230B8408411862B02EB8AB6C7">3</mods:number>
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<mods:start id="EC0DD0BA9C192368ED631C7E08175E00">251</mods:start>
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<mods:end id="0054820CA21C838451128F517F4D28EA">307</mods:end>
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<mods:location id="F01A2EE9487C7C5F159A7BBD1BE70483">
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<mods:url id="6AEDCDD7FDA7F77DEC193787488DB615">http://dx.doi.org/10.11646/zootaxa.5424.3.1</mods:url>
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</mods:location>
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<mods:classification id="5380C454C40EE641E6D21FD2ED3DB3BD">journal article</mods:classification>
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<mods:identifier id="B2E75161E7304D851BDE9DC3D4B0898A" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
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<mods:identifier id="34887FEDDFE6FE27ABEBDA8F5F39FFD0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A56ECAFAAF06026B60C8DF49F8526E41" type="Zenodo-Dep">10821214</mods:identifier>
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<mods:identifier id="D5FF4C6D1393294AFFE840378635DC6A" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
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</mods:mods>
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<treatment id="A15C878DFFC06D73D6DDDE10BD6D03DE" LSID="urn:lsid:plazi:treatment:A15C878DFFC06D73D6DDDE10BD6D03DE" httpUri="http://treatment.plazi.org/id/A15C878DFFC06D73D6DDDE10BD6D03DE" lastPageId="43" lastPageNumber="294" pageId="41" pageNumber="292">
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<subSubSection id="61EF6510FFC06D71D6DDDE10BFFD0101" box="[151,506,1831,1858]" pageId="41" pageNumber="292" type="nomenclature">
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<paragraph id="294A369BFFC06D71D6DDDE10BFFD0101" blockId="41.[151,506,1831,1894]" box="[151,506,1831,1858]" pageId="41" pageNumber="292">
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<heading id="720281F7FFC06D71D6DDDE10BFFD0101" bold="true" box="[151,506,1831,1858]" fontSize="11" level="1" pageId="41" pageNumber="292" reason="1">
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<emphasis id="1B81EA89FFC06D71D6DDDE10BFFD0101" bold="true" box="[151,506,1831,1858]" pageId="41" pageNumber="292">
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<taxonomicName id="EEF54D18FFC06D71D6DDDE10BFB60101" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,433,1831,1858]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">
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<emphasis id="1B81EA89FFC06D71D6DDDE10BFB60101" bold="true" box="[151,433,1831,1858]" italics="true" pageId="41" pageNumber="292">Paraphloeostiba rufula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00B257F2FFC06D71D7F2DE1FBFFD0101" box="[440,506,1832,1858]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFC06D71D6DDDE7BBF6A0125" box="[151,365,1868,1894]" pageId="41" pageNumber="292" type="description">
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<paragraph id="294A369BFFC06D71D6DDDE7BBF6A0125" blockId="41.[151,506,1831,1894]" box="[151,365,1868,1894]" pageId="41" pageNumber="292">
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(
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<figureCitation id="B1CE2A1EFFC06D71D6D5DE7BBEF10125" box="[159,246,1868,1894]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="41" pageNumber="292">Figs 80</figureCitation>
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,
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<figureCitation id="B1CE2A1EFFC06D71D748DE7BBF620125" box="[258,357,1868,1894]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="41" pageNumber="292">126–132</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFC06D72D6DDDEA3BAD606B4" lastPageId="42" lastPageNumber="293" pageId="41" pageNumber="292" type="materials_examined">
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<paragraph id="294A369BFFC06D71D6DDDEA3BFAA01B5" blockId="41.[151,1436,1940,2038]" pageId="41" pageNumber="292">
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<emphasis id="1B81EA89FFC06D71D6DDDEA3BF4101ED" bold="true" box="[151,326,1940,1966]" pageId="41" pageNumber="292">Type material.</emphasis>
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<typeStatus id="F64E8839FFC06D71D704DEA3BFB101ED" box="[334,438,1940,1966]" pageId="41" pageNumber="292" type="holotype">Holotype</typeStatus>
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<specimenCount id="3FF3FD12FFC06D71D7F4DEA2BFD401EC" box="[446,467,1941,1967]" count="1" pageId="41" pageNumber="292" type="male">♂</specimenCount>
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(dissected): ‘
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<collectionCode id="4FE4AE5EFFC06D71D422DEA3BCAB01ED" box="[616,684,1940,1966]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="41" pageNumber="292">NEW</collectionCode>
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<collectingCountry id="51E2760BFFC06D71D4FEDEA3BD1901ED" box="[692,798,1940,1966]" name="Guinea-Bissau" pageId="41" pageNumber="292">GUINEA</collectingCountry>
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| Okapa,
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<date id="5D4B105BFFC06D71D5C6DEA3BA0901EC" box="[908,1038,1940,1967]" pageId="41" pageNumber="292" value="1965-06-16">16.VI.1965</date>
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, | flying in daylight’ <printed>, ‘
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<collectionCode id="4FE4AE5EFFC06D71D335DEA3BB9F01ED" box="[1407,1432,1940,1966]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="41" pageNumber="292">R</collectionCode>
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. Hornabrook | BM 1970-232.’ <printed>, ‘
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<typeStatus id="F64E8839FFC06D71D426DE8FBD020191" box="[620,773,1976,2002]" pageId="41" pageNumber="292" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="EEF54D18FFC06D71D556DE8FBDD00191" authorityName="Steel" authorityYear="1960" box="[796,983,1976,2002]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="genus">
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<emphasis id="1B81EA89FFC06D71D556DE8FBDD00191" box="[796,983,1976,2002]" italics="true" pageId="41" pageNumber="292">Paraphloeostiba</emphasis>
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</taxonomicName>
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<taxonomicName id="EEF54D18FFC06D71D5A6DE8FBA370191" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1004,1072,1976,2002]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">
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<emphasis id="1B81EA89FFC06D71D5A6DE8FBA370191" box="[1004,1072,1976,2002]" italics="true" pageId="41" pageNumber="292">rufula</emphasis>
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</taxonomicName>
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<emphasis id="1B81EA89FFC06D71D273DE8FBA7D0191" bold="true" box="[1081,1146,1976,2002]" pageId="41" pageNumber="292">
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<taxonomicNameLabel id="00B257F2FFC06D71D273DE8FBA7D0191" box="[1081,1146,1976,2002]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.
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<collectionCode id="4FE4AE5EFFC06D71D34DDE8FBB1A0191" box="[1287,1309,1976,2002]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="41" pageNumber="292" type="Museum">V</collectionCode>
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. des. 2024’ <red, printed> (
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<collectionCode id="4FE4AE5EFFC06D71D70DDEEBBFA601B5" box="[327,417,2012,2038]" pageId="41" pageNumber="292">BMNH</collectionCode>
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).
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</paragraph>
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<paragraph id="294A369BFFC36D72D68DD9A0BAD606B4" blockId="42.[151,1437,151,632]" pageId="42" pageNumber="293">
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<typeStatus id="F64E8839FFC36D72D68DD9A0BF3F06F2" box="[199,312,151,177]" pageId="42" pageNumber="293" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="3FF3FD12FFC36D72D702D9A0BF8B06F2" box="[328,396,151,177]" count="3" pageId="42" pageNumber="293" type="female">3 ♀♀</specimenCount>
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(
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<specimenCount id="3FF3FD12FFC36D72D7D5D9A0BC3806F2" box="[415,575,151,177]" count="1" pageId="42" pageNumber="293" type="generic">one specimen</specimenCount>
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dissected;
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<specimenCount id="3FF3FD12FFC36D72D48FD9A0BD6406F2" box="[709,867,151,177]" count="1" pageId="42" pageNumber="293" type="generic">one specimen</specimenCount>
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without right antennomeres 6–11); underside of the card of
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<specimenCount id="3FF3FD12FFC36D72D751D98DBFC50690" box="[283,450,185,212]" count="2" pageId="42" pageNumber="293" type="generic">two specimens</specimenCount>
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with handwritten ‘1/64’): same data as the holotype, with additional red printed label: ‘
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<typeStatus id="F64E8839FFC36D72D6D1D9EABF2C06B4" box="[155,299,221,247]" pageId="42" pageNumber="293" type="paratype">PARATYPE</typeStatus>
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<taxonomicName id="EEF54D18FFC36D72D775D9EBBFFD06B5" authorityName="Steel" authorityYear="1960" box="[319,506,220,246]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="293" phylum="Arthropoda" rank="genus">
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<emphasis id="1B81EA89FFC36D72D775D9EBBFFD06B5" box="[319,506,220,246]" italics="true" pageId="42" pageNumber="293">Paraphloeostiba</emphasis>
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</taxonomicName>
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<taxonomicName id="EEF54D18FFC36D72D447D9EBBC5606B5" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[525,593,220,246]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="293" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">
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<emphasis id="1B81EA89FFC36D72D447D9EBBC5606B5" box="[525,593,220,246]" italics="true" pageId="42" pageNumber="293">rufula</emphasis>
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</taxonomicName>
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<emphasis id="1B81EA89FFC36D72D413D9EABC9F06B4" bold="true" box="[601,664,221,247]" pageId="42" pageNumber="293">
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<taxonomicNameLabel id="00B257F2FFC36D72D413D9EABC9F06B4" box="[601,664,221,247]" pageId="42" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.
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<collectionCode id="4FE4AE5EFFC36D72D568D9EABD3F06B4" box="[802,824,221,247]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="42" pageNumber="293" type="Museum">V</collectionCode>
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. des. 2024’ <red, printed> (
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<collectionCode id="4FE4AE5EFFC36D72D221D9EABAC206B4" box="[1131,1221,221,247]" pageId="42" pageNumber="293">BMNH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFC36D73D68DD9C8BF0202AA" lastPageId="43" lastPageNumber="294" pageId="42" pageNumber="293" type="description">
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<paragraph id="294A369BFFC36D72D68DD9C8BB6E0723" blockId="42.[151,1437,151,632]" pageId="42" pageNumber="293">
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<emphasis id="1B81EA89FFC36D72D68DD9C8BF50075A" bold="true" box="[199,343,255,281]" pageId="42" pageNumber="293">Description.</emphasis>
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Measurements (n=4): HW: 0.45–0.46; HL: 0.23–0.24; OL: 0.15; TL: 0.04; AL (
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<typeStatus id="F64E8839FFC36D72D2F4D9C8BB21075A" box="[1214,1318,255,281]" pageId="42" pageNumber="293" type="holotype">holotype</typeStatus>
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): 0.61; PL: 0.37–0.41; PWmax: 0.55–0.61; PWmin: 0.48–0.53; ESL: 0.63–0.76; EW: 0.67–0.77; MTbL (
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<typeStatus id="F64E8839FFC36D72D2E8D815BB0D077F" box="[1186,1290,290,316]" pageId="42" pageNumber="293" type="holotype">holotype</typeStatus>
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): 0.35; MTrL (
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<typeStatus id="F64E8839FFC36D72D6D5D872BF00071C" box="[159,263,325,351]" pageId="42" pageNumber="293" type="holotype">holotype</typeStatus>
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): 0.18 (MTrL 1–4: 0.07; MTrL 5: 0.11); AW: 0.72–0.78; AedL: 0.55; BL: 2.00–2.32 (
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<typeStatus id="F64E8839FFC36D72D2FDD872BB19071C" box="[1207,1310,325,351]" pageId="42" pageNumber="293" type="holotype">holotype</typeStatus>
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: 2.15).
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</paragraph>
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<paragraph id="294A369BFFC36D72D68DD85FBF90043B" blockId="42.[151,1437,151,632]" pageId="42" pageNumber="293">
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Habitus as in
|
|||
|
|
<figureCitation id="B1CE2A1EFFC36D72D716D85FBFAB07C1" box="[348,428,360,386]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="42" pageNumber="293">Fig. 80</figureCitation>
|
|||
|
|
. Head, pronotum and abdomen reddish brown; elytra yellow-brown, with slightly darkened latero-apical and apical portion of pronotum; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellowish. Middle portion of head with very fine and sparse punctation; neck without visible punctures; pronotum with fine and sparse punctation, slightly larger than that on middle portion of head, sparser in middle and indistinct in mediobasal and mediolateral portions; elytra with moderately sparse punctation about as that on pronotum, finer and sparser in parascutellar portion and along suture. Pronotum with dense and isodiametric microsculpture, distinctly coarser than that in middle portion of head; microsculpture of elytra dense, slightly larger and less coarser than that on pronotum.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="7D8A6613FFC36D72D6DDDEACBD1B01BA" ID-DOI="http://doi.org/10.5281/zenodo.10821277" ID-Zenodo-Dep="10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="42" pageNumber="293" startId="42.[151,264,1947,1971]" targetBox="[159,1428,660,1914]" targetPageId="42" targetType="figure">
|
|||
|
|
<paragraph id="294A369BFFC36D72D6DDDEACBD1B01BA" blockId="42.[151,1436,1947,2042]" pageId="42" pageNumber="293">
|
|||
|
|
<emphasis id="1B81EA89FFC36D72D6DDDEACBF6B01F0" bold="true" box="[151,364,1947,1971]" pageId="42" pageNumber="293">FIGURES 126–132.</emphasis>
|
|||
|
|
<taxonomicName id="EEF54D18FFC36D72D73ADEACBC5B01F0" authorityName="Shavrin" authorityYear="2024" box="[368,604,1947,1971]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="293" phylum="Arthropoda" rank="species" species="rufula">
|
|||
|
|
<emphasis id="1B81EA89FFC36D72D73ADEACBC5B01F0" box="[368,604,1947,1971]" italics="true" pageId="42" pageNumber="293">Paraphloeostiba rufula</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DD9A0BB55075E" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">Head 1.9 times as broad as long, with short and deep anteocellar foveae as long as diameter of ocellus. Ocelli large, located slightly below level of postocular carinae; distance between ocelli about as long as distance between ocellus and posterior margin of eyes. Antennomeres 7–10 distinctly transverse; antennomere 5 indisticntly broader than 4, 6 slightly longer and distinctly broader than 5, 7 slightly shorter than 6, 8–10 slightly broader than 7.</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DD810BA3807CA" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
Pronotum 1.4 times as broad as long, 1.2–1.3 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; middle portion without impressions, but
|
|||
|
|
<specimenCount id="3FF3FD12FFC26D73D53ED87CBA160726" box="[884,1041,331,357]" count="2" pageId="43" pageNumber="294" type="generic" typeStatus="paratype">two paratypes</specimenCount>
|
|||
|
|
with traces of very indistinct wide impressions in mediobasal third; lateral and laterobasal portions slightly impressed.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DD8A4BC9C07EE" blockId="43.[151,1437,151,1437]" box="[199,667,403,429]" pageId="43" pageNumber="294">Elytra 1.7–1.8 times as long as pronotum.</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DD880BECF04C6" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
Male. Posterior margin of abdominal tergite VIII truncate (
|
|||
|
|
<figureCitation id="B1CE2A1EFFC26D73D521D880BDC80792" box="[875,975,439,465]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 128</figureCitation>
|
|||
|
|
). Posterior margin of abdominal sternite VIII concave (
|
|||
|
|
<figureCitation id="B1CE2A1EFFC26D73D70ED8ECBFAB07B6" box="[324,428,475,501]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 129</figureCitation>
|
|||
|
|
). Aedeagus with wide basal portion, gradually narrowed toward small rounded apex; parameres reaching apex of median lobe, distinctly broadened apically, with three moderately long apical and fourfive preapical setae, median parts of each paramere with three moderately long setae; internal sac moderately short and narrow, with elongate sclerotized structures in basal part (
|
|||
|
|
<figureCitation id="B1CE2A1EFFC26D73D510DB70BDB90422" box="[858,958,583,609]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 126</figureCitation>
|
|||
|
|
). Lateral aspect of the aedeagus as in
|
|||
|
|
<figureCitation id="B1CE2A1EFFC26D73D33ADB70BEC304C6" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 127</figureCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDBB8BC9104B2" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
Female. Posterior margin of abdominal tergite VIII truncate (
|
|||
|
|
<figureCitation id="B1CE2A1EFFC26D73D53FDBB8BDDF04EA" box="[885,984,655,681]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 130</figureCitation>
|
|||
|
|
). Posterior margin of abdominal sternite VIII rounded (
|
|||
|
|
<figureCitation id="B1CE2A1EFFC26D73D776DB84BF98048E" box="[316,415,691,717]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 131</figureCitation>
|
|||
|
|
). Accessory sclerite short, from widest apical portion gradually narrowed toward rounded apex (
|
|||
|
|
<figureCitation id="B1CE2A1EFFC26D73D691DBE0BF3904B2" box="[219,318,727,753]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="43" pageNumber="294">Fig. 132</figureCitation>
|
|||
|
|
). Spermatheca not recognized.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDBCDBA1E051E" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D68DDBCDBFB60556" bold="true" box="[199,433,762,789]" pageId="43" pageNumber="294">Comparative notes.</emphasis>
|
|||
|
|
Based on the very broad median lobe and apical portions of the parameres,
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D345DBCCBB740556" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1295,1395,763,789]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D345DBCCBB270557" box="[1295,1312,763,788]" italics="true" pageId="43" pageNumber="294">P</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D365DBCCBB740556" box="[1327,1395,763,789]" italics="true" pageId="43" pageNumber="294">rufula</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D331DBCCBEA9057A" bold="true" pageId="43" pageNumber="294">
|
|||
|
|
<taxonomicNameLabel id="00B257F2FFC26D73D331DBCCBEA9057A" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is similar to
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D774DA28BFB2057A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[318,437,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D774DA28BFB2057A" box="[318,437,799,825]" italics="true" pageId="43" pageNumber="294">P. assimile</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D7F1DA28BFFD057A" bold="true" box="[443,506,799,825]" pageId="43" pageNumber="294">
|
|||
|
|
<taxonomicNameLabel id="00B257F2FFC26D73D7F1DA28BFFD057A" box="[443,506,799,825]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D44DDA28BD1B057A" authority="Steel, 1960" authorityName="Steel" authorityYear="1960" box="[519,796,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="celebensis">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D44DDA28BC92057A" box="[519,661,799,825]" italics="true" pageId="43" pageNumber="294">P. celebensis</emphasis>
|
|||
|
|
<bibRefCitation id="4D644B6AFFC26D73D4D1DA28BD1B057A" author="Steel, W. O." box="[667,796,799,825]" pageId="43" pageNumber="294" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel, 1960</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D56CDA28BDA4057B" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[806,931,799,824]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="papuana">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D56CDA28BDA4057B" box="[806,931,799,824]" italics="true" pageId="43" pageNumber="294">P. papuana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D5FBDA28BA4B057A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[945,1100,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D5FBDA28BA4B057A" box="[945,1100,799,825]" italics="true" pageId="43" pageNumber="294">P. penelopeae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D218DA28BA96057A" bold="true" box="[1106,1169,799,825]" pageId="43" pageNumber="294">
|
|||
|
|
<taxonomicNameLabel id="00B257F2FFC26D73D218DA28BA96057A" box="[1106,1169,799,825]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D28FDA28BB57057A" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[1221,1360,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="tonkinesis">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D28FDA28BB57057A" box="[1221,1360,799,825]" italics="true" pageId="43" pageNumber="294">P. tonkinesis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. It can be distinguished from all these species by the following morphological features:
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDA50BD5F05E6" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
from
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D74FDA50BF7B05C2" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,380,871,897]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D74FDA50BF7B05C2" box="[261,380,871,897]" italics="true" pageId="43" pageNumber="294">P. assimile</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D7C9DA50BFC405C2" bold="true" box="[387,451,871,897]" pageId="43" pageNumber="294">
|
|||
|
|
<taxonomicNameLabel id="00B257F2FFC26D73D7C9DA50BFC405C2" box="[387,451,871,897]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
by the larger body, darker head, pronotum and abdomen, the presence of the punctation on the head, the narrower pronotum and the shorter parameres;
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDA98BA7F058A" blockId="43.[151,1437,151,1437]" box="[199,1144,943,969]" pageId="43" pageNumber="294">
|
|||
|
|
from
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D74FDA98BF94058A" authorityName="Steel" authorityYear="1960" box="[261,403,943,969]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="celebensis">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D74FDA98BF94058A" box="[261,403,943,969]" italics="true" pageId="43" pageNumber="294">P. celebensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the larger body and narrower apical portions of the parameres;
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDAE4BFB60252" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
from
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D74DDAE4BF8005AF" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[263,391,979,1004]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="papuana">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D74DDAE4BF8005AF" box="[263,391,979,1004]" italics="true" pageId="43" pageNumber="294">P. papuana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the larger and darker body, the narrower pronotum, and slightly longer parameres with narrower apical portions;
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDD2CBAA9021A" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
from
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D74DDD2CBFA10276" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[263,422,1051,1077]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D74DDD2CBFA10276" box="[263,422,1051,1077]" italics="true" pageId="43" pageNumber="294">P. penelopeae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D7E5DD2CBFF60276" bold="true" box="[431,497,1051,1077]" pageId="43" pageNumber="294">
|
|||
|
|
<taxonomicNameLabel id="00B257F2FFC26D73D7E5DD2CBFF60276" box="[431,497,1051,1077]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
by the darker body, the presence of the punctation on the head and the narrower pronotum, the longer elytra, smaller apical part of the median lobe and the shorter parameres;
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDD54BAB102E2" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
from
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D74EDD54BF88023E" baseAuthorityName="Cameron" baseAuthorityYear="1940" box="[260,399,1123,1149]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="tonkinesis">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D74EDD54BF88023E" box="[260,399,1123,1149]" italics="true" pageId="43" pageNumber="294">P. tonkinesis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the distinctly larger body, the absence of the longitudinal impressions in the middle of the narrower pronotum, the narrower apical portion of the median lobe and the shorter parameres.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDD9CBF0202AA" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
From all these species
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D791DD9CBC3A0286" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[475,573,1195,1221]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">P. rufula</taxonomicName>
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D403DD9CBC890286" bold="true" box="[585,654,1195,1221]" pageId="43" pageNumber="294">
|
|||
|
|
<taxonomicNameLabel id="00B257F2FFC26D73D403DD9CBC890286" box="[585,654,1195,1221]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
differs by the details of the external and internal structure of the aedeagus.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="61EF6510FFC26D73D68DDDC5BEF70372" pageId="43" pageNumber="294" type="distribution">
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDDC5BEF70372" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D68DDDC5BF66034F" bold="true" box="[199,353,1266,1292]" pageId="43" pageNumber="294">Distribution.</emphasis>
|
|||
|
|
<taxonomicName id="EEF54D18FFC26D73D721DDC4BC73034E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[363,628,1267,1293]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="294" phylum="Arthropoda" rank="species" species="rufula" status="sp. nov.">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D721DDC4BC73034E" box="[363,628,1267,1293]" italics="true" pageId="43" pageNumber="294">Paraphloeostiba rufula</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D4CADDC4BCC4034E" bold="true" box="[640,707,1267,1293]" pageId="43" pageNumber="294">
|
|||
|
|
<taxonomicNameLabel id="00B257F2FFC26D73D4CADDC4BCC4034E" box="[640,707,1267,1293]" pageId="43" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is known from the
|
|||
|
|
<typeStatus id="F64E8839FFC26D73D5E5DDC4BDD8034E" box="[943,991,1267,1293]" pageId="43" pageNumber="294">type</typeStatus>
|
|||
|
|
locality in central part of
|
|||
|
|
<collectingCountry id="51E2760BFFC26D73D352DDC4BEEB0372" name="Papua New Guinea" pageId="43" pageNumber="294">Papua New Guinea</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="61EF6510FFC26D73D68DDC0CBD6D03DE" pageId="43" pageNumber="294" type="etymology">
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDC0CBEF9033A" blockId="43.[151,1437,151,1437]" pageId="43" pageNumber="294">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D68DDC0CBF490316" bold="true" box="[199,334,1339,1365]" pageId="43" pageNumber="294">Etymology.</emphasis>
|
|||
|
|
The specific epithet is the Latin
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D48DDC0CBD110316" box="[711,790,1339,1365]" italics="true" pageId="43" pageNumber="294">rufulus</emphasis>
|
|||
|
|
, -
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D57ADC0CBD390317" box="[816,830,1339,1364]" italics="true" pageId="43" pageNumber="294">a</emphasis>
|
|||
|
|
, -
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D51CDC0CBD7F0317" box="[854,888,1339,1364]" italics="true" pageId="43" pageNumber="294">um</emphasis>
|
|||
|
|
(reddish). It alludes to the general coloration of the body.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="294A369BFFC26D73D68DDCB5BD6D03DE" blockId="43.[151,1437,151,1437]" box="[199,874,1410,1437]" pageId="43" pageNumber="294">
|
|||
|
|
<emphasis id="1B81EA89FFC26D73D68DDCB5BF4C03DF" bold="true" box="[199,331,1410,1436]" pageId="43" pageNumber="294">Bionomics.</emphasis>
|
|||
|
|
All specimens were collected flying in daylight.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|